Seasonal variation in the energy and water exchanges above and below a larch forest in eastern Siberia

The water and energy exchanges in forests form one of the most important hydro‐meteorological systems. There have been far fewer investigations of the water and heat exchange in high latitude forests than of those in warm, humid regions. There have been few observations of this system in Siberia for an entire growing season, including the snowmelt and leaf‐fall seasons. In this study, the characteristics of the energy and water budgets in an eastern Siberian larch forest were investigated from the snowmelt season to the leaf‐fall season. The latent heat flux was strongly affected by the transpiration activity of the larch trees and increased quickly as the larch stand began to foliate. The sensible heat dropped at that time, although the net all‐wave radiation increased. Consequently, the seasonal variation in the Bowen ratio was clearly ‘U’‐shaped, and the minimum value (1·0) occurred in June and July. The Bowen ratio was very high (10–25) in early spring, just before leaf opening. The canopy resistance for a big leaf model far exceeded the aerodynamic resistance and fluctuated over a much wider range. The canopy resistance was strongly restricted by the saturation deficit, and its minimum value was 100 s m^?1 (10 mm s^?1 in conductance). This minimum canopy resistance is higher than values obtained for forests in warm, humid regions, but is similar to those measured in other boreal conifer forests. It has been suggested that the senescence of leaves also affects the canopy resistance, which was higher in the leaf‐fall season than in the foliated season. The mean evapotranspiration rate from 21 April 1998 to 7 September 1998 was 1·16 mm day^?1, and the maximum rate, 2·9 mm day^?1, occurred at the beginning of July. For the growing season from 1 June to 31 August, this rate was 1·5 mm day^?1. The total evapotranspiration from the forest (151 mm) exceeded the amount of precipitation (106 mm) and was equal to 73% of the total water input (211 mm), including the snow water equivalent. The understory evapotranspiration reached 35% of the total evapotranspiration, and the interception evaporation was 15% of the gross precipitation. The understory evapotranspiration was high and the interception evaporation was low because the canopy was sparse and the leaf area index was low. Copyright © 2001 John Wiley & Sons, Ltd.     

Evaporation and the subcanopy energy environment in a flooded forest

The combination of tree canopy cover and a free water surface makes the subcanopy environment of flooded forested wetlands unlike other aquatic or terrestrial systems. Subcanopy vapour fluxes and energy budgets represent key controls on water level and understorey climate but are not well understood. In a permanently flooded forest in south‐eastern Louisiana, USA, an energy balance approach was used to address (a) whether evaporation from floodwater under a forest canopy is solely energy limited and (b) how energy availability was modulated by radiation and changes in floodwater heat storage. A 5‐month continuous measurement period (June–November) was used to sample across seasonal changes in canopy activity and temperature regimes. Over this period, the subcanopy airspace was humid, maintaining saturation vapour pressure for 28% of the total record. High humidity coupled with the thermal inertia of surface water altered both seasonal and diel energy exchanges, including atypical phenomena such as frequent day‐time vapour pressure gradients towards the water surface. Throughout the study period, nearly all available energy was partitioned to evaporation, with minimal sensible heat exchange. Monthly mean evaporation ranged from 0.7 to 1.7 mm/day, peaking in fall when canopy senescence allowed greater radiation transmission; contemporaneous seasonal temperature shifts and a net release of stored heat from the surface water resulted in energy availability exceeding net radiation by 30% in October and November. Relatively stable energy partitioning matches Priestley–Taylor assumptions for a general model of evaporation in this ecosystem.     

Forest water use in the initial stages of reclamation in the Athabasca Oil Sands Region

