On some fabaceous fruits from the Siwalik sediments (Middle Miocene-Lower Pleistocene) of Eastern Himalaya

Four legume fruits (Fabaceae) from the Siwalik sediments (middle Miocene to lower Pleistocene) of Darjeeling and Arunachal Pradesh of Eastern Himalaya are described here. One fossil fruit, Dalbergia prelatifolia sp. nov., is recovered from the lower part of the Siwalik succession (Gish Clay Formation of Sevok Group; middle to upper Miocene) of Darjeeling foothills. Mastertia neoassamica sp. nov. and Acacia miocatechuoides sp. nov. are collected from the lower part of the Siwalik succession (Dafla Formation; middle Miocene to upper Miocene), while Pongamia kathgodamensis Prasad is recovered from the upper part of the Siwalik succession (Kimin Formation; upper Pliocenelower Pleistocene) of Arunachal sub Himalaya. Their presence indicates a warm and humid tropical environment in the region during the Siwalik sedimentation.     

云南下侏罗统禄丰组恐龙足迹的发现和张北足迹属新观察

记述了云南省禄丰县腰站乡下侏罗统禄丰组2个恐龙足迹,命名了棋盘张北足迹（Changpeipus pareschequier ichnosp. nov.）一新种。这是在禄丰组中首次发现恐龙足迹化石,重新观察并修订了张北足迹一属的属征。滦平张北足迹（Changpeipus luanpingeris）应为石炭张北足迹（Changpeipus carbonicus）的亚成年个体,为同物异名。此前被认为是石炭张北足迹（Changpeipus carbonicus）前足迹的IVPP V2472 2a,应为其未成年体的后足迹。张北足迹与卡岩塔足迹（Kayentapus）类似。综合种种特征,棋盘张北足迹（新种）的造迹恐龙很可能为禄丰组的腔骨龙类（Coelophysoidea）     

An unusual theropod track assemblage from the Cretaceous of the Zhucheng area, Shandong Province, China

More than 125 footprints of theropods from the Cretaceous Longwangzhuang Formation have been mapped in a preliminary study at a site in the Zhucheng region of China. The tracks represent at least three morphotypes. The largest morphotype is a large theropod (footprint length ∼30 cm) represented by a single trackway and an isolated natural cast. At least 10 trackways assigned to the new ichnospecies Corpulentapus lilasia represent a medium-sized biped (footprint length ∼13 cm) with very short, wide, robust, ‘tulip-shaped’ tracks and long steps (∼5 × footprint length), and a short central digit (III) indicating weak mesaxony. Corpulentapus trackways are narrow and theropod-like even though track morphology is convergent with the footprints of some ornithopods. The third morphotype, made by a medium-sized grallatorid track maker (ichnogenus Paragrallator), is about the same size (∼13 cm) as the robust morphotype, but far more elongate and gracile, with an elongate central digit (III) indicating strong mesaxony. This ichnotaxon requires detailed comparison with Grallator sensu stricto. The contrast in morphology between the two common morphotypes is striking and demonstrates that two distinct medium-sized taxa of presumed theropod affinity frequented the same habitat in significant numbers.     

Gastrochaenolites ornatus Kelly and Bromley and other trace fossils on a Middle Miocene omission surface from the Aures Massif,Algeria

Trace fossils provide detailed palaeoenvironmental and palaeoecological information of both ancient and modern sedimentary systems. During middle Miocene times the Aures Massif located in the northeastern part of Algeria, was affected by, at least, one marine transgression. The latter led to the installation of a carbonate platform, which is placed, for the first time, in a Mediterranean context. In the Rhassira basin, the Middle Miocene marine succession is characterised by carbonate platform deposits dominated by rhodolith beds, typical of those known throughout the Mediterranean area. This succession can be divided into many units separated by discontinuities interpreted here as omission surfaces. The Djebel Arhane section shows two omission surfaces characterised by a pre-omission suite (firmground) represented by Balanoglossites burrows for the first surface and Gastrochaenolites ornatus burrows/borings for the second one, and an omission suite (hardground) as evidenced by the bioerosive structures Trypanites and Caulostrepsis, in both surfaces, respectively. Gastrochaenolites ornatus traces were formed and preserved in firm, compact, semi-lithified and fine-grained substrates (firm- to hardground), indicating the Glossifungites ichnofacies. They show bioglyphs which have been formed during contraction of the posterior adductor muscles. These suggest that their tracemakers were represented by suspension-feeding bivalves, most probably Pholadidae or Mytilidae, which rotated during penetration. The fill of these traces is composed of marine deposits related to a transgressive lag. The omission suite is divided into two ichnocoenoses: (i) pre-lithification burrows/borings, and (ii) post-lithification borings. This is the first report of the ichnotaxon G. ornatus from Algeria.     

