西藏特提斯喜马拉雅海相白垩纪—古近纪生物地层与重大地质事件研究进展

地质历史时期曾发过许多对生命的演化进程造成过重大影响与制约的全球性地质事件,白垩纪—古近纪就是一个重大地质事件频发的时期。随着冈瓦纳大陆在中生代时期的解体,全球海陆格局发生了根本的变化,地球的表层和岩石圈层均发生了重大的改变,由此引发了构造运动空前活跃,发生过诸如大洋缺氧事件(OAE)、大洋富氧事件(CORBs)、白垩纪/古近纪(K/Pg)生物大灭绝事件、古新世—始新世极热(PETM)事件、印度-亚洲板块碰撞、新特提斯洋的演化及最终消亡等一系列的全球性重大地质事件。对这些重大地质事件的研究,有助于加深我们对古海洋、古地理、古环境的认识。尝试追踪和捕捉这些重大地质事件,恢复和重建古地理,其基础是建立精确的年代地层格架。西藏南部保存了中国最为完整的海相白垩纪—古近纪沉积,完整地记录了上述的全球性重大地质事件,通过对札达、岗巴、定日、亚东、江孜、萨嘎和吉隆等地区高分辨率浮游和底栖有孔虫、介形虫、钙质超微化石和放射虫生物地层学研究,可直接约束全球性重大地质事件发生的时间,并为重建新特提斯洋古海洋环境和古地理提供证据。此外,在重大地质环境突变期间生物的演化过程,也可为探明极端环境变化发生时期气候-环境-生物之间的协同演化关系提供证据。本文系统总结了课题组为主的近年来对藏南白垩纪—古近纪海相地层中微体古生物学的研究成果及重要进展,并对未来研究提出展望。     

塔里木盆地西部白垩纪—古近纪海相地层框架及对重大地质事件的记录

塔里木盆地西部白垩纪—古近纪发生了大规模的海侵事件,形成一个喇叭状向西开口的海湾,该海湾属于东特提斯洋的一个分支。该地区白垩纪—古近纪海相地层记录了东特提斯洋演化和一系列重大地质事件,具有重要的研究价值,但对其地层的研究仍相对薄弱,对重大地质事件的研究还不够深入。本文拟通过详细的岩石地层、生物地层和其他地层方法,完善地层划分与对比框架,并在此基础上讨论Cenomanian/ Turonian界线大洋缺氧事件(OAE2)、白垩纪/古近纪界线(K/Pg)、古新世—始新世极热事件(PETM)、特提斯海进与海退等一系列重大地质事件。塔里木盆地白垩纪—古近纪海相或海陆过渡相地层自下而上为克孜勒苏群、库克拜组、乌依塔克组、依格孜牙组、吐依洛克组、阿尔塔什组、齐姆根组、盖吉塔格组、卡拉塔尔组、乌拉根组和巴什布拉克组,上述地层中含有丰富的有孔虫、介形虫、钙质超微、沟鞭藻、孢粉、双壳类、腹足类等化石,以及少量菊石、腕足类、海胆和鲨鱼牙齿等化石。综合的生物地层和年代地层研究表明,克孜勒苏群的时代为早白垩世Barremian-Albian期,库克拜组—依格孜牙组的时代为晚白垩世Cenomanian-Maastrichtian期(Cenomanian/Turonian界线可能位于库克拜组三段),吐依洛克组的时代为白垩纪—古近纪过渡期;阿尔塔什组的时代为古新世早中期,齐姆根组为古新世晚期—始新世最早期,盖吉塔格组—乌拉根组的时代为中始新世中晚期,巴什布拉克组的时代为晚始新世,但不排除最上部进入渐新世早期。塔里木盆地的海侵开始于克孜勒苏群中上部沉积期(Albian晚期—Aptian早期),但规模很有限,大规模的海侵始于晚白垩世Cenomanian早期;从晚白垩世—古新世,共经历了5次大规模的海侵—海退事件;大约41 Ma前后,海水退出盆地南部的昆仑山山前,34 Ma前后,海水退出盆地北部的天山山前。上述海侵—海退事件可能受构造和全球海平面变化的双重影响,但构造事件对海侵的启动和结束可能更具决定性的影响。阐述了塔里木盆地西部白垩纪—古近纪海相地层所记录的OAE2、K/Pg界线、PETM和特提斯海侵—海退等事件,其中笔者及团队第一次在塔里木盆地西部齐姆根组中所发现和报道的PETM事件,将有助于揭示全球近岸地区PETM的特征和生物-环境响应。在未来的研究中,需要进一步厘清塔里木盆地西部地层序列,建立更加精细的生物地层和年代地层框架,加强对PETM和特提斯海侵—海退等重大地质事件的研究。     

