2010—2020及影响因素分析年青岛海滩剖面稳定性

海滩的稳定性及其演变是砂质海滩研究的重要内容。在全球砂质海滩普遍遭受侵蚀的背景下,分析部分稳定海滩的特征和原因可以为海滩维护和科学管理提供重要参考。本文基于近 11 年 （2010—2020 年） 青岛第一、第三和石老人海水浴场共 8 条典型剖面的实地监测高程数据,对海滩滩肩宽度和单宽体积进行计算对比,研究表明,除石老人海水浴场一个剖面变化较大外,其他剖面的滩肩宽度只变化了 0~14 m,是 11 年前的 0%~23.3%;单宽体积变化了 8.5~46.5 m3,是 11 年前的1.0%~9.4%;海滩剖面垂向扰动范围一般在 30~60 cm 范围内。海滩存在局部小尺度淤积或侵蚀,剖面形态总体较为稳定。海平面变化、台风和风暴潮等自然因素是海岸轻微侵蚀或调整的重要影响因素,但对海滩的优化管理 （包括定期监测海滩、适度展开养护、制定完善的管理政策和加强执行力度等人为因素） 可以应对这些海滩变化,并且已成为近年来该研究海滩保持稳定的关键因素。     

Mapping beach morphodynamics remotely: A novel application tested on South African sandy shores

Sandy beaches have been identified as threatened ecosystems but despite the need to conserve them, they have been generally overlooked. Systematic conservation planning (SCP) has emerged as an efficient method of selecting areas for conservation priority. However, SCP analyses require digital shapefiles of habitat and species diversity. Mapping these attributes for beaches from field data can take years and requires exhaustive resources. This study thus sought to derive a methodology to classify and map beach morphodynamic types from satellite imagery. Since beach morphodynamics is a strong predictor of macrofauna diversity, they could be considered a good surrogate for mapping beach biodiversity. A dataset was generated for 45 microtidal beaches (of known morphodynamic type) by measuring or coding for several physical characteristics from imagery acquired from Google Earth. Conditional inference trees revealed beach width to be the only factor that significantly predicted beach morphodynamic type, giving four categories: dissipative, dissipative-intermediate, intermediate and reflective. The derived model was tested by using it to predict the morphodynamic type of 28 other beaches of known classification. Model performance was good (75% prediction accuracy) but misclassifications occurred at the three breaks between the four categories. For beaches around these breaks, consideration of surf zone characteristics in addition to beach width ameliorated the misclassifications. The final methodology yielded a 93% prediction accuracy of beach morphodynamic type. Overlaying other considerations on this classification scheme could provide additional value to the layer, such that it also describes species’ spatial patterns. These could include: biogeographic regions, estuarine versus sandy beaches and short versus long beaches. The classification scheme was applied to the South African shoreline as a case study. The distribution of the beach morphodynamic types was partly influenced by geography. Most of the long, dissipative beaches are found along the west coast of the country, the south coast beaches are mostly dissipative-intermediate, and the east coast beaches range from short, estuarine pocket and embayed beaches in the former Transkei (south east), to longer intermediate and reflective beaches in KwaZulu-Natal (in the north east). Once combined with the three biogeographic regions, and distinguishing between estuarine and sandy shores, the South African coast comprised 24 different beach types. Representing shorelines in this form opens up potential for numerous spatial analyses that can not only further our understanding of sandy beach ecology at large spatial scales but also aid in deriving conservation strategies for this threatened ecosystem.     

Assessing inter-beach differences in semi-terrestrial arthropod assemblages on Maltese pocket sandy beaches (Central Mediterranean)

The distinctiveness of macrofaunal assemblages on different sandy beaches in the Maltese Islands was previously suggested by different single-season studies. A multi-seasonal sampling programme using pitfall trapping was implemented on four Maltese beaches to test the occurrence of this phenomenon. A total of 29,302 individuals belonging to 191 species were collected over a 2-year period, during which the beaches were sampled once per calendar season. A total of 77 species were recorded from single Maltese beaches only, of which nine were psammophiles. Non-metric multidimensional scaling analyses of pitfall trap species-abundance data resulted in a weak separation pattern, with samples grouping mainly in terms of beach and island rather than in terms of season or year of sampling, No physical variable could conclusively explain these patterns. It is concluded that although operating on Maltese beaches, macrofaunal assemblage distinctiveness is weaker than originally thought and can be attributed to the presence/absence or abundance of just a few psammophilic species. It is postulated that this phenomenon may be related to the 'pocket beach' nature of Maltese beaches, where headlands on either side of the beach to a large extent prevent the occurrence of longshore currents, resulting in semi-isolation of the populations of psammophilic species. A large number of single-beach records reported in this study highlight the high degree of beta diversity and spatial heterogeneity of Maltese beaches, and the conservation importance of the individual beach macrofaunal assemblages.     