Following large‐scale surface oil sands mining, large tracts of the boreal forest in the Athabasca Oil Sands Region of Western Canada are legally required to be reclaimed. A greater understanding of how these novel ecosystems function and develop with regard to water use is crucial to aid in the development of regulatory practices and protocols based on information from ecosystem recovery. In this paper, a 12‐year (2003–2014) eddy covariance measurement record of latent and sensible heat fluxes and gross ecosystem productivity of carbon dioxide is analysed to evaluate how a reclaimed boreal forest has developed during its initial growth period. The study site is a reclaimed oil sands saline‐sodic clay shale overburden deposit that was topped with 100 cm of glacial till and 20 cm of peat mineral mix. The site was seeded with barley (Hordeum spp.) in 2001 to reduce erosion of the soil cover whereas aspen (Populus tremuloides Michx.) and spruce (Picea glauca [Moench] Voss) boreal tree species were planted in 2004. Changes in structure and function corresponded to the transition of dominant vegetation cover from early successional species to forest. Leaf area index increased from a growing season peak of 0.9 in 2003 to 4.0 in 2014 and was associated with an increased growing season gross ecosystem productivity (4.9 to 8.9 g C m^?2 day^?1), an increased evapotranspiration (1.6 to 3.4 mm day^?1), and a decreased partitioning of energy to sensible heat (Bowen's ratio decreased from 1.1 to 0.4). Although canopy conductance increased throughout the 12 years, the shift from early successional species to trees with more conservative water use resulted in a decrease in conductance normalized by leaf area. Water use efficiency has increased slightly since 2008 with an average of 10.0 g CO₂ kg^?1 H₂O for the last 6 years. No prolonged dry periods were observed during the study period. The functioning of this novel ecosystem is evolving as expected on the basis of the trends observed for other natural and disturbed boreal forests.     

Evapotranspiration and canopy characteristics of two lodgepole pine stands following mountain pine beetle attack

Over the past decade, British Columbia (BC), has experienced the largest mountain pine beetle (MPB) outbreak on record. This study used the eddy‐covariance (EC) technique to examine the impact of the MPB attack on evapotranspiration (E) and associated canopy characteristics of two lodgepole pine stands with secondary structure (trees, saplings and seedlings surviving the attack) located in central BC. MPB‐06, an 85‐year‐old almost pure stand of pine trees, was first attacked in 2006, and by 2010, ~80% of the trees had been killed. MPB‐03, a 110‐year‐old stand with an overstory consisting of over 90% pine and a developed sub‐canopy, was first attacked in 2003 and by 2007 had > 95% pine canopy mortality. EC measurements began in August 2006 at MPB‐06 and in March 2007 at MPB‐03, and continued for four years. Annual total E ranged from 226 mm to 237 mm at MPB‐06, and from 280 to 297 mm at MPB‐03, showing relatively little year‐to‐year change at both sites over the four years. Increased E from the accelerated growth of the surviving vegetation (secondary structure, shrubs and herbs) compensated for reduction in E due to the death of the overstory. Monthly average daytime canopy conductance, the Priestley–Taylor (α), and the canopy–atmosphere decoupling coefficient (Ω) steadily increased during the growing season reaching approximate maximum values of 5 mm s^?1, 0.75 and 0.12, respectively. Potential evapotranspiration was approximated using a vapour pressure deficit‐dependent α obtained at high soil water content. Calculated water deficits indicated some water‐supply limitation to the surviving trees and understory at both sites. Rates of root zone drainage during the growing season were low relative to precipitation. Copyright © 2013 John Wiley & Sons, Ltd.     

Relationship between canopy height and the reference value of surface conductance for closed coniferous stands

When estimating the dry‐canopy evaporation rate of coniferous stands using the Penman–Monteith equation, it is crucial to determine the reference value of surface conductance G_s. This paper examines the relationship between canopy height and the reference value of G_s based on the maximum value of G_s with a vapour pressure deficit ≥ 1·0 kPa, ?_{s max}. There is a clear correlation between canopy height and ?_{s max} when the projected leaf area index ≥3·0. This suggests that using this relationship will enable more accurate determination of the reference value of G_s for closed stands. Copyright © 2003 John Wiley & Sons, Ltd.     

Water and heat fluxes above a lowland dipterocarp forest in Peninsular Malaysia

We measured the fluxes of sensible and latent heat between a low‐land dipterocarp forest in Peninsular Malaysia and the atmosphere. No clear seasonal or interannual changes in latent heat flux were found from 2003 to 2005, while sensible heat flux sometimes fluctuated depending on the fluctuation of incoming radiation between wet and dry seasons. The evapotranspiration rates averaged for the period between 2003 and 2005 were 2·77 and 3·61 mm day^?1 using eddy covariance data without and with an energy balance correction, respectively. Average precipitation was 4·74 mm day^?1. Midday surface conductance decreased with an increasing atmospheric water vapour pressure deficit and thus restricted the excess water loss on sunny days in the dry season. However, the relationship between the surface conductance and vapour pressure deficit did not significantly decline with an increase in volumetric soil water content even during a period of extremely low rainfall. Copyright © 2009 John Wiley & Sons, Ltd.     