塔里木盆地西部白垩纪—古近纪海相地层框架及对重大地质事件的记录

塔里木盆地西部白垩纪—古近纪发生了大规模的海侵事件,形成一个喇叭状向西开口的海湾,该海湾属于东特提斯洋的一个分支。该地区白垩纪—古近纪海相地层记录了东特提斯洋演化和一系列重大地质事件,具有重要的研究价值,但对其地层的研究仍相对薄弱,对重大地质事件的研究还不够深入。本文拟通过详细的岩石地层、生物地层和其他地层方法,完善地层划分与对比框架,并在此基础上讨论Cenomanian/Turonian界线大洋缺氧事件(OAE2)、白垩纪/古近纪界线(K/Pg)、古新世—始新世极热事件(PETM)、特提斯海进与海退等一系列重大地质事件。塔里木盆地白垩纪—古近纪海相或海陆过渡相地层自下而上为克孜勒苏群、库克拜组、乌依塔克组、依格孜牙组、吐依洛克组、阿尔塔什组、齐姆根组、盖吉塔格组、卡拉塔尔组、乌拉根组和巴什布拉克组,上述地层中含有丰富的有孔虫、介形虫、钙质超微、沟鞭藻、孢粉、双壳类、腹足类等化石,以及少量菊石、腕足类、海胆和鲨鱼牙齿等化石。综合的生物地层和年代地层研究表明,克孜勒苏群的时代为早白垩世Barremian-Albian期,库克拜组—依格孜牙组的时代为晚白垩世Cenomanian-Maastrichtian期(Cenomanian/Turonian界线可能位于库克拜组三段),吐依洛克组的时代为白垩纪—古近纪过渡期;阿尔塔什组的时代为古新世早中期,齐姆根组为古新世晚期—始新世最早期,盖吉塔格组—乌拉根组的时代为中始新世中晚期,巴什布拉克组的时代为晚始新世,但不排除最上部进入渐新世早期。塔里木盆地的海侵开始于克孜勒苏群中上部沉积期(Albian晚期—Aptian早期),但规模很有限,大规模的海侵始于晚白垩世Cenomanian早期;从晚白垩世—古新世,共经历了5次大规模的海侵—海退事件;大约41 Ma前后,海水退出盆地南部的昆仑山山前,34Ma前后,海水退出盆地北部的天山山前。上述海侵—海退事件可能受构造和全球海平面变化的双重影响,但构造事件对海侵的启动和结束可能更具决定性的影响。阐述了塔里木盆地西部白垩纪—古近纪海相地层所记录的OAE2、K/Pg界线、PETM和特提斯海侵—海退等事件,其中笔者及团队第一次在塔里木盆地西部齐姆根组中所发现和报道的PETM事件,将有助于揭示全球近岸地区PETM的特征和生物环境响应。在未来的研究中,需要进一步厘清塔里木盆地西部地层序列,建立更加精细的生物地层和年代地层框架,加强对PETM和特提斯海侵—海退等重大地质事件的研究。     