Measuring species richness on sandy beach transects: extrapolative estimators and their implications for sampling effort

Species richness is a measure that is fundamental to many studies in ecology, and it is particularly important on sandy beaches, where it underlies patterns described by the broadly accepted swash exclusion hypothesis. However, its estimation in practice is problematic. This has led ecologists in other fields to adopt extrapolative estimators of species richness, which project the total number of species present in a habitat by adjusting upward the number of species observed by an amount related to the number of rare species encountered in the samples. In so doing, the species richness can be estimated, with confidence intervals, at any level of sampling effort. Despite the availability and advantages of these methods, beach ecologists have continued to use the observed species richness as a point estimate of biodiversity for beaches. Here, we employ a Monte Carlo resampling approach over a range of routine transect designs used to sample sandy beaches, and evaluate the performance of seven non-parametric extrapolative estimators for species richness relative to that of the more conventionally used observed species richness. We find that the first-order Jackknife estimator (Jack 1) is the least biased, most accurate and most consistent across sites. Employing this estimator would allow accurate estimation of species richness on short (tens of metres) stretches of beach without exceeding the acceptable levels of sampling effort (4–5 m²). Spreading this effort evenly over three across-shore transects, each with a minimum of 13 equally spaced levels seems appropriately efficient. Although a greater number of research studies is required to ascertain the generality of these results beyond the beaches we sampled, we tentatively recommend the application of our results in biodiversity surveys on sandy beaches.     

Responses of talitrid amphipods to a gradient of recreational pressure caused by beach urbanization

Demands for public outdoor recreation and urbanization are a potential source of environmental disturbances, particularly on sandy beaches. These habitats have therefore experienced rapid degradation worldwide. Talitrid amphipods, which are common inhabitants of sandy beaches, are probably good indicators of such disturbances. In this study, we evaluated the responses of the two amphipods Talitrus saltator and Atlantorchestoidea brasiliensis to human pressure on Spanish and a Brazilian beach, hypothesizing that disregarding species differences, their population sizes will decline under urbanized conditions. The Barra da Tijuca (Brazil) and Levante (Spain) beaches have mixed landscapes with varying levels of development, and each has a protected area (MPA) adjacent to urbanized sites. Numbers of talitrids as well as visitors were compared among sites on each beach. Talitrids were most abundant at the protected sites. This was true of both beaches, even though temporal patterns of human activity were quite different, with Levante experiencing sharp declines in the number of visitors during its harsh winter. Urbanized sites did not experience high Talitrid densities even during a period of low human pressure. On this basis, T. saltator and A. brasiliensis were considered good indicators of urbanization and recreational activities on sandy beaches. MPAs were found to be very important for the successful conservation of the talitrid populations. A feasible method of managing human pressure and thereby conserving beach biodiversity therefore seems to be the establishment of beach MPAs.     

Effects of morphodynamic state on macrofauna community of exposed sandy beaches on Galician coast (NW Spain)

It is known that the fauna of the exposed sandy beaches are primarily controlled by physical variables; but how these variables operate along and across the beach still remains fairly under discussion. In our study, we sampled a range of exposed sandy beaches along the Northwest coast of Spain to determine the relationship between the principal physical variables of the beaches (including beach morphodynamic state), and the macrofaunal community. The fauna of these beaches comprise truly marine species along the intertidal zone as well as semi-terrestrial species in the upper and supratidal environments. These two groups respond in a different manner to the physical environment. The first group is directly controlled by the morphodynamic state of the beach, and variations in the physical environment; the second group is not clearly affected by these physical conditions. In this case, other variables such as food availability and the human uses of the upper limits of the beach seem to be more relevant in explaining the patterns observed in the macrofaunal community.     

Sandy beach ecosystems: key features, sampling issues, management challenges and climate change impacts

Escalating pressures caused by the combined effects of population growth, demographic shifts, economic development and global climate change pose unprecedented threats to sandy beach ecosystems worldwide. Conservation of beaches as functional ecosystems and protection of their unique biodiversity requires management interventions that not only mitigate threats to physical properties of sandy shores, but also include ecological dimensions. Yet, beach management remains overwhelmingly focused on engineering interventions. Here we summarise the key outcomes of several workshops, held during the 2006 Sandy Beach Ecology Symposium in Vigo, Spain, that addressed issues of climate change, beach management and sampling methodology. Because efficient communication between managers and ecologists is critical, we summarise the salient features of sandy beaches as functional ecosystems in 50 'key statements'; these provide a succinct synopsis of the main structural and functional characteristics of these highly dynamic systems. Key outcomes of the workshops include a set of recommendations on designs and methods for sampling the benthic infaunal communities of beaches, the identification of the main ecological effects caused by direct and indirect human interventions, the predicted consequence of climate change for beach ecosystems, and priority areas for future research.     

