U‐series dating of the Late Pleistocene mammalian fauna from Wood Quarry (Steetley), Nottinghamshire,UK

We present the U‐series dating of bones from Wood Quarry (‘Steetley Quarry Cave’) using the diffusion–adsorption model to account for uranium uptake. The results give a weighted mean date of 66.8 ± 3.0 kyr, placing this assemblage within or just before Marine Oxygen Isotope Stage 4. The fauna is thought to correlate with the Banwell Bone Cave mammal assemblage‐zone of the Early Devensian in Britain. Our results support the idea that this assemblage‐zone immediately precedes the assemblage represented nearby at Pin Hole in Creswell Crags which is contemporary with the Mid‐Devensian and correlates with MIS 3. Our dates, and dates for the Banwell Bone Cave mammal assemblage‐zone from Stump Cross Cavern and evidence from other sites may indicate a longevity for this fauna. Copyright © 2005 John Wiley & Sons, Ltd.     

Palynological assemblages of non-marine rocks at the Permian–Triassic boundary, western Guizhou and eastern Yunnan, South China

Marine and non-marine facies of the Permian–Triassic boundary stratigraphic set (PTBST) are well developed in South China. Palynological assemblages enable subdivision and correlation of the Permian–Triassic boundary (PTB) rocks. Three palynological assemblages are recognized across the PTBST in two terrestrial PTB sections in western Guizhou and eastern Yunnan, South China. Assemblage 1 (Xuanwei Formation) is a Late Permian palynological assemblage dominated by ferns and pteridosperms, with minor gymnosperms. Most taxa are typical long-ranging Paleozoic forms, but the appearance of Lueckisporites confirms a Late Permian age for this assemblage. Assemblage 2 (PTBST) is marked by an abrupt decrease in palynomorph abundance and diversity, and thriving fungal/algal(?) spores. Assemblage 2 is still dominated by ferns and pteridosperms, with a few gymnosperms, but is characterized by a mixed palynoflora containing both Late Permian and Early Triassic elements. Most taxa are typical Late Permian ones also found in Assemblage 1, however, some taxa of Early Triassic aspect, e.g. Lundbladispora and Taeniaesporites, appeared for the first time. In Assemblage 3 (top Xuanwei Formation and Kayitou Formation), the proportion of gymnosperm pollen increases rapidly, exceeding that of ferns and pteridosperms, but the abundance of palynomorphs is still low. Typical Early Triassic taxa (such as Lundbladispora, Aratrisporites and Taeniaesporites) are present in greater abundance and confirms an Early Triassic age for this assemblage.     

Species identification and decay assessment of Late Pleistocene fragmentary vertebrate remains from Pin Hole Cave (Creswell Crags,UK) using collagen fingerprinting

Ancient bone remains are widely utilized when investigating vertebrate biodiversity of past animal populations but are often so highly fragmented that the majority of specimens cannot be identified to any meaningful taxonomic level. Recently, high‐throughput methods for objective species identification using collagen peptide mass fingerprinting have been created to overcome this with the added indication that they could also offer a means of relative ageing through decay measurement. Here we explore both species identification and decay measurements for the Pin Hole Cave ‘microfaunal’ assemblage, the site that has been designated as a representative for Marine Oxygen Isotope Stage 3 in Britain in terms of its suite of mammalian fauna. We explore the technique's potential to corroborate the faunal diversity established previously by macroscopic studies and evaluate the decay measurements across the species boundary. The results support that the analysis of fragmentary remains by collagen fingerprinting can yield a more diverse set of fauna, and offer additional information relating to taphonomy, than the analysis of morphologically intact bones on their own. However, although useful for identifying likely contaminations of an assemblage, there was an unexpected decrease in the decay measurements observed for some megafauna compared with much younger microfauna, indicating that other factors need to be carefully monitored before it could be used as a relative ageing technique in Quaternary deposits.     

山东济宁煤田(东区)晚古生代孢粉组合

鉴定了山东济宁煤田(东区)晚古生代孢粉组合中的孢粉化石70属189种.据孢粉化石在地层中的分布和垂向的变化规律自上而下建立了5个孢粉组合:Ⅴ. Cyclogranisporites pressus-Lycospora pellucida组合, Ⅳ. Granulatisporites piroformis组合,Ⅲ. Sinulatisporites sinensis组合,Ⅱ. Thymospora pseudothiessenii-T. thiessenii组合,Ⅰ. Densosporites annulatus组合.通过与邻区及华北部分地区对比,确定本区孢粉组合的地质时代应为:晚石炭世晚期(组合Ⅰ),早二叠世早期(组合Ⅱ),早二叠世中期(组合Ⅲ),早二叠世晚期(组合Ⅳ)和晚二叠世早期(组合Ⅴ),本区石炭系-二叠系界线应置于太原组第11层灰岩之底.     