年代古地理学: 理论与实践*

古地理学研究和探讨的是地质历史时期的地表环境,是构造—环境(气候)—物质相互作用的产物。传统的古地理学及相应的构造古地理、生物古地理学等对大地构造演化及古大陆重建、油气勘探等起到了重要的作用。随着地球演化的深入研究,在多层圈的相互作用,特别是古气候、古环境等研究基础上,需要有机地把地表环境和构造环境相结合,并且综合考虑深层及浅层地质过程对古地理格局的形成和演化的制约。在传统的古地理学基础上,提出一个新的学科——年代古地理学,用以研究和探讨地质历史时期的地表过程。该学科是精细的同位素年代格架下的不同动力学—运动学—物质迁移—环境变迁形成古地理的研究,或者物质—力学—环境(气候)相互作用下的古地理格局的研究。旨在研究和探讨地表过程,亦即形成地表环境的过程,强调同一时间的多元性、不同空间的差异性和不同尺度的穿时性。文中以近年来研究的台湾、扬子及华北克拉通等地区的精细时间尺度下的古地理环境为例,进一步剖析年代古地理学对于重新理解和认识古构造格局演化的重要性。     

年代古地理学: 理论与实践*

古地理学研究和探讨的是地质历史时期的地表环境,是构造—环境(气候)—物质相互作用的产物。传统的古地理学及相应的构造古地理、生物古地理学等对大地构造演化及古大陆重建、油气勘探等起到了重要的作用。随着地球演化的深入研究,在多层圈的相互作用,特别是古气候、古环境等研究基础上,需要有机地把地表环境和构造环境相结合,并且综合考虑深层及浅层地质过程对古地理格局的形成和演化的制约。在传统的古地理学基础上,提出一个新的学科——年代古地理学,用以研究和探讨地质历史时期的地表过程。该学科是精细的同位素年代格架下的不同动力学—运动学—物质迁移—环境变迁形成古地理的研究,或者物质—力学—环境(气候)相互作用下的古地理格局的研究。旨在研究和探讨地表过程,亦即形成地表环境的过程,强调同一时间的多元性、不同空间的差异性和不同尺度的穿时性。文中以近年来研究的台湾、扬子及华北克拉通等地区的精细时间尺度下的古地理环境为例,进一步剖析年代古地理学对于重新理解和认识古构造格局演化的重要性。     

塔里木盆地西北缘乌恰地区晚白垩世-渐新世微相分析及沉积演化

塔里木盆地西部是我国发育中新生代海相地层的少数地区之一。以沉积微相分析为手段,通过对塔里木盆地西北缘乌恰地区库孜贡苏剖面晚白垩世-古近纪岩性、生物组合、颗粒成分、基质类型及其沉积结构和构造特征等仔细研究,重点针对碳酸盐岩划分出（含）骨屑隐晶灰岩、隐晶灰岩、微（隐）晶白云岩、鸟眼隐晶灰岩、微晶鲕粒白云岩、生物碎屑灰岩、微晶球粒白云岩、生物灰岩、亮晶鲕粒灰岩等9个岩相类型。根据碳酸盐岩微相类型组合和剖面结构沉积特征,划分出潮上带、潮间带、潮下带、台地边缘浅滩和生物礁5个沉积相,并恢复了该区晚白垩世-古近纪的沉积环境演化过程,为查清白垩纪-古近纪特提斯洋演化以及重建该地区古环境和油气勘探研究提供重要的基础材料。     

白垩纪—古近纪界限时期的陆地气候重建进展

白垩纪—古近纪(K-Pg)界限时期发生了地质历史中最严重的生物大灭绝事件之一,重建该时期气候演化历史及其对生态系统的影响对理解深时气候演化规律和预测未来气候变化都具有重要意义。相比海相剖面而言,基于陆相剖面的白垩纪—古近纪界限时期古气候重建研究开展较晚,相关成果缺乏系统总结和集成。本文系统回顾了全球陆相白垩纪—古近纪界限剖面的分布情况和事件年代学新进展,总结了基于以上剖面重建的定量古气候记录和德干火山喷发记录,建立了界限时期中纬度陆地年平均温度和大气CO₂浓度时间演化序列。陆相剖面分布相对集中且数量较少,中国是该时期陆相剖面发育最多的国家之一,多个盆地已经建立了德干火山喷发记录,在重建该时期陆地气候上具有很大潜力。基于陆相沉积物汞元素地球化学记录表明,德干火山作用导致了晚马斯特里赫特期变暖事件并触发了白垩纪末期生物大灭绝。中纬度陆地年平均温度重建结果显示马斯特里赫特期中期全球经历了～10℃的降温,到马斯特里赫特期晚期德干火山初始喷发前显著升温至～20℃,并在白垩纪—古近纪界限前后发生多次快速波动。尽管65～63 Ma之间陆地温度记录相对缺失,69～65 Ma之间重...     