A New Fossil Termite(Isoptera,Stolotermitidae,Stolotermes)from the Early Miocene of Otago,New Zealand

<正>The forewing of a termite from Early Miocene lake sediments in Otago,southern New Zealand is figured and described.It exhibits the generic characters of the damp-wood termite Stolotermes Hagen,but differs from forewings of the known species in size and venation pattern and is described as Stolotermes kupe sp.nov.S.kupe represents the first confident record of fossil Stolotermitidae and extends the fossil record of the family back to the Early Miocene.It also is the first direct evidence of fossil Isoptera from New Zealand,though silicified termite faecal pellets,referable to Kalotermes brauni,have been previously described.S.kupe indicates that Stolotermitidae has been present in the Australasian region since at least the Early Miocene.     

First record of the beetle family Lasiosynidae (Insecta: Coleoptera) from the Lower Cretaceous of China

A new genus, Cretasyne gen. nov., in the family Lasiosynidae, with two species C. lata sp. nov. and C. longa sp. nov., is described from the Lower Cretaceous Yixian Formation of Inner Mongolia, China. These fossils are the first record of the family from the Yixian Formation and provide significant information on lasiosynid morphology and diversity. The fossil record of Lasiosynidae is also reviewed.     

记广东南雄盆地上白垩统主田组鸟脚类足迹及中国已命名的大型鸟脚类足迹的属级新观察

记述了广东省南雄盆地上白垩统主田组3组恐龙行迹,由杨梅坑足迹点的足迹命名了南雄鸭嘴龙足迹（Hadrosauropodus nanxiongensis ichnosp. nov.）一新种;古市足迹点的足迹归入鸭嘴龙足迹一未定种（Hadrosauropodus isp.）。这是中国鸭嘴龙足迹属的首次描述,将该足迹的分布从北美拓展至中国,并为南雄盆地存在大型的鸭嘴龙提供了新的化石证据。还整理了中国已描述的大型鸟脚类足迹：中国足迹（Sinoichnites）被定为非正式的塑模标本;扬子足印（Yangtzepus）被归入兽脚类足迹;禽龙足迹（Iguanodonopus）被认为是裸名,归入禽龙足迹（Iguanodontipus）;嘉荫足迹（Jiayinosauropus）增加了新材料并得以具体描述;内乡足迹（Neixiang footprint）得以重新描述。     

A new tribe of Galerucinae leaf beetle (Insecta: Coleoptera: Chrysomelidae) from the Upper Cretaceous Taimyr amber

The new tribe Taimyralticini trib. nov., new genus Taimyraltica gen. nov. and new species Taimyraltica calcarata sp. nov. (Coleoptera: Chrysomelidae: Galerucinae) are described from Upper Cretaceous (Santonian) Taimyr amber of northern Siberia (Yantardakh locality). The new genus shares characters with genera of the tribe Alticini (small size, transverse antebasal groove on pronotum, regular punctate-striate elytra) and the Galerucini (narrow, non-swollen metafemora). A possible apomorphy of the new genus and tribe is a large and acute metatibial spur, erect and oriented transversely to the longitudinal axis of tibia. This character state is unknown in recent and fossil genera of Galerucinae. Early evolution and divergence of leaf beetles are discussed, and a Jurassic and Early Cretaceous subfamily radiation is proposed. Fossil galerucines are reviewed. The oldest Phyllotreta is recorded from the lower Miocene of Izarra (Spain). Our findings show an extreme rarity of Phytophaga in Taimyr amber from Yantardakh. Absence of Phytophaga was recorded earlier at Obeschayuschiy (Santonian-Campanian of the Magadan Region) and compared with their much larger share in Khetana (north of Khabarovsk Krai); we find nearly the same differences between Taimyr amber from Yantardakh and Canadian amber. In Obeschayuschiy this appears due to the Mesophytic character of the flora. It is assumed that the Santonian amber gymnosperm forest of the Khatanga Basin had the same character.     

