棉花冠层阴阳叶空间分布的日变化及其对光合作用的影响

在2007年7—8月棉花生长旺季,将冠层按高度分多层,通过简单的试验方法确定了棉冠内叶片丛聚指数(clumping index,Ω),以此区分计算出田间各层逐时阳叶(受光叶)和阴叶(被遮荫叶)叶面积指数的动态值。在8月23日用LI-6400R便携式光合测量系统分多层分别测定阴、阳叶单张叶片净光合作用的日变化,结合阴、阳叶面积权重,探讨利用Ω区分阴阳叶之后对整个冠层日总光合作用的影响。试验结果表明:(1)花铃期棉花冠层内Ω约为0.68;(2)利用Ω计算得到8月23日冠层日间(09:00—18:00)总净光合作用日平均值大约为20.3μmol·m-2·s-1,其中阳叶贡献约占总量的72%,阴叶约占总量的28%;(3)上层叶片贡献约占总量的75%,中层叶片约占总量的22%,下层叶片约占总量的3%。     

基于Ｌｏｔｋａ Ｖｏｌｔｅｒｒａ方程的基因调控网络的稳定性分析

在2007年7—8月棉花生长旺季,将冠层按高度分多层,通过简单的试验方法确定了棉冠内叶片丛聚指数（clumping index,Ω）,以此区分计算出田间各层逐时阳叶（受光叶）和阴叶（被遮荫叶）叶面积指数的动态值。在8月23日用LI-6400R便携式光合测量系统分多层分别测定阴、阳叶单张叶片净光合作用的日变化,结合阴、阳叶面积权重,探讨利用Ω区分阴阳叶之后对整个冠层日总光合作用的影响。试验结果表明：（1）花铃期棉花冠层内Ω约为0.68;（2）利用Ω计算得到8月23日冠层日间（09：00—18：00）总净光合作用日平均值大约为20.3μmol·m-2·s-1,其中阳叶贡献约占总量的72%,阴叶约占总量的28%;（3）上层叶片贡献约占总量的75%,中层叶片约占总量的22%,下层叶片约占总量的3%。     

基于碳水通量耦合原理改进Penman-Monteith蒸散发模型

陆地蒸散(ET)涵括地表和潮湿叶片的蒸发和植物的蒸散发,是陆地水循环的重要组成部分。Penman-Monteith方程是估算陆地蒸散的重要方法,方程中的叶片或冠层气孔导度是提高估算精度的关键因子。根据碳水循环的耦合原理,植物光合作用模型可用于估算叶片或冠层气孔导度。植物光合作用模型可分为三类:1)使用总冠层导度的大叶模型(BL),2)区别阴、阳叶冠层导度的双大叶模型(TBL),3)区别阴、阳叶叶片导度的双叶模型(TL)。与这三类光合作用模型相对应,衍生出基于不同导度计算方法的三种蒸散估算模型。三种蒸散模型之间的主要区别在于是否进行从叶片尺度到冠层尺度的气孔导度集成。这三种模型中,双叶模型使用叶片尺度的气孔导度,集成度最低。反之,大叶模型使用冠层尺度的气孔导度,集成度最高。由于在Penman-Monteith中,蒸腾和气孔导度之间的关系是非线性的,气孔导度的集合会导致负偏差。因此,与通量测量相比,大叶蒸散模型的估算偏差最大,而双叶蒸散模型的估算偏差最小。     

北方黑云杉林冠内空气CO2浓度及其上方通量模拟

考虑植物阴叶和阳叶对环境的反应、植物叶片丛聚特性以及叶片氮素水平的垂直差异,建立了黑云杉林与大气之间物质输送和能量交换的一维完全多层模式.模式中对阳叶和阴叶的叶面积指数借用Chen J M等提出的方法,并经过修改以适用于多层模拟.叶水平的光合作用c运用Farq uhar方程并与CO2传导方程联列获得.模式验证的资料取自BOREAS(Boreal Ecosystem-fAtmosphere Study)研究计划中加拿大Saskatchewan的南试验区,属于温带气候.通过对冠.层上方的显热、感热和CO2通量、以及植物光合作用、蒸腾作用和气孔传导等生理作用t过程的实测值进行了计算结果的验证,结果表明计算的显热通量比实测值偏低,潜热通量则略高于实测值,而CO2通量只有在较高水平时偏高,3个通量的计算值与实测值比较接近,R 2分别为0.71,0.78和0.65,取得了较好的效果.气孔导度、光合作用和蒸腾作用的模拟结果表明,三者的实测与模拟值之间的R 2分别达到0.57,0.69和0.66,均通过0.01显著度检验,且无显著的系统偏差.因此,可以认为在多层模拟中充分考虑叶片不同受光状况,不同氮素水平以及叶片丛聚特性的影响有助于更好地研究植被与大气的交换过程.     