白垩纪地球表层系统重大地质事件与温室气候变化研究

国家重点基础研究发展计划（973）项目“白垩纪地球表层系统重大地质事件与温室气候变化”，将以白垩纪与碳循环相关的重大地质事件和温室气候变化的关系为主线，以大洋缺氧事件—富氧作用转变过程和机制研究为突破口，进行海陆相整合研究，重点追溯东特提斯洋和我国大陆地球表层系统重大地质事件的记录，探讨这些事件与碳循环、快速气候变化的正/负反馈机制。项目将充分利用中国大陆发育完好的白垩纪海相、陆相地层及古生物记录，通过松辽盆地白垩系科学钻探全岩芯取样和多学科综合研究的途径，着眼于厘定反映地质事件和气候变化的层位及标志和时间格架，解决高分辨率海、陆相沉积事件的精确对比，分析地层记录中气候标志和古生物类群的地理分布，集中研究陆地和海洋环境对同一事件的响应机制，重溯白垩纪地球表层系统重大地质事件过程及成因，探究陆相烃源岩大规模形成、陆地生物群更替与温室气候变化和碳循环之间的正/负反馈关系和机制等科学问题，为预测全球长时间尺度上的气候变化趋势提供科学依据。     

序

<正>特提斯地质的研究涉及到全球构造、地壳和岩石圈演化、洋陆变迁等重大地质学理论问题,因此特提斯演化是一个长期被关注的科学问题。近年来,越来越多的研究者认为"三江"地区特提斯演化阶段存在多个陆壳地块与洋盆相间排列的多岛洋构造-古地理格局,从洋盆扩张到俯冲汇聚碰撞造山,经历了从特提斯大洋岩石圈向大陆岩石圈转换及其大陆边缘多岛弧     

洋板块地质研究进展

中国存在多个时代、多种类型的造山带,发育了多种多样的俯冲增生杂岩带,经历了复杂多变的洋陆转换过程,如何揭示包括洋内演化和洋陆转换等的造山过程一直是一个难题。为此,中国区域地质志项目组提出了洋板块地质研究,试图通过对造山系俯冲增生杂岩带、蛇绿岩带等洋岩石圈地质建造、结构构造进行系统研究,再造洋岩石圈从洋中脊形成到海沟俯冲消亡、转换成陆的地质作用全过程。本文介绍了洋板块地质提出到现今主要的研究进展,包括四个方面。一是,初步建立了洋板块地质格架,洋板块地质的研究包括俯冲增生杂岩的物质组成、蛇绿岩类型及其形成的构造环境、洋板块沉积组合和洋板块地层、岛弧火成岩组合、洋陆转换的过程和机制、洋-陆转换过程与成矿作用等重要内容。二是,识别出北山牛圈子—马鬃山、嘉荫—依兰、陈蔡、东昆仑布青山—阿尼玛卿、鹰扬关、大洪山、甘孜—理塘、新余神山—新干神政桥等中国陆域62条主要的俯冲增生杂岩带/增生杂岩带。俯冲增生杂岩带是认识、理解造山系时空结构、组成和演化的关键。三是,在祁连地区识别出较为完整的洋内弧岩石组合。洋盆演化形成大陆过程中的洋内俯冲带是大陆的诞生地,洋内俯冲作用形成的洋内弧是洋盆演化形成大陆的初始弧。洋内弧火成岩组合序列的发现为研究洋陆转换过程提供了岩石学依据。祁连造山带是洋板块地质研究的经典地区之一。研究显示,当金山出露完整的洋内弧岩石组合,这些岩石记录了洋内弧从初始俯冲到发育成熟的全过程,为探讨祁连造山带原特提斯洋构造演化提供了新的依据。四是,制定了洋板块地质构造图编图方案,编图内容主要包括俯冲增生杂岩带、岩浆弧、高压-超高压带、俯冲期和碰撞期构造形变要素和构造演化等。编图单元分为三级:一级为俯冲增生杂岩带;二级为岩片;三级包括基质和岩块。编图过程中需要明确岩浆弧的性质和归属,明确图面上某一岩浆弧与哪个蛇绿混杂岩或大洋配套。图面上对于构造要素的表达重点是区分俯冲和碰撞阶段。通过构造变形的时态、相态、位态研究,识别俯冲期和碰撞期的构造变形形迹。这是洋板块地质初步的研究成果,以俯冲增生杂岩带的研究为基础,探讨特提斯洋等大洋的演化、中国东部古太平洋/太平洋转换与中新生代成矿关系等重大基础地质问题是洋板块地质研究下一步的工作方向。目前,洋板块地质的研究还处于试点阶段,洋板块地质与成矿的成因联系等重大地质问题尚需今后更深入地研究。     