New cirripedes (Crustacea,Thoracica) from the Jurassic and Cretaceous of the United Kingdom

Extensive new material of scalpelliform cirripedes, including articulated individuals, from the Late Jurassic (Tithonian) Kimmeridge Clay of Dorset, southern England, is described and used as a basis for taxonomic revision of the family Zeugmatolepadidae. Two subfamilies, the Zeugmatolepadinae and Martillepadinae nov., are established and two new genera (Martillepas, Concinnalepas) in the latter subfamily are described from the Upper Jurassic and one (Icenilepas) from the Upper Cretaceous Chalk. Material from the Kimmeridge Clay of Dorset also includes the oldest fossil representative of the family Pollicipedidae for which a new genus and species, Etcheslepas durotrigensis, are erected, and the oldest calanticid, Cretiscalpellum sp. nov. Jurassic taxa established by T.H. Withers in 1928, on the basis of sparse material, are redescribed, and referred to the new genera as Martillepas ovalis, M. costata, M. hollisi, Concinnalepas concinna and Etcheslepas fragilis.     

Biostratigraphy and depositional environment of the Miocene limestone bed of Baripada,Mayurbhanj district,Odisha: Foraminiferal,sedimentological and bulk organic geochemical evidences

Baripada Marine Beds (BMB) have been studied extensively in terms of its mega fossil content. However, not much has been discussed about the foraminiferal content and the organic matter assemblage in these beds. The fossiliferous sequence of BMB consists of sandstone, shale and limestone units. The present study is persued on the limestone unit of BMB. Foraminiferal, thin section and rock-eval pyrolysis studies were performed on the 33 surface samples collected from five sections of Jamdapal and Mukurmatia region along the Budhabalang river bank. Foraminifera are less abundant in the samples of Jamdapal, whereas Mukurmatia is comparatively rich. First appearance datum of Globorotalia menardii[ranges since middle Miocene (12.6 Ma); FAD at planktic foraminiferal zone N12] and last appearance datum of Neogloboquadrina continuosa [ranges between early Miocene (23.2 Ma) to late Miocene (8.3 Ma); FAD at planktic foraminiferal zone N4B and LAD at N16] together suggest that the limestone unit was deposited in between 12.6 to 8.3 Ma within the upper Miocene. Also, the association of shallow water benthic foraminifera (Species of Ammonia, Asterorotalia, Bolivina, Buliminella, Cibicides, Challengerella, Criboelphidium, Cribononion, Elphidium, Hanzawaia, Nonionella) and planktic foraminifera (Globigerina falcoensis, Globigerina woodi, Globorotalia menardii, Neogloboquadrina continuosa) together with oyster bank and shark teeth suggest deposition of limestone within well oxygenated, tropical, shallow water, open marine condition (within 40m water depth). Lithological and thin section studies together with global sea level fluctuation history advocate that upper Miocene marine transgression promoted the formation of this unit. Bulk organic geochemical data obtained by the rock-eval pyrolysis studies on selected samples indicate a low total organic carbon (TOC), with low hydrogen index (HI), high oxygen index (OI). The organic facies is characterised by type-IV kerogen with major contribution from near shore terrestrial plants. This also suggests deposition in shallow, oxygenated environments that did not promote significant accumulation and preservation of organic content in sediments.     