Comparison of tree transpiration under wet and dry canopy conditions in a Costa Rican premontane tropical forest

Spatial and temporal variation in wet canopy conditions following precipitation events can influence processes such as transpiration and photosynthesis, which can be further enhanced as upper canopy leaves dry more rapidly than the understory following each event. As part of a larger study aimed at improving land surface modelling of evapotranspiration processes in wet tropical forests, we compared transpiration among trees with exposed and shaded crowns under both wet and dry canopy conditions in central Costa Rica, which has an average 4200 mm annual rainfall. Transpiration was estimated for 5 months using 43 sap flux sensors in eight dominant, ten midstory and eight suppressed trees in a mature forest stand surrounding a 40‐m tower equipped with micrometeorological sensors. Dominant trees were 13% of the plot's trees and contributed around 76% to total transpiration at this site, whereas midstory and suppressed trees contributed 18 and 5%, respectively. After accounting for vapour pressure deficit and solar radiation, leaf wetness was a significant driver of sap flux, reducing it by as much as 28%. Under dry conditions, sap flux rates (J_s) of dominant trees were similar to midstory trees and were almost double that of suppressed trees. On wet days, all trees had similarly low J_s. As expected, semi‐dry conditions (dry upper canopy) led to higher J_s in dominant trees than midstory, which had wetter leaves, but semi‐dry conditions only reduced total stand transpiration slightly and did not change the relative proportion of transpiration from dominant and midstory. Therefore, models that better capture forest stand wet–dry canopy dynamics and individual tree water use strategies are needed to improve accuracy of predictions of water recycling over tropical forests. Copyright © 2016 John Wiley & Sons, Ltd.     

Evapotranspiration from a wet temperate grassland and its sensitivity to microenvironmental variables

The eddy covariance and energy balance method was employed to determine evapotranspiration (LE) over a wet temperate C3–C4 co‐existing grassland in Japan. After sensible heat flux (H) was estimated via the eddy covariance technique, LE was calculated as the residual of the energy budget with calibration against the direct measurements of LE by a lysimeter. Daily mean LE varied from 0·8 to 10·5 MJ d⁻¹, with a peak at 16·5 MJ d⁻¹ in late July to early August. Day‐to‐day and seasonal variability in LE was affected appreciably by net radiation (R_n), atmospheric vapour pressure deficit (VPD), canopy surface conductance (g_c) and leaf area index (LAI). Before the canopy closure, LE responded to LAI in a linear manner. However, LE decreased with increasing LAI later in summer. Daytime variation in the decoupling coefficient (Ω) demonstrates that the canopy decoupled from the atmosphere in the morning and LE was primarily driven by the available energy, while in the afternoon the canopy partially coupled to the atmosphere so that LE was sensitive to VPD and g_c. Throughout the whole measurement period, Ω was generally larger than 0·5, suggesting that the available energy contributes more to LE than VPD. Copyright © 2004 John Wiley & Sons, Ltd.     

Canopy conductance in a two‐storey Siberian boreal larch forest,Russia

A larch forest in eastern Siberia was characterized by the presence of two distinct storeys, the overstorey with a small leaf area index (LAI) and a dense understorey with a relatively large LAI. To understand the roles of the overstorey and understorey in forest–atmosphere water exchange, canopy conductance (G_c), a critical parameter used in determining the energy and mass exchange, was calculated on the basis of latent heat flux above the overstorey and understorey, measured separately. Results showed that G_c for the overstorey (G_co) and understorey (G_cu) experienced different seasonal fluctuations. G_co was smaller than G_cu during periods of leaf expansion and leaf fall and showed an increasing trend until 1 month after the onset of leaf expansion. In contrast, a sharp decrease in G_co was observed immediately before onset of leaf fall. Furthermore, G_co was slightly larger than G_cu during the fully foliated period. A simple model using solar radiation and vapour pressure deficit (D) as inputs successfully reproduced the G_c in fully foliated periods with acceptable accuracy. Furthermore, both the understorey and overstorey in this study have a large reference G_c (G_c at D = 1 KPa) than their counterparts of other boreal forests and would not be able to sustain a constant leaf–soil water potential difference as D increases. We speculated that this confers the forest with an advantage allowing it to be able to sustain carbon assimilation during large D days and thus provides for the survival of the ecosystem during the short growing season at this site. Copyright © 2014 John Wiley & Sons, Ltd.     