Dietary niche separation of three Late Pleistocene bear species from Vancouver Island,on the Pacific Northwest Coast of North America

Competition between taxa related to climate changes has been proposed as a possible factor in Pleistocene megafaunal extinctions, and here we present isotope evidence of the diets of three co-existing bear species [black bear (Ursus americanus), brown bear (Ursus arctos), and the now extinct short-faced bear (Arctodus simus)] from a locale in western North America dating to the Late (Terminal) Pleistocene (~14.5–11.7 ka). The three bear species were found at several sites on Vancouver Island, on the western coast of Canada. To examine the chronological overlap and niche partitioning between these species of bear, we used direct radiocarbon dating, stable isotope analysis and ZooMS proteomic identification methods. Here we present new radiocarbon evidence from Terminal Pleistocene U. americanus, U. arctos and A. simus from several sites on the island, along with both bulk collagen and compound-specific isotope data for these species. Radiocarbon dates confirm the chronological overlap of Arctodus and both Ursus species in the montane regions of the island at the end of the Pleistocene. Stable isotope data reveal niche differentiation between these species, with U. americanus occupying a distinctly lower trophic position than the other two taxa.     

Pleistocene oceanographie changes indicated by deep sea benthic foraminifera in the northern Indian Ocean

An attempt has been made to understand the Pleistocene bottom water history in response to the paleoclimatic changes in the northern Indian Ocean employing quantitative analyses of deep sea benthic foraminifera at the DSDP sites 219 and 238. Among the 150 benthic foraminifera recorded a few species show dominance with changing percent frequencies during most of the sequence. The dominant benthic foraminiferal assemblages suggest that most of the Pleistocene bottom waters at site 219 and Early Pleistocene bottom waters at site 238 are of North Indian Deep Water (NIDW) origin. However, Late Pleistocene assemblage at site 238 appears to be closely associated with a water mass intermediate between North Indian Deep Water (NIDW) and Antarctic Bottom Water (AABW). Uvigerina proboscidea is the most dominant benthic foraminiferal species present during the Pleistocene at both the sites. A marked increase in the relative abundance ofU. proboscidea along with less diverse and equitable fauna during Early Pleistocene suggests a relative cooling, an intensified oceanic circulation and upwelling of nutrient rich bottom waters resulting in high surface productivity. At the same time, low sediment accumulation rate during Early Pleistocene reveals increased winnowing of the sediments possibly due to more corrosive and cold bottom waters. The Late Pleistocene in general, is marked by relatively warm and stable bottom waters as reflected by low abundance ofU. proboscidea and more diverse and equitable benthic fauna. The lower depth range for the occurrence ofBulimina aculeate in the Indian Ocean is around 2300 m, similar to that of many other areas.B. aculeata also shows marked increase in its abundance near the Pliocene/Pleistocene boundary while a sudden decrease in the relative abundance ofStilostomella lepidula occurs close to the Early/Late Pleistocene boundary.     

New records of temperate mollusks in two Late Pleistocene terrestrial localities from northeastern Oaxaca,Southern Mexico

The Mixteca Alta Oaxaqueña is in the state of Oaxaca, southern Mexico. This region is characterized by numerous Pleistocene fossiliferous localities. The objective of this study is to describe a diverse assemblage of Late Pleistocene freshwater and terrestrial mollusks in two localities from northeastern Oaxaca, Coixtlahuaca District. We identified 10 taxa of gastropods and one of bivalves. By the sedimentological characteristics and the mollusks assemblage, it is possible to relate the first locality with meandriform river deposits, without vegetation. The second locality was associated with a floodplain with short-lived associated vegetation. Five identified species constitute the most austral records of these taxa in Neartic Realm. In all the taxa, the Late Pleistocene occurrences constitute the last records of the identified mollusks in the study zone.     