A new fossil species of snakeflies (Raphidioptera: Mesoraphidiidae) from the Late Cretaceous of Kazakhstan,with notes on Turonian Neuropterida

Alloraphidia kyzylzharica sp. nov. is described from the Late Cretaceous (early Turonian) locality at Kyzylzhar, southern Kazakhstan. It is very similar to A. dorfi, but easily distinguished from it by wing shape and venation. We restrict the genus Alloraphidia to these two species. Ascalapharia raphidiformis is considered a member of Baissopteridae, sit. nov. Turonian Neuropterida are mainly characterized by a mixture of specialized genera of extinct families, and genera (sometimes modern) of highly advanced taxa, reflecting a sequence of global mid-Cretaceous crisis of non-marine biocoenoses.     

First fossil evidence of Connaraceae R. Br. from Indian Cenozoic and its phytogeographical significance

Fossil leaflet impression described here as a new species Rourea miocaudata sp. nov., showing close resemblance with the modern leaflets of Rourea caudata Planch. (Connaraceae R. Br.), has been recorded from the lower part of the Siwalik sediments (Dafla Formation, middle–upper Miocene) exposed at the road-cutting section of Pinjoli area in West Kameng district, Arunachal Pradesh. The important morphological characters of the fossil are its narrow elliptic leaflet, cuneate base, long caudate apex, entire margin, eucamptodromous to brochidodromous secondary veins, presence of intersecondary veins, percurrent and reticulate tertiary veins and orthogonally reticulate quaternary veins. This is the first authentic record of the occurrence of leaflet comparable to R. caudata of Connaraceae from the Cenozoic sediments of India and abroad. At present R. caudata does not grow in India and is restricted only in southeast Asia especially in China and Myanmar. This taxon probably migrated to these southeast Asian regions after lower Siwalik sedimentation (middle–upper Miocene) due to climatic change caused by post-Miocene orogenic movement of the Himalaya. The recovery of this species and other earlier-described evergreen taxa from the same formation, suggests the existence of a tropical, warm and humid climatic conditions during the depositional period.     

New lacewings (Insecta: Neuroptera) from the Lower Cretaceous Wealden supergroup of Southern England

Eight new genera and thirteen new species of lacewings (Neuroptera) are described from the Lower Cretaceous Wealden Supergroup, Weald Sub-basin: Principiala rudgwickensis sp. nov. (Ithonidae), Stenomylina medialis gen. et sp. nov., Protosmylina bifasciata gen. et sp. nov. (Osmylidae), Cretapsychops corami gen. et sp. nov., Micropsychops parallelus gen. et sp. nov., Valdipsychops logunovi gen. et sp. nov., V. brigidae gen. et sp. nov., V. maculosus gen. et sp. nov., V. proudlovei gen. et sp. nov., V. minimus gen. et sp. nov., Psychopsites rolandi gen. et sp. nov. (Psychopsidae), Actinophlebioides valdensis gen. et sp. nov. and Protohemerobius perexiguus gen. et sp. nov. (Neuroptera familia incertae sedis). Psychopsidae are shown to dominate the neuropteran fauna, while the presence of Brongniartiellidae has not been confirmed in the Wealden, due to previously thought representatives of this family being in fact psychopsids. This is the first fossil record of Ithonidae and Osmylidae in Britain. The range of Kalligrammatidae has been extended in the Cretaceous of Britain from the mid-Valanginian to early Barremian.     

A new record and a new ichnospecies of Arachnostega from the Middle Miocene of Egypt

The ichnogenus Arachnostega Bertling, 1992 is recorded for the first time from a bioeroded, coral-rich, Middle Miocene limestone bed in the middle Siwa Escarpment Member (Marmarica Formation) at Siwa Oasis, northern Western Desert of Egypt. These burrowing traces are preserved on the surface of a few internal moulds of bivalves and gastropods. Until now, the ichnogenus Arachnostega included a single ichnospecies, A. gastrochaenae Bertling, 1992. In the material studied herein, two ichnospecies are identified, described, illustrated and compared with other, similar traces. One of these is a new ichnospecies of Arachnostega; this is here named A. siwaensis ichnosp. nov. It is distinguished from A. gastrochaenae in the possession of tunnels that usually branch in a V-pattern; intervals of ramification are longest in the main branch and decrease in second- and third-order branches. Arachnostega traces were produced in a warm, low-energy, shallow-marine (<10?m water depth) environment. Polychaetes are the most likely producers of marine Arachnostega, which are commonly occurred in the same studied bed. This new occurrence is significant in extending the known stratigraphical and geographical ranges of Arachnostega into the Middle Miocene of Egypt.     