双叶模型在冬小麦田冠层CO2通量多层模拟中的应用

综合考虑农田生态系统中水、热、CO2输送所涉及的大气、水文、生物等生物物理过程,以Farquhar等提出的叶片尺度光合作用生物化学过程机理模型为理论基础,对其进行空间尺度扩展,并改进冠层分层方法,建立了均匀农田与大气之间物质输送和能量交换的多层模式,在模式中运用双叶模型,同时考虑叶片氮素水平垂直差异,对2008年4—5月华北平原冬小麦生长旺季农田生态系统中冠层CO2通量进行了模拟研究,并利用涡度相关观测的通量数据对模型的有效性加以验证,结果表明:在冠层多层空间,小麦拔节至孕穗期和开花至乳熟期叶片氮含量随冠层高度的衰减系数分别为0.793(R2=0.698)和1.374(R2=0.728),冠层内叶片氮含量的空间分布可以用以相对累积叶面积指数为自变量的函数来描述;模型分别计算各层阴、阳叶的光截取、气孔传导、光合作用等,最终计算冠层上方CO2通量,冬小麦农田净生态系统生产力模拟值与实测值相关显著(R2=0.78),模拟的CO2通量日变化特征晴天昼间比阴雨天和夜间的效果好;在考虑丛聚影响的叶片非随机分布的密集农田中,阴叶对总初始生产力的贡献率在35.7%左右,对生产力贡献很重要。分层统计显示,作物最终产量的形成主要...     

棉花阴、阳叶的气孔导度和光合作用观测对比及模型应用

以实验观测数据为基础,对棉花阳叶和阴叶光合速率、气孔导度的变化进行比较,通过双叶模型和大叶模型对冠层光合作用进行对比分析,结果表明,大叶模型的计算结果普遍高于双叶模型,总体高出13％。由此可见,过去在研究植被一大气碳交换中使用大叶模型估算植物光合作用会夸大碳汇的作用。     

盘锦湿地芦苇叶片气孔导度的模拟

基于2005年5～9月盘锦湿地芦苇叶片气体交换观测数据,针对芦苇叶片气孔导度与光合速率以及光合速率与光合有效辐射之间的关系进行分析。结果表明:芦苇叶片气孔导度与光合速率的关系可应用Ball-Berry模型描述,光合速率与光合有效辐射的关系可应用非直角双曲线光合模型描述,联合Ball-Berry模型与非直角双曲线光合模型可通过环境变量求解叶片气孔导度。模型考虑了气孔导度与光合之间的相互作用。利用实验数据对气孔导度模型验证表明,叶片气孔导度模拟值和观测值回归方程的斜率为0.95,方程决定系数R=0.82(P<0.05)。     

黑河下游影响荒漠河岸胡杨林蒸腾的冠层与大气耦合分析

基于2014年胡杨主要生长季内树形特征、树干液流、环境因子的实际观测数据,利用经验公式,计算了胡杨冠层蒸腾速率、冠层气孔导度与解耦系数的值,分析了其日变化与季节变化特征。结果表明:(1)从日变化趋势来看,解耦系数在早晨和傍晚时较小,中午达到最大值,这主要是由于早晨和傍晚时太阳辐射比较弱、作物气孔开度小,使冠层气孔导度降低造成的;而中午时冠层气孔导度达到全天的最大值,解耦系数值也达到最大。(2)从季节变化趋势来看,解耦系数与冠层气孔导度变化趋势相近,在生长季内均呈先增大后减小,之后略有浮动增大,最后减小的趋势。本研究对影响荒漠河岸胡杨林蒸腾的冠层与下层大气进行相关性推断,认为影响胡杨林蒸腾的冠层与大气耦合度较高。尽管试验地处于极端干旱区,下层大气十分干燥,林冠层叶气界面水分散失很快,但黑河下游河岸林供水良好,林冠层空气动力学条件相近,使得胡杨林蒸腾主要受叶面气孔控制。     