Changes in ostracod assemblages and morphologies during lake‐level variations of Lago Cardiel (49°S), Patagonia,Argentina, over the last 15.6 ka

Changes in the ostracod assemblages from two sediment cores collected from Lago Cardiel in southeastern Patagonia (49°S) reflect the main regional abrupt climatic changes over the last 15.6 cal. ka BP. Shifts in species abundance and switches in dominances suggest that these were mainly driven by variable salinity. During the Late Pleistocene, Limnocythere rionegroensis was abundant and dominant, indicating waters with high salinity and prevalence of evaporative processes. Between 12.6 and 10.8 cal. ka BP, Lago Cardiel expanded markedly and reached an Early Holocene highstand of +55 m above present lake level. A major change in ostracod assemblage in which Limnocythere patagonica appears as the dominant species in parallel with the disappearance of L. rionegroensis and Eucypris aff. cecryphalium mirrored this transitional period between the cold and dry Late Pleistocene and the humid and warm Early Holocene. Over the last 4 cal. ka BP, L. rionegroensis returned to the species assemblage and Riocypris whatleyi increased its abundance pointing towards increasing salinities. The variations in size, shape and ornamentation of L. rionegroensis and R. whatleyi fossil valves were examined using geometric morphometric techniques and further compared to those of modern Patagonian sites. Limnocythere rionegroensis specimens displayed high morphological variability during the evolution of Lago Cardiel. More specifically, the switch in reproductive mode – from sexual to parthenogenetic– and the increase in valve ornamentation around 12.7 cal. ka BP suggest that these changes were promoted by the hydrological alteration that occurred in the Late Pleistocene. This exercise provides a robust range of morphological variation for these proxies, which will be useful in further taxonomic and palaeoenvironmental studies adding more information about different factors influencing the observed morphological trends.     

Coastal curios? An analysis of ex situ beach finds for mapping new Palaeolithic sites at Happisburgh,UK

Recent archaeological discoveries from exposures of the Cromer Forest-bed Formation at Happisburgh, UK, have radically changed interpretations of the nature and timing of early hominin occupation of northern latitudes, but this in situ archaeology is only one part of the picture. Surface finds of Pleistocene mammalian remains have been found along this coastline for centuries, with stone tools adding to this record over the past 7 years. The ex situ nature of these finds, however, means they are often seen as limited in the information they can provide. This work contributes to a growing body of research from a range of landscape and environmental contexts that seeks to demonstrate the value and importance of these ex situ assemblages. Here the focus is on Palaeolithic flint artefacts and Pleistocene mammalian remains recovered by a group of local collectors through systematic, GPS-recorded beach collection from 2013 to 2017, and their use in developing a methodology for working with ex situ Palaeolithic finds in coastal locations. The results demonstrate significant patterning that identifies unexplored exposures both onshore and offshore, considerably expanding the known extent of deposits and facilitating new insights into the wider archaeological landscape associated with the earliest occupation of northern Europe.     

Distinguishing between naturally and culturally flaked cobbles: A test case from Alberta,Canada

Distinguishing between naturally and culturally produced, simply flaked cobbles has been a problem for proponents of a pre‐Clovis occupation in the Americas. Several sites in Alberta have been assigned a pre‐Clovis status based on the presence of simply flaked cobbles found in Late Pleistocene till deposits. Historically, these types of assemblages have been assigned a cultural status based on subjective criteria and appeals to the analyst's expertise. To determine the archaeological status of two such assemblages from Alberta (Varsity Estates and Silver Springs), they were compared to a known natural assemblage and two known cultural assemblages. Chi‐square testing was used to evaluate several lithic attributes. Only those attributes that statistically differentiated between natural and cultural assemblages were used for further analyses. All cobbles were then scored using these attributes. A point was awarded when a statistically significant attribute of human‐manufacture was present. These points were then totaled, providing an aggregate score for each cobble. These scores were plotted to determine whether the test assemblages had closer affinities with the known natural or known cultural assemblages. The results indicate that the proposed pre‐Clovis assemblages have closer affinities to known natural assemblages than to cultural assemblages. Our results suggest that these sites provide no evidence for a pre‐Clovis occupation in the Americas. © 2004 Wiley Periodicals, Inc.     

What have natural and human changes wrought on the foraminifera of San Francisco Bay late Quaternary estuaries?