Biostratigraphy of the Garau Formation (Berriasian?–lower Cenomanian) in central part of Lurestan zone,northwest of Zagros,Iran

Deposition of organic rich black shales and dark gray argillaceous limestones in the Berriasian–Turonian interval has been documented in many parts of the world. Northwest of Zagros, Iran (Lurestan zone), thin bedded black shales and marls, dark gray argillaceous limestones and fissile limestone layers, having bitumen, of the Garau Formation are deposited. For biostratigraphic studies two stratigraphic sections including one surface section (Kuzaran) and one subsurface section (Naft well) were selected, respectively. In this study, 61 foraminiferal species belonging to 17 genera have been identified, and 12 biozones were recognized. Based on fossils distribution and biozones identification, the age of the Garau Formation is Berriasian?–early Cenomanian. In addition, the micropalaeontological study demonstrated a variety of widespread morphological changes in planktonic foraminifera assemblages (e.g., the elongation of the final chambers, appearance of twin chambers in the last whorl). These changes coincide with deposition of argillaceous limestones and marls rich in organic matter, indicating oceanic anoxic events. On this basis, three oceanic anoxic events such as OAE1a, OAE1b and OAE1d were recognized in Naft well section and two (OAE1b and OAE1d) in Kuzaran section.     

Cenomanian-Coniacian Sea-level Change and Dissolved Oxygen Fluctuations in Tethys-Himalaya: Evidences from Benthic Foraminifera of Gamba, Tibet

Benthic foraminifera, preserved in the Late Cretaceous organic carbon-rich sediments of Gamba, southern Tibet, provide high-resolution proxies for sea-level changes and dissolved oxygen fluctuations of southeastern Tethys. The fossils were statistically analyzed and divided into three faunas of "Cenomanian fauna", "Turonian fauna", and "Coniacian fauna". A middle neritic-upper bathal environment (50-250m) was estimated considering the ratios of planktonic and epifaunal benthic foraminifera (P/(P+E)), the morphological analysis according to the studies of recent foraminifera and the abundant distributions of depth-related species such as Alabamina creta, Laevidentalina sp., Praebulimina spp., Pleurostomella cf. naranjoensis, Pyrulina sp., Quinqueloculina spp., Haplophragmoides spp., etc. The result shows an almost parallel trend with the global transgressive and regressive cycles, but the former fluctuates more frequently at upper Cenomanian, which probably indicates tectonic instability of the continental margin. According to the benthic foraminiferal richness (BFN), Shannon-Weiner diversity (H(s)), as well as benthic foraminiferal oxygen index (BFOI), five periods of oxygen depleted conditions (dysoxic-anoxic) have been recognized. They correspond to the OAE2, the lower Turonian, the upper Turonian, the Turonian-Caniacian boundary event and the probably OAE3. In addition, the oxygen fluctuations in Gamba might be controlled directly by sea-level changes, while the paleoproductivity and oxygen conditions interacted with each other under oxygen deficiency environments.     