陆面模拟中植被辐射传输参数化方案研究

在冠层二流辐射传输模式基础上新发展了一个描述太阳短波辐射在植被中传输的冠层四流辐射传输模式.冠层四流辐射传输模式是在大气辐射传输理论的基础上得到一组描述短波辐射在植被中传输过程的冠层辐射传输基本方程,引进大气中求解辐射传输方程的四流近似解法,并求得冠层四流辐射传输方程的解析解.方程中各项参量能够反映叶子或冠层特殊的几何和光学特征.冠层向上、向下辐射通量取决于冠层散射相函数、叶子在入射光方向投影面积、单个叶子反射率和透射率、叶面积指数以及直射光入射太阳高度角等.四流模式计算叶子水平倾角时对太阳短波辐射的反照率,与二流模式结果比较可以验证模式的理论推导和建模都是正确的:计算结果的比较,表明四流模式在水平叶角分布时计算的冠层反照率与二流模式结果一致,同时直射光从任何太阳高度角入射的冠层反照率结果也一致,从而证明发展的冠层四流辐射传输模式是成功的.模拟试验中将两种模型同时耦合到同一个陆面过程模式中进行比较试验,结果表明,冠层四流辐射传输模式能够得到更精确的植被反照率,从而使得陆面模式计算的地表吸收的净太阳辐射通量更接近于观测值.     

C3植物光合作用日变化的模拟

对前人光合作用-气孔导度耦合模型进行了修正,建立了光合作用-蒸腾作用-气孔导度的耦合模型,它概括了叶片上各主要物理过程和生理过程之间的相互联系和制约关系。 用数值方法研究了不同环境因子（太阳辐射、温度、湿度和风速等）对光合作用、蒸腾作用和气孔导度的日变化及中午降低（midday depression）的影响。 主要结果是:（1）当边界层导度减小时,光合“午睡”加剧,蒸腾作用减弱,但作为反馈调节,气孔导度增加。 （2）气孔导度的最适温度最低,光合作用次之,蒸腾作用最适温度最高。当光合作用中午受到高温的胁迫时,气孔导度下降的幅度最大,光合作用次之,蒸腾作用的降幅最小。一天中,气孔导度降低的持续时间最长,蒸腾作用降低的持续时间最短。（3）空气绝对湿度越低,气孔导度越低,光合午睡越明显。蒸腾作用则决定于饱和水汽压差(Vpd)和气孔导度两个因素的相反的作用。蒸腾作用随Vpd增加而增大,但Vpd超过一定值后,反而使蒸腾作用下降。 （4）当温度在光合最适温度以上时,太阳辐射的增加使叶温增加,引起光合“午睡”的加剧和气孔导度的降低。（5）ci/cs在中午的降低表明气孔的关闭是光合作用“午睡”现象的原因。     

Estimation of Biomass Heat Storage Using Thermal Infrared Imagery: Application to a Walnut Orchard

A new method to estimate tree biomass heat storage from thermal infrared (TIR) imaging of biomass surface temperature is presented. TIR images of the canopy are classified into trunk, branches, and leaves. The one-dimensional heat equation in cylindrical coordinates is forced with trunk and branch surface temperatures to simulate the temperature distribution and heat storage in tree trunks and branches. Assuming uniform leaf temperatures, heat storage in leaves is computed from the surface temperature of the leaves separately for the sunlit upper and shaded lower canopy. The sum of trunk, branches, leaf, and air heat storage gives the canopy heat storage. Measurements in a walnut orchard near Davis, California, in early June 2007 showed that biomass heat storage was of the same order as air heat storage and about 1% of daytime and 9% of nighttime net radiation.     

不同光质对温室甜椒光合特性的影响

以甜椒＂苏椒13号＂品种为试材,于2009年在江苏南京设计不同彩色塑料薄膜（红、绿、黄、紫、蓝、无色（CK））覆盖处理试验,系统研究了不同光质对温室甜椒叶片光合作用特性的影响。结果表明：不同光质处理的甜椒叶片光补偿点和光饱和点分别在4560μmol.m-2.s-1和1 0001 200μmol.m-2.s-1范围内;红膜处理的单叶最大光合速率最高达8.4μmol.m-2.s-1,紫膜处理最低仅为2.89μmol.m-2.s-1;红膜和CK处理的甜椒叶片CO2饱和点明显高于紫膜和黄膜处理,所有处理的CO2补偿点均在100μmol.mol-1左右。CK的叶绿素含量最高,绿膜处理最低。不同色膜处理的晴天叶片净光合速率、气孔导度、蒸腾速率日变化均呈单峰型。除蓝膜外,其他色膜处理胞间CO2浓度日变化曲线均呈＂W＂型。水分利用效率日平均值以红膜处理最高、紫膜最低。气孔限制值以紫膜处理最高、红膜处理最低。红膜、黄膜处理可促进甜椒光合作用,而紫膜则具有明显的抑制作用。     