In this study we compare the foraminifera of modern South San Francisco Bay with fossils from sediments of a previous estuary at 125 ka to provide a basis for interpreting the impact of natural and human change on the benthic ecosystem. All the species found in the Pleistocene sediments of this study are estuarine and/or shallow-water species occurring commonly in San Francisco Bay today, except for the introduced foraminifer Trochammina hadai, a native of Japan that was not found in samples taken in San Francisco Bay before 1983. The biodiversity and species composition of the fossil and modern assemblages before the introduction of T. hadai are nearly identical, suggesting that the environmental and physical changes in the 125,000-year-old and modern estuaries have not had a significant effect on the meiofauna of the Bay. In contrast, modern anthropogenic change in the form of species introductions has impacted the modern foraminiferal assemblage: T. hadai began to dominate the modern assemblage a decade after its introduction. Similar to the recorded impacts of introductions of marine metazoan invertebrate species, the dominance of T. hadai changed species proportions in the post-1980s foraminiferal assemblage, however no known extinctions in the native foraminiferal fauna occurred.     

Exploring Paleogeographic Conditions at Two Paleolithic Sites in Navarino,Southwest Greece,Dated by Optically Stimulated Luminescence

In this paper, we employed optically stimulated luminescence (OSL) dating of sediments from two archaeological sites located in Navarino, Messenia, southwestern Greece, to deduce a chronology for the archaeological sites. Archaeological surveys identified two Paleolithic sites on fossilized coastal dunes. Chipped stone tool assemblages were identified eroding out of paleosols developed in the dunes. The assemblage from one site lacked distinct typological features and hence it was difficult to assign to a chronological period. The lithic assemblage from the other site contained artifacts that typologically can be assigned to the Levallois‐Mousterian. Previous efforts to date the artifact‐bearing sediments at these sites were unsuccessful. Using newer OSL dating methods (i.e., the Single‐Aliquot‐Regenerated Dose protocol and thermally transferred‐OSL[TT‐OSL]), we attempted to construct a chronological framework for Late Pleistocene human activity in the southwest Peloponnese. The revised OSL chronology for the first site is 28 ± 5 ka, while a luminescence age of 8 ± 1 ka for the second site only represents a later deflation event. Within the framework of Quaternary environmental change, the location of Paleolithic sites relative to the coast would have changed during the course of the Pleistocene. As a result, Paleolithic exploitation strategies would have been strongly influenced by the changing coastal geomorphology, encouraging hominids to adapt to new distributions of resources. OSL dating of the archaeological sites allowed us to connect traces of hominid activity with climatic stadials/interstadials of the later Pleistocene derived from existing relative sea‐level curves. Ultimately, these data permitted the reconstruction of regional Late Pleistocene paleogeography. © 2012 Wiley Periodicals, Inc.     

Bearing up well? Understanding the past,present and future of Australia's koalas

The modern Koala Phascolarctos cinereus is the last surviving member of a once diverse family Phascolarctidae (Marsupialia, Phascolarctomorphia). Nine genera and at least 16 species of koala are known. Late Oligocene sediments of central Australia record the oldest fossils and highest species diversity. Five species are known from the early to middle Miocene rainforest assemblages of the Riversleigh World Heritage Area, Queensland. With the onset of dryer conditions after the middle Miocene climatic optimum (~ 16 Ma), rainforest habitats contracted resulting in the apparent extinction of three koala lineages (Litokoala, Nimiokoala, Priscakoala). Phascolarctos first appears in the fossil record during the Pliocene and the modern species around 350 ka. Despite a dramatic decline in taxonomic diversity to a single extant species, the fossil record indicates that at most only three koala species coexisted in any given faunal assemblage throughout their 24 million year history. Within these assemblages, the vast majority of extinct koalas are extremely rare (some known from only a single specimen) which may reflect a general rarity within their palaeohabitats compared with the modern species which is represented by an estimated 400,000 individuals spread over most of eastern mainland Australia. Be that as it may, P. cinereus, although once geographically more widespread, occurring for example in Western Australia in the Pleistocene, underwent significant range contractions and localized population extinctions during the stressful climatic conditions of the late Pleistocene and more recently through human-induced habitat destruction. Combined with threats of disease, reduced genetic diversity and climate change, the survival of this iconic Australian marsupial is arguably a cause for concern.     