Stratigraphy of the Cenomanian–Turonian Oceanic Anoxic Event OAE2 in shallow shelf sequences of NE Egypt

Two shallow water late Cenomanian to early Turonian sequences of NE Egypt have been investigated to evaluate the response to OAE2. Age control based on calcareous nannoplankton, planktic foraminifera and ammonite biostratigraphies integrated with δ¹³C stratigraphy is relatively good despite low diversity and sporadic occurrences. Planktic and benthic foraminiferal faunas are characterized by dysoxic, brackish and mesotrophic conditions, as indicated by low species diversity, low oxygen and low salinity tolerant planktic and benthic species, along with oyster-rich limestone layers. In these subtidal to inner neritic environments the OAE2 δ¹³C excursion appears comparable and coeval to that of open marine environments. However, in contrast to open marine environments where anoxic conditions begin after the first δ¹³C peak and end at or near the Cenomanian–Turonian boundary, in shallow coastal environments anoxic conditions do not appear until the early Turonian. This delay in anoxia appears to be related to the sea-level transgression that reached its maximum in the early Turonian, as observed in shallow water sections from Egypt to Morocco.     

Oxygen level,primary productivity,and water turbulence during the OAE2 interval of Zagros Basin (SW Iran): Benthic foraminiferal variations in the carbonate microfacies

Cenomanian/Turonian boundary (upper Sarvak Formation) benthic foraminiferal assemblages were analyzed to reconstruct oxygen level, primary productivity, and water turbulence in the Izeh Zone, Zagros Basin. The interplay between environmental perturbations during the Oceanic Anoxic Event 2 (OAE2) and regional tectonic activities in the Zagros Basin resulted in formation of various benthic foraminiferal assemblages in the study section. The OAE2 interval at the region of study starts with extinction of rotaliporids at the onset of δ¹³C positive excursion (peak “a”), which is associated with population of infaunal benthic foraminifera (especially Bolivina alata). The following interval at the onset of Whiteinella archaeocretacea Biozone is characterized by the total absence of benthic taxa and dominance of planoheterohelicids (“Heterohelix shift”) in the black shale strata, indicating expansion of oxygen minimum zone and unhospitable conditions for both benthic and planktic foraminifera. The upper part of OAE2 interval (including δ¹³C peaks “b” and “c”) coincides with harbinger of Neo-Tethys closure in the Arabian Plate, causing a compressional tectonic regime, and creation of uplifted terrains in the basin. The relative sea level started to locally fall in this succession, which was accompanied by a better ventilation of seafloor, lower TOC contents, and reappearance of benthic foraminifera.     

古新世/始新世全球增温在西藏岗巴地区的表现

随着深海钻探计划(DSDP)和大洋钻探计划(ODP)的开展,在古海洋学领域取得了许多重大发现。古近纪古新世与始新世(P/E)界线间的地质事件的发现便是其重要成果之一。西藏南部岗巴地区发育有良好的海相界线地层。对生物群、碳、氧、锶稳定同位素以及磁化率的研究显示,全球界线事件在西藏南部岗巴地区具有明显的影响。古新统宗浦组顶部底栖有孔虫动物群阶段性绝灭,总灭绝率为69%。始新统遮普惹组底部有孔虫逐渐复苏,除12种为古新世的残存类型外,始新世生物组合全为新的属种,并呈现三个复苏阶段。碳稳定同位素表现为三期负向偏移,在界线处出现-4‰的异常。这一异常与全球碳稳定同位素事件表现一致。锶同位素在界线位置没有明显的变化,而明显的峰值出现在界线之上,并与全球53Ma出现的锶同位素变化相对应。磁化率在界线之上2m处出现一个很明显的波动,与第3期碳同位素异常吻合。所有研究结果表明,西藏南部(特提斯-喜马拉雅低纬度浅海环境)在古新世与始新世界线时期出现明显的气候变化,说明全球深海温度增高在浅海环境具有相同表现,只是影响时间较长。     

Stratigraphy of the Cenomanian–Turonian Oceanic Anoxic Event OAE2 in shallow shelf sequences of NE Egypt