The ecology of sandy beaches of the Benguela ecosystem

Sandy beaches constitute nearly 46 per cent of the coastline between the Cape of Good Hope and the Orange River along the west coast of South Africa. In addition, shores of mixed sand and rock make up a further 24 per cent although these are not considered here. Sandy beaches are therefore the dominant shore type along the coastline, and most are subject to high wave energy. There are two main ecological beach types along the study coastline: those that receive a high input of organic matter in the form of stranded kelp and those that do not. Neither type appears to support large stocks of surf-zone phytoplankton, but despite this, even beaches receiving no stranded kelp bear high standing stocks of infauna. This fact may be related to the location of the beaches alongside a highly productive upwelling region. Existing ecological information on sandy beaches along the Benguela coastline is reviewed and integrated to form a composite picture of present understanding of these beaches. The definition of a sandy beach includes not only the sandy intertidal zone but also the surf zone and sand dunes associated with it. Sandy beaches are characterized by the absence of attached primary producers, although in some parts of the world primary production by surf-zone phytoplankton has been found to be important. Secondary production by the infauna usually depends on matter imported into the system, except on beaches supporting important stocks of surf-zone phytoplankton. Imported organic matter is retained by beach sediments which act as a physical sieve, filtering large quantities of water with each wave and tide.     

Assessing the response of sandy-beach macrobenthos to recreation and the ecological status of the beach ecosystem at Liandao,China

Sandy beaches, prime recreational assets that attract more people than other types of shorelines, are the main assets underpinning many coastal economies and developments. However, the intense stressors to the ecology of sandy beaches caused by recreational activities are generally underestimated or ignored. This study was the first to emphasize the ecological characteristics of recreational beaches in China and assess the response of macrobenthos at Liandao Beach, a typical sandy beach in the central part of China. Using the Before/During/After/Control/Impact (BDACI) method, differences in multiple variables (species richness, mean density, diversity) between a heavily recreationally impacted transect (TrH/Impact) and a less-impacted transect (TrL/Control) in different impact periods (Before, During & After) were applied to assess the response of sandy-beach macrobenthos to recreation and tourism. Meanwhile, a comparison between the macrobenthos conditions of the studied beach and reference conditions calculating from historical data of similar beaches with little recreation and tourism disturbances was carried out. Furthermore, the quality control chart approach was used to detect in detail the macrobenthos stable state and yearly response to recreational activities. The results showed that recreation and tourism posed a serious threat to the beach ecosystem; macrobenthos on the studied beach were seriously disturbed by recreational activities, and their variations were highly negatively related to the human recreational level. After monthly recreational impacts, the macrobenthos decreased sharply and recovered with difficulty during 1 year. Over the long-term, the macrobenthos had a sharp decreasing trend, which indicated that the ecological environment of the beach was highly disturbed. Conservation measures for recreational beaches are very challenging because of the economic value and popularity of sandy beaches as recreational sites. However, practicable conservation measures could be implemented that prevent or slow ecological degradation. A number of management and conservation suggestions are put forward, such as the establishment of the ecological carrying capacity of Liandao Beach in terms of direct recreational use, creation of themed exhibitions related to the beach macrobenthos and beach ecosystem and dissemination of this knowledge to visitors, implementation of spatial zoning of beach use, visitor management and environmental education.     

Nematodes from wave-dominated sandy beaches: diversity,zonation patterns and testing of the isocommunities concept

Spatial patterns of nematode community structure from two geographically spaced intermediate, micro-tidal beaches (i.e. Mediterranean and Baltic) were investigated. Differences in the nematode assemblages were found to be significantly different and related to the morphodynamic characteristics of the studied zones (upper beach, swash/breakers and subtidal). Highest nematode densities and species diversities were recorded on the coarse-grained, more physically controlled, Italian beach in contrast to the more chemically controlled Polish beach. This is in contrast to the worldwide patterns of macrofaunal communities. As demonstrated by higher taxonomic distinctness measurements, upper beaches were found to harbour species from both the marine and terrestrial ecosystem and are considered to be important ecotones between these adjacent systems. The swash/breaker zones are characterised by the loss of distinctive species caused by the high water percolation in these zones. The concept of parallel ecological communities ‘isocommunities’ is only supported for the upper beach zones.     