Net radiation frequency distribution in a corn crop

Net radiation measurements were made with a traversing system in a 250 cm tall (LAI = 4.0) corn crop with east-west oriented rows. Frequency distributions of flux density of net radiation were plotted using 0.075 cal cm^-2 min^-1 class intervals. These distributions showed a shift from predominantly sunlit to predominantly shaded conditions with depth into the plant canopy. The frequency distributions showed that net radiation in sunflecks in a canopy can exceed that above the canopy, and that negative values of net radiation can exist in shadows. A Soil-Plant-Atmosphere Model (SPAM) also predicted that net radiation in sunflecks in the canopy would exceed net radiation above the canopy. The effects of spots with high radiation load within the plant canopy on leaf temperature and convective heat exchange were investigated theoretically.Contributed by the Northeast Branch, Soil and Water Conservation Research Division, Agricultural Research Service, U.S. Department of Agriculture, in cooperation with the Cornell University Agricultural Experiment Station, Ithaca, New York. Supported in part by the Atmospheric Sciences Laboratory, U.S. Army Electronics Command, Fort Huachuca, Arizona. Deparment of Agronomy Series, Paper No. 907.     

A Generalized Layered Radiative Transfer Model in the Vegetation Canopy

In this paper, a generalized layered model for radiation transfer in canopy with high vertical resolution is developed. Differing from the two-stream approximate radiation transfer model commonly used in the land surface models, the generalized model takes into account the effect of complicated canopy morphology and inhomogeneous optical properties of leaves on radiation transfer within the canopy. In the model, the total leaf area index （LAI） of the canopy is divided into many layers. At a given layer, the influences of diffuse radiation angle distributions and leaf angle distributions on radiation transfer within the canopy are considered. The derivation of equations serving the model are described in detail, and these can deal with various diffuse radiation transfers in quite broad categories of canopy with quite inhomogeneons vertical structures and uneven leaves with substantially different optical properties of adaxial and abaxial faces of the leaves. The model is used to simulate the radiation transfer for canopies with horizontal leaves to validate the generalized model. Results from the model are compared with those from the two-stream scheme, and differences between these two models are discussed.     

Influence of foliar density and thermal stability on profiles of Reynolds stress and turbulence intensity in a deciduous forest

Observations were made of turbulence in an extensive deciduous forest on level terrain using a vertical array of seven three-dimensional sonic anemometer/thermometers within and above the canopy. Data were collected through the period of leaf fall and over a range of thermal stabilities. A bulk canopy drag coefficient was nearly independent of the density of the forest but decreased greatly with the onset of nocturnal stability. The depth of penetration of momentum into the forest increased with leaf fall but, again, was greatly curtailed by stable conditions. Turbulent velocities decreased with increasing depth in the forest but relative turbulence intensities increased to mid-canopy levels. Leaf density influenced turbulence levels but not as strongly as did thermal stability. Thermal effects were adequately described by the single parameter h/L, where h is the canopy height and L is the Monin-Obukhov length. The longitudinal and vertical velocity correlation coefficient was larger in magnitude than expected in the upper layers of the forest but decreased to a small value in the lowest layers where the Reynolds stress was small. The ratio _w /u ^*, where u ^* is the local friction velocity, reflected changes in the uw correlation, becoming smaller than usual in the upper canopy layers. It is believed that these effects result from the intermittent, spatially coherent structures that are responsible for a large fraction of the momentum flux to the forest.     

Characteristics of air flow above and within soybean canopies

Air flow was observed above and within canopies of a number of kinds of soybeans. The Clark cultivar and two isolines of the Harosoy cultivar were studied in 1979 and 1980, respectively. Wind speed above the canopy was measured with cup anemometers. Heated thermistor anemometers were used to measure air flow within the canopy. Above-canopy air flow was characterized in terms of the zero-plane displacement (d), roughness parameter (z _o) and drag coefficient (C _d). d and z _o were dependent on canopy height but were independent of friction velocity in the range 0.55 to 0.75 m s^?1 · C _d for the various canopies ranged from 0.027 to 0.035. Greater C _d values were measured over an erectophile canopy than over a planophile canopy. C _d was not measurably affected by differences in leaf pubescence. Within-canopy wind profiles were measured at two locations: within and between rows. The wind profile was characterized by a region of great wind shear in the upper canopy and by a region of relatively weak wind shear in the middle canopy. Considerable spatial variability in wind speed was evident, however. This result has significant implications for canopy flow modeling efforts aimed at evaluating transport in the canopy. In the lower canopy, wind speed within a row increased with depth whereas wind speed between two rows decreased with depth. The wind speeds at the two locations tended to converge to a common value at a height near 0.10 m. The attenuation of within-canopy air flow was stronger in canopies with greater foliage density. Canopy flow attenuation seemed to decrease with increasing wind speed, suggesting that high winds distorted the shape of the canopy in such a manner that the penetration of wind into the canopy increased.