Two shallow water late Cenomanian to early Turonian sequences of NE Egypt have been investigated to evaluate the response to OAE2. Age control based on calcareous nannoplankton, planktic foraminifera and ammonite biostratigraphies integrated with δ¹³C stratigraphy is relatively good despite low diversity and sporadic occurrences. Planktic and benthic foraminiferal faunas are characterized by dysoxic, brackish and mesotrophic conditions, as indicated by low species diversity, low oxygen and low salinity tolerant planktic and benthic species, along with oyster-rich limestone layers. In these subtidal to inner neritic environments the OAE2 δ¹³C excursion appears comparable and coeval to that of open marine environments. However, in contrast to open marine environments where anoxic conditions begin after the first δ¹³C peak and end at or near the Cenomanian–Turonian boundary, in shallow coastal environments anoxic conditions do not appear until the early Turonian. This delay in anoxia appears to be related to the sea-level transgression that reached its maximum in the early Turonian, as observed in shallow water sections from Egypt to Morocco.     

Planktonic foraminiferal biostratigraphy and assemblage composition across the Cenomanian–Turonian boundary interval at Clot Chevalier (Vocontian Basin,SE France)

The Cenomanian–Turonian boundary interval is generally considered a critical time for planktonic foraminifera due to the environmental perturbations associated with Oceanic Anoxic Event 2. However, only the rotaliporids became extinct at the onset of the event, whilst several lineages evolved and/or diversified. This remarkable morphologic plasticity is often overlooked in the literature, partly because a number of stratigraphic sections have only been studied in thin-section due to the degree of lithification of the samples. Improved documentation of the morphological variability of planktonic foraminifera and better defined species concepts are required in order to improve biostratigraphy, particularly as Helvetoglobotruncana helvetica is an unreliable marker for the base of the Turonian. At the same time, detailed study of the planktonic foraminiferal response to OAE 2 demands a more profound knowledge of the assemblage composition.We present new biostratigraphic, taxonomic, and quantitative data for planktonic foraminiferal species from the Clot Chevalier section (Vocontian Basin, SE France), with the aim of (1) providing a detailed biostratigraphic analysis of the section, (2) documenting the morphological plasticity of specimens in this time interval and stabilizing species concepts, and (3) identifying promising markers to improve the resolution of the present biozonation and allow regional correlation. Samples were processed with acetic acid to extract isolated planktonic foraminifera. Assemblages were assigned to the upper Cenomanian Rotalipora cushmani Zone and to the uppermost Cenomanian–lowermost Turonian Whiteinella archaeocretacea Zone. Planktonic foraminiferal bioevents and assemblage composition identified at Clot Chevalier are compared with the well-studied Pont d'Issole section located ca. 15 km to the NE, highlighting similarities and differences in the species occurrences that may complicate the stratigraphic correlation between the two sections.The results of our study support the validity and common occurrence of species that have been misidentified and/or overlooked in the literature (i.e., Dicarinella roddai, Praeglobotruncana oraviensis, Marginotruncana caronae) and indicate that primitive marginotruncanids evolved before the onset of OAE 2, although species diversification occurred only after the event. Moreover, we believe that the first appearance of P. oraviensis might represent a promising bioevent for approximating the Cenomanian/Turonian boundary, after calibration with bio- and chemostratigraphically well-constrained sections. Finally, we describe three new trochospiral species, named “Pseudoclavihedbergella” chevaliensis, Praeglobotruncana pseudoalgeriana and Praeglobotruncana clotensis.     