Sandy beach macrofaunal communities on the western coast of Portugal – Is there a steady structure under similar exposed conditions?

The structure of macrofaunal communities at two similar exposed sandy beaches on the western coast of Portugal was monitored for approximately 18 months by sampling all the beach area, from the shoreline to the base of the dunes. The beaches’ physical environment, as well as community density and composition, seasonal variations and the potential relationships between biological data and environmental parameters were studied. The two beaches had similar exposure to wave action, but differed in terms of sediment grain size, extent of the intertidal area, sediment moisture content and, especially, in the potential food availability in the form of allochthonous debris. Differences were observed with regard to the communities’ structure, namely regarding composition and relative contribution of the dominant species. Seasonality, especially temperature variations, and the interaction between seasons and the beach zones (supralittoral vs. intertidal) also had a strong influence on communities: controlling dominant species’ density and the horizontal distribution of the dominant species, and promoting a differential utilization of the beach by several resident macrofaunal animals. The present study allows the identification of key species in exposed sandy beaches of western Portugal and demonstrates that a steady community structure does not persist in the similarly exposed conditions observed, which may be mainly a response to distinct detritus subsidies, combined with differences in sediment grain size, sediment moisture content and extent of the intertidal area.     

Relationships Between Community Structure of the Intertidal Macroinfauna and Sandy Beach Characteristics Along the Chilean Coast

Abstract. Eight sandy beaches were seasonally sampled along the coast of Chile, from ca. 21 to 42° S (about 3000 km) to study the relationship between community structure of the intertidal macroinfauna and beach characteristics. Sediment samples (0.1 m², 30 cm deep) were collected (July – September 1998 and December 1998 – January 1999) with plastic cylinders at 15 equally spaced levels along three replicated transects extending from above the drift line to the swash zone. The sediment was sieved through a 1 mm mesh and the organisms collected stored in 5 % formalin. To define beach types, Dean's parameter (Ω) was calculated from wave heights and periods, and fall velocity of sand particles from the swash zone. Crustaceans (mainly peracarids) were the most diverse group with 14 species, followed by polychaetes with 5 species. The talitrid amphipod Orchestoidea tuberculata , the cirolanid isopods Excirolana braziliensis and E. hirsuticauda and the anomuran decapod Emerita analoga were the most widely distributed and common species. Regression analyses between species richness, abundance and biomass of the whole macroinfauna versus sediment characteristics, beach face slopes and morphodynamic beach states showed no significant relationships. Thus, macroinfaunal community characteristics did not increase linearly from lower intermediate to higher intermediate or dissipative beach states as had been found before in Chile or in other coasts. A comparative analysis with data from sandy beaches of other world regions showed that the number of species inhabiting Chilean sandy beaches was generally lower, whereas total population abundances were generally higher compared with values reported elsewhere.     

Meiofauna as descriptor of tourism-induced changes at sandy beaches

Tourism has long been considered as a 'clean industry' with almost no negative effects on the environment. This study demonstrated, in two different coastal systems (Mediterranean and Baltic), that tourism related activities are particularly affecting the sandy beach meio- and nematofauna in the upper beach zone, the specific ecotone in which many meiofauna species from both the marine and the terrestrial environment congregate. Tourist upper beaches are characterized by a lower % total organic matter (%TOM), lower densities, lower diversities (absence of Insecta, Harpacticoida, Oligochaeta, terrestrial nematodes and marine Ironidae nematodes) and higher community stress compared to nearby non-tourist locations. The %TOM was found to be the single most important factor for the observed differences in meiofauna assemblage structure at tourist versus non-tourist beaches in both the Mediterranean and the Baltic region. The free-living nematode assemblages from tourist upper zones depart significantly from expectations based on random selections from the regional nematode species pool. Furthermore upper zone assemblages are characterised by a low species diversity consisting of taxonomically closely related nematode species with r-strategist features. Generally, faunal differences between tourist and non-tourist beaches are decreasing towards the lower beach zones.     