A new family of scorpionflies (Insecta; Mecoptera) from the Lower Cretaceous of England

Seven specimens of fossil scorpionflies (Mecoptera) not assignable to any known family were discovered in the Wealden Supergroup (Lower Cretaceous) of southern England. They were found at Rudgwick Brickworks, West Sussex and Smokejacks Brickworks, Surrey and came from the Upper Weald Clay Formation, dated as Barremian (∼129.4–125 Ma). A new family – Englathaumatidae fam. nov., new genus – Englathauma gen. nov. and two new species E. crabbi sp. nov. and E. mellishae sp. nov. are described. A discussion of systematic position of these new taxa within the order Mecoptera is given. Englathaumatidae fam. nov. has been a nomen nudum since the year 2002, due to the first author's untimely death.     

Fossil neocheilostomine bryozoans from the tertiary rocks of western Kachchh, Gujarat

Paper describes six species of neocheilostomine bryozoans occurring in the Middle Eocene-Early Miocene rocks of Western Kachchh, Gujarat, India. Taxa belong to the families Quadricellariidae Gordon, 1984, Vinculariidae Busk, 1852 and Poricellariidae, Harmer, 1926. Of these three species are new. These are Vincularia feddeni nov sp. Poricellaria waioriensis nov sp. and Poricellaria sakurkari nov sp. Two species, Nellia tenella (Lamarck, 1816) and Vincularia kutchensis (Tewari & Srivastava, 1967) already known from this region, are further reported here. Poricellaria complicata (Reuss, 1869)? is reported for the first time and indicates oldest record from India so far. Phylogenetic analysis using PAST programme indicates that Poricellariids and Vinculariids have not evolved from the same ancestor Nellia tenella (Lamarck, 1816)     

New genera and species of bark-gnawing beetles (Coleoptera: Trogossitidae) from the Yixian Formation (Lower Cretaceous) of Western Liaoning,China

Two new genera from the Lower Cretaceous Yixian Formation of Beipiao City, Liaoning Province, northeastern China are described and illustrated. Paracretocateres gen. nov. (type species P. bellus, sp. nov.) and Yixianteres gen. nov. (type species Y. beipiaoensis sp. nov.) are assigned to the subfamily Lophocaterinae based on the state of the procoxae which are not projecting, the transverse procoxal cavities, and the exposed protochantins. The new taxa broaden the diversity of this family and provide further morphological characters for phylogenetic studies of Trogossitidae. They also provide evidence that lophocaterines well-diversified prior to the trogossitines and their biologies may have been similar to extant forms.     

Revision of the Cretaceous echinoids Heterodiadema and Trochodiadema,and description of two new genera from the Middle East and Arabian Peninsula

The Cretaceous hetero diadematid echinoids Heterodiadema Cotteau, 1862 and Trochodiadema de Loriol, 1900 are revised and provided with amended diagnoses. Of six species previously referred to Heterodiadema, only three are now retained in this genus, namely Heterodiadema libycum (Desor in Agassiz & Desor, 1846), Heterodiadema ouremense de Loriol, 1884 and Heterodiadema buhaysensis Smith, 1995. Similarly, seven species were previously assigned to Trochodiadema, but here only Trochodiadema abramense de Loriol, 1900 is left in this genus. Based on material previously referred to Trochodiadema, two new genera and one new species are established: Loriolidiadema gen. nov. from the Lower Cretaceous of the Middle East, and Arabicodiadema gen. nov. and Arabicodiadema alii sp. nov from the Albian and Cenomanian of the Arabian Peninsula. Trochodiadema isidis Fourtau, 1921 is transferred to Aegyptiaris Abdelhamid & Abdelghany, 2015. New morphological data on primary and secondary spines and perignathic girdle are presented; these improve our understanding of the evolution and relationships of the family Heterodiadematidae Smith & Wright, 1993.