New Stigmaphronidae and Megaspilidae (Hymenoptera: Ceraphronoidea) from Canadian Cretaceous amber

Three new species within the stigmaphronid genus TagsmiphronEngel and Grimaldi, 2009, and one new species within the megaspilid genus ConostigmusDahlbom, 1858 are described from Upper Cretaceous (Campanian) amber originating at the Grassy Lake locality in Alberta, Canada. New taxa include Tagsmiphron spiculum sp. nov., Tagsmiphron leucki sp. nov., Tagsmiphron exitorum sp. nov., and Conostigmus cavannus sp. nov. The new Conostigmus species is a rare discovery. It is the third megaspilid species to be found in Cretaceous amber, with the two specimens described herein effectively doubling the number of known Mesozoic exemplars for the family. We provide the first comprehensive report of known Ceraphronoidea within Canadian amber, and contrast this against other Cretaceous amber assemblages, discussing the potential palaeobiogeographic and palaeoenvironmental implications of the Canadian amber assemblage.     

Plio-Pleistocene mammals from Mille-Logya,Ethiopia, and the post-Hadar faunal change

We describe the non-primate mammalian fauna from the late Pliocene to earliest Pleistocene deposits of Mille-Logya in the Lower Awash Valley, Ethiopia, dated to c. 2.9–2.4 Ma, and divided into three successive units: Gafura, Seraitu, and Uraitele. We identify 41 mammalian taxa (including rodents), the most diverse group being the Bovidae, with 17 taxa. While the Gafura assemblage still resembles those from the earlier Hadar Formation, the younger Seraitu assemblage documents a major turnover. While there is little change in the species present across this interval, the relative abundances of various taxa change dramatically, with suids being largely replaced by open-country bovids (Alcelaphini and Antilopini). We interpret this faunal change as reflective of an environmental shift, contemporaneous with the replacement of Australopithecus afarensis by Homo in the area.     

The stratigraphical significance of deer species in the cromer forest-bed formation

Deer species provide a valuable biostratigraphical tool through Cromer Forest-bed times, due particularly to species turnover between the Early and early Middle Pleistocene. This study is based on the reidentification of 348 fossil antlers. The provenance of most large mammal fossils from the CF-bF, collected over 150 years, was recorded only by the nearest coastal village. None the less, analysis of the cervid taxa by these ‘localities’ reveals interesting patterns. The fauna of the West Runton Freshwater Bed is of early Middle Pleistocene complexion, and that from the foreshore at East Runton is of Early Pleistocene. Pure or nearly pure early Middle Pleistocene assemblages also occur at Kessingland-Pakefield and at Trimingham. At Overstrand, Sidestrand, Mundesley, Bacton and Happisburgh, there is a mixture of Early and early Middle Pleistocene elements. Analysis of Savin's data shows that fossils of earlier species were generally found further down the beach than those of later ones. Late nineteenth century geological surveys, made when bone collecting was at its peak, give additional information about fossil horizons, which in several cases can be related to modern stratigraphical units. Most Early Pleistocene large-mammal bones came from Pastonian conglomerates, in contrast to small-mammal and molluscan assemblages mostly extracted from Pre-Pastonian Crag. However, the diversity of Early Pleistocene CF-bF cervid species in comparison with continental faunas, and their pattern of distribution between sites, suggests they may span more than one chronostratigraphic stage. Early Middle Pleistocene assemblages came from strata now referred to the Cromerian, and the differing proportions of taxa between sites provide limited evidence of time-transgression.     

Permian megaspores from Kuraloi Area,Ib-River coalfield,Mahanadi Basin,Orissa

Heterogenous megaspores recovered from the Upper Permian sediments of Kuraloi Area, Ib-River coalfield, have been systematically described. The megaspore assemblage includes seven genera with seventeen species viz. Bokarosporites rotundus, Banksisporites indicus, B. utkalensis, Biharisporites spinosus, Biharisporites sp.A, Biharisporites sp.B, Jhariatriletes baculosus, J. srivastavae, J. damudicus, Manumisporites distinctus, Singhisporites baculatus, S. radialis, S. surangei, Ramispinatispora indica, R. nautiyalii and Ramispinatispora sp. Additionally, a new species of Ramispinatispora viz. R.mahanadiensis has been identified. Associated spore pollen assemblages are suggestive of a Late Permian age corresponding to Raniganj/Kamthi Formation. Raniganj megaspores are known from Damodar and Godavari Basins. The newly recorded megaspore assemblage from the Ib-River coalfield is comparable to that of Raniganj/Kamthi Formation of Godavari Graben. The diversity in exosporium ornamentations indicates structural diversity. Moreover, their prolific presence ultimately points towards occurrence of heterosporous source vegetation in the area.     