台风季节朱家尖岛砂、砾质岬湾海滩的不同沉积地貌动态变化

岬湾砂、砾质海岸是海岛、海岸带重要的旅游资源, 具有较高的社会经济和生态价值, 长期以来备受关注。本文以浙江朱家尖岛东岸的5个代表性岬湾海滩为例, 基于2019年台风季节早期、中期及晚期测量获得的海滩地形和沉积物数据, 结合海滩近岸的水动力数据, 分析了砂质海滩和砾石海滩这两种不同类型岬湾海滩的沉积地貌动态变化。结果表明, 朱家尖岛东岸5个海滩在台风季节出现了不同的体积变化, 大沙里、东沙、千沙、乌石塘和小乌石塘海滩的单宽体积变化分别为11.93m³·m^-1、-54.41m³·m^-1、-19.75m³·m^-1、2.19m³·m^-1和-1.96m³·m^-1。砾石滩较砂质海滩更为稳定, 无人类活动干扰的砂质海滩在台风季节侵蚀更少、变化更小。台风季节大沙里、东沙、千沙、乌石塘和小乌石塘海滩表层沉积物的平均粒径分别为2.47Փ、2.24Փ、2.64Փ、-5.96Փ和-6.03Փ, 粒径粗化和离岸输运是5个海滩表层沉积物在台风季节的主要表现, 砂质海滩的沉积物粒度特征变化比砾石海滩要大。沉积物粒径、台风强度及台风期间的主要波向与海滩走向之间的关系、海岸工程这3种因素都可能对海滩在台风季节的沉积地貌动态变化产生影响。本文研究结果可为台风季节的海滩管理提供参考。     

Can sandy beach molluscs show an endogenously controlled circatidal migrating behaviour? Hints from a swash rig experiment

Many macrofaunal species inhabiting exposed sandy beaches are dependent on the swash for their nutrition and migration and are highly adapted to the harsh physical conditions of the swash. The most important physical factors that determine the distribution and behaviour of swash related fauna, next to the swash itself, are sediment grain size and beach slope. Crucial swash parameters are swash period and swash velocity. Studying the influence of these factors on the animals in the field is often very difficult and it is almost impossible to identify which factor causes what effect. Crucial knowledge about the direct role of the swash itself is lacking. Therefore, a device that generates swash waves on an artificial beach under laboratory conditions was designed: the swash rig. In the swash rig, full and independent control on sediment grain size, beach slope, swash velocity and swash period is present. This will allow us to do a variety of experiments on the influence of each of these factors, independently or combined, on swash fauna. In one such experiment, Olivella semistriata – a dominant surfing gastropod on Ecuadorian sandy beaches – was placed in the swash rig during rising and falling tide and subjected – under constant conditions – to an equal wave regime. During falling tide, and in absence of any tidal cue, almost all specimens moved downshore, as they would in the field; hardly any specimens moved upshore. During upcoming tide, however, there was noticeable upward migration in the swash rig, and half of the runs showed a net upward migration. Contrary to the common understanding that the behaviour of sandy beach molluscs is entirely environmentally driven, this experiment hints at the presence of an endogenous circatidal clock, which is used to direct the tidal migration of the species.     

Community and Population Responses of the Macroinfauna to Physical Factors over a Range of Exposed Sandy Beaches in South-central Chile

Ten exposed sandy sites covering a range from reflective to dissipative beaches were sampled in south-central Chile to evaluate: (1) spatial changes in species richness, abundance and biomass of the intertidal macroinfauna in response to changes in mean grain size, beach face slope and beach type, and (2) spatial changes in abundance, biomass and body sizes of the most abundant species in response to changes in the physical factors. The number of species, abundance and biomass per beach in general decreased with increasing particle size and beach face slope (steeper beaches) and increased from reflective to dissipative conditions. The best fit for number of species was with Dean's parameter, a measure of beach type, whereas for abundance and biomass the best fits were found with particle size. The isopod Excirolana braziliensis and the anomuran Emerita analoga increased in abundance and biomass towards dissipative conditions, whereas Excirolana hirsuticauda showed the opposite trend in biomass and was significantly larger in beaches with steeper profiles. It is concluded that responses to changes in beach type are more pronounced at community level than within species populations.     

强潮环境下的海滩剖面沉积特征——以春季广西北海银滩为例

海滩地貌动力状态是海滩过程和陆海相互作用研究中的重要内容,但目前国内对北部湾强潮环境下的砂质海滩地貌与沉积的研究相对较少。对2010年3月采集的北海银滩剖面高程与沉积物变化的资料进行分析,探讨此类海滩响应外在驱动力作用的地貌动力状态特征,结果表明:1)北海银滩地貌结构简单,地貌状态属于消散型类型。2)银滩沉积物类型为砂,并以细砂、中砂为主,沉积物分选性较好;此外,沉积物中砂的百分含量和中值粒径自陆向海逐渐减小,而极细砂的百分含量则逐渐增加。3)北海银滩横向剖面沉积物可区分为后滨、滩面和内滨3个沉积单元。后滨沉积物在风的作用下以跃移质为主;滩面沉积物因波浪形成的冲流作用而具有双向运动的性质;内滨沉积物主要受控于波浪和潮流的联合驱动作用而以跃移质组分为主。