中—新生代有孔虫古地理对西藏特提斯演变的动态响应

有孔虫化石资料是地质历史的真实记录,对不同地质时期古地理格局和生态环境的变迁具有动态响应。西藏特提斯构造带的演化、板块相对地理位置变迁等诸多问题一直是地学界关注的热点。研究西藏特提斯沉积盆地内有孔虫动物群的古生态特征和古地理分布,能够识别生物地理区系,进而恢复不同时期的大地构造演化格局。西藏地区中、新生代古生物地理区系的分化是西藏特提斯地质演变的具体反映。西藏南部早侏罗世产底栖大有孔虫Orbitopsella喜暖动物群,晚侏罗世出现双壳类Buchia喜冷动物群。由此推测,侏罗纪新特提斯洋扩张尤其是中大西洋的开张,将位于印度大陆北缘的特提斯喜马拉雅带,从早侏罗世较低纬度的温暖位置向南推移至较高纬度的低温地区。白垩纪中期Orbitolina有孔虫类群繁盛于特提斯北侧亚洲大陆的拉萨地块和羌塘盆地,但没有出现在印度大陆。这说明当时印度大陆已脱离冈瓦纳大陆向北漂移,受四周深水环境的阻隔,Orbitolina动物群未能向印度大陆扩散。此时深水环境中生活着浮游有孔虫Ticinella-Rotalipora动物群。Turonian晚期开始形成海退,拉萨地块的海洋环境基本消失。Coniacian-Campanian早期印度大陆北缘浮游有孔虫继续占优势,繁盛Marginotruncana-Globotruncana动物群。直至白垩纪末,印度和欧亚大陆之间的深海阻隔仍然存在,雅鲁藏布江缝合带两侧动物群一直存在根本性差异。印度大陆北缘发育着Orbitoides-Omphaloceclus 动物群,冈底斯南缘则以Lepidorbitoides-Pseudorbitoides动物群为特征。古新世Danian期生态环境发生变化,显示大印度与亚洲大陆发生初始碰撞(66~61 Ma)。Selandian期之后,缝合带两侧才出现相同的Miscellanea-Daviesina有孔虫类群,生物区系的分异基本结束。始新世早期缝合带两侧为完全相同的生物区系,共同发育底栖大有孔虫Nummulites-Discocyclina动物群。有孔虫古地理证据表明,大印度与欧亚大陆的初始碰撞在古新世早期发生,时间大致在Danian期,沿雅鲁藏布缝合带的深海演变为残留海环境。小个体货币虫Nummulites willcoxi和浮游有孔虫Globigerina ouachitaensis的存在,代表特提斯喜马拉雅最高海相沉积,时代属于始新世Priabonian晚期(35~34 Ma)。随后,特提斯喜马拉雅海封闭,海水完全退出西藏境内。     

对西藏岗巴上白垩统的新认识

在前人研究的基础上对岗巴地区上白垩统的划分做了厘定 ,进一步建立了晚白垩世的 1 2个浮游有孔虫化石带。根据岩性特征及化石带的研究 ,上白垩统被划分为赛诺曼期至土仑早期的冷青热组 ;土仑中期至三冬期的岗巴村口组和康潘期至马斯特里赫特期的宗山组。该区赛诺曼期与土仑期的界线位于冷青热组上部 ,以 H elvetoglobotruncana praehelvetica的初现为标志。     

Repeated anoxia–extinction episodes progressing from slope to shelf during the latest Cenomanian

Oceanic Anoxic Event 2 (OAE 2) during the Cenomanian–Turonian (C/T) transition caused stepwise marine extinctions. Using organic compounds, stable carbon and oxygen isotopes, and foraminifera from three depth-transect sections in northern Spain, this study revealed repeated anoxic/euxinic events coinciding with warming and stepwise extinctions of planktonic and/or benthic foraminifera within intermediate to surface waters in the proto-North Atlantic during the C/T transition. Those short-duration euxinic events occurred four times: at 93.95 Ma, marked by the extinction of Rotalipora greenhornensis; at 93.90 Ma, marked by the extinction of Rotalipora cushmani; at the mid-time maximum of the plateau of the δ¹³C of carbonates (93.70 Ma); and at the time of the C/T boundary (93.55 Ma). Furthermore, the main benthic foraminiferal extinctions occurred during the first and second euxinic events in the upper slope, during the second and third euxinic events in the outer to middle shelf, and during the third and fourth events in the middle shelf. The main euxinic events in each section also showed a progression to the shallow shelf. The main anoxia–extinction events occurred in the upper slope and outer shelf then moved to the middle shelf. The shallowest section had relatively weak anoxia and a proportionally low extinction rate. These new findings indicate that foraminiferal extinctions started from the intermediate water and the continental slope and then moved to the continental shelf. This was the result of the repeated progression of euxinic–anoxic water from the upper slope to the middle shelf on the eastern continental margin of the proto-North Atlantic four times during a 400 kyr period, to the end of the Cenomanian.