Pleistocene geochronologies for fluvial sedimentary sequences: an archaeologist's perspective

Current models of Pleistocene fluvial system development and dynamics are assessed from the perspective of European Lower and Middle Palaeolithic stone tool assemblages recovered from fluvial secondary contexts. Fluvial activity is reviewed both in terms of Milankovitch‐scale processes across the glacial/interglacial cycles of the Middle and Late Pleistocene, and in response to sub‐Milankovitch scale, high‐frequency, low‐magnitude climatic oscillations. The chronological magnitude of individual phases of fluvial activity is explored in terms of radiocarbon‐dated sequences from the Late Glacial and early Holocene periods. It is apparent that fluvial activity is associated with periods of climatic transition, both high and low magnitude, although system response is far more universal in the case of the high magnitude glacial/interglacial transitions. Current geochronological tools do not permit the development of high‐resolution sequences for Middle Pleistocene sediments, while localised erosion and variable system responses do not facilitate direct comparison with the ice core records. However, Late Glacial and early Holocene sequences indicate that individual fluvial activity phases are relatively brief in duration (e.g. 10² and 10³ yr). From an archaeological perspective, secondary context assemblages can only be interpreted in terms of a floating geochronology, although the data also permit a reinvestigation of the problems of artefact reworking. Copyright © 2005 John Wiley & Sons, Ltd.     

Taxonomic composition,paleoecology and biostratigraphy of Late Cretaceous diatoms from Devon Island,Nunavut, Canadian High Arctic

Upper Cretaceous sediments of the Kanguk Formation exposed in Eidsbotn and Viks Fiord grabens on Devon Island, Nunavut, Canadian High Arctic, yielded 91 fossil marine diatom species and varieties (including indeterminate taxa), representing 41 genera. Excellent preservation of the assemblages was aided by shallow burial, protection in downfaulted linear grabens, and the presence of abundant volcanic material. Planktonic species and resting spores comprise nearly 70% of the diatom assemblage, and provided abundant food resources for the Late Cretaceous Arctic ecosystem. Deposition of the approximately 225 m-thick stratigraphic sequence was predominantly in a shallow marine neritic setting, with an upward progression to interbedded terrestrial deposits of the Expedition Fiord Formation, reflecting a regression and eventual persistence of terrestrial facies into the Early Cenozoic. The Kanguk Formation is widespread across the Canadian Arctic, and diatom biostratigraphy indicates a Santonian–Campanian age for the sequences reported herein, based on the presence of Gladius antiquus in the lowermost strata and occurrence of Costopyxis antiqua throughout the succession. However, Amblypyrgus sp. A and Archepyrgus sp. aff. A. melosiroides, encountered in the lower part of the succession, are known exclusively from the Lower Cretaceous. This may suggest a slightly older age. New information on shallow shelf diatom assemblages from this study is compared to reports on two other Late Cretacous Arctic diatom assemblages. These three sites represent an environmental transect from shallow to distal shelf settings and into the oceanic realm.     

Late Pleistocene paleoecology of arctic ground squirrel (Urocitellus parryii) caches and nests from Interior Alaska's mammoth steppe ecosystem, USA

Botanical analyses of fossil and modern arctic ground squirrel (Urocitellus parryii) caches and nests have been used to reconstruct the past vegetation from some parts of Beringia, but such archives are understudied in Alaska. Five modern and four fossil samples from arctic ground squirrel caches and nests provide information on late Pleistocene vegetation in Eastern Beringia. Modern arctic ground squirrel caches from Alaska's arctic tundra were dominated by willow and grass leaves and grass seeds and bearberries, which were widespread in the local vegetation as confirmed by vegetation surveys. Late Pleistocene caches from Interior Alaska were primarily composed of steppe and dry tundra graminoid and herb seeds. Graminoid cuticle analysis of fossil leaves identified Calamagrostis canadensis, Koeleria sp. and Carex albonigra as being common in the fossil samples. Stable carbon isotopes analysis of these graminoid specimens indicated that plants using the C₃ photosynthetic pathways were present and functioning with medium to high water-use efficiency. Fossil plant taxa and environments from ground squirrel caches in Alaska are similar to other macrofossil assemblages from the Yukon Territory, which supports the existence of a widespread mammoth steppe ecosystem type in Eastern Beringia that persisted throughout much of the late Pleistocene.