Rainfall partitioning into throughfall,stemflow, and interception within a single beech (Fagus sylvatica L.) canopy: influence of foliation,rain event characteristics,and meteorology

While the hydrological balance of forest ecosystems has often been studied at the annual level, quantitative studies on the factors determining rainfall partitioning of individual rain events are less frequently reported. Therefore, the effect of the seasonal variation in canopy cover on rainfall partitioning was studied for a mature deciduous beech (Fagus sylvatica L.) tree over a 2‐year period. At the annual level, throughfall amounted to 71% of precipitation, stemflow 8%, and interception 21%. Rainfall partitioning at the event level depended strongly on the amount of rainfall and differed significantly (p < 0·001) between the leafed and the leafless period of the year. Therefore, water fluxes of individual events were described using a multiple regression analysis (r_a² > 0·85, n = 205) with foliation, rainfall characteristics and meteorological variables as predictor variables. For a given amount of rainfall, foliation significantly increased interception and decreased throughfall and stemflow amounts. In addition, rainfall duration, maximum rainfall rate, vapour pressure deficit, and wind speed significantly affected rainfall partitioning at the event level. Increasing maximum hourly rainfall rate increased throughfall and decreased stemflow generation, while higher hourly vapour pressure deficit decreased event throughfall and stemflow amounts. Wind speed decreased throughfall in the growing period only. Since foliation and the event rainfall amount largely determined interception loss, the observed net water input under the deciduous canopy was sensitive to the temporal distribution of rainfall. Copyright © 2007 John Wiley & Sons, Ltd.     

Hydrometeorological behaviour of pine and larch forests in eastern Siberia

Seasonal changes in the water and energy exchanges over a pine forest in eastern Siberia were investigated and compared with published data from a nearby larch forest. Continuous observations (April to August 2000) were made of the eddy‐correlation sensible heat flux and latent heat flux above the canopy. The energy balance was almost closed, although the sum of the turbulent fluxes sometimes exceeded the available energy flux (R_n ? G) when the latent heat flux was large; this was related to the wind direction. We examined the seasonal variation in energy balance components at this site. The seasonal variation and magnitude of the sensible heat flux (H) was similar to that of the latent heat flux (λE), with maximum values occurring in mid‐June. Consequently, the Bowen ratio was around 1·0 on many days during the study period. On some clear days just after rainfall, λE was very large and the sum of H and λE exceeded R_n ? G. The evapotranspiration rate above the dry canopy from May to August was 2·2 mm day^?1. The contributions of understory evapotranspiration (E_u) and overstory transpiration (E_o) to the evapotranspiration of the entire ecosystem (E_t) were both from 25 to 50% throughout the period analysed. These results suggest that E_u plays a very important role in the water cycle at this site. From snowmelt through the tree growth season (23 April to 19 August 2000), the total incoming water, comprised of the sum of precipitation and the water equivalent of the snow at the beginning of the melt season, was 228 mm. Total evapotranspiration from the forest, including interception loss and evaporation from the soil when the canopy was wet, was 208–254 mm. The difference between the incoming and outgoing amounts in the water balance was from +20 to ?26 mm. The water and energy exchanges of the pine and larch forest differed in that λE and H increased slowly in the pine forest, whereas λE increased rapidly in the larch forest and H decreased sharply after the melting season. Consequently, the shape of the Bowen ratio curves at the two sites differed over the period analysed, as a result of the differences in the species in each forest and in soil thawing. Copyright © 2003 John Wiley & Sons, Ltd.     

Observations of canopy storage capacity and wet canopy evaporation in a humid boreal forest

Evaporation of intercepted rain by a canopy is an important component of evapotranspiration, particularly in the humid boreal forest, which is subject to frequent precipitation and where conifers have a large surface water storage capacity. Unfortunately, our knowledge of interception processes for this type of environment is limited by the many challenges associated with experimental monitoring of the canopy water balance. The objective of this study is to observe and estimate canopy storage capacity and wet canopy evaporation at the sub-daily and seasonal time scales in a humid boreal forest. This study relies on field-based estimates of rainfall interception and evapotranspiration partitioning at the Montmorency Forest, Québec, Canada (mean annual precipitation: 1600 mm, mean annual evapotranspiration: 550 mm), in two balsam fir-white birch forest stands. Evapotranspiration was monitored using eddy covariance sensors and sap flow systems, whereas rainfall interception was measured using 12 sets of throughfall and six stemflow collectors randomly placed inside six 400-m² plots. Changes in the amount of water stored on the canopy were also directly monitored using the stem compression method. The amount of water intercepted by the forest canopy was 11 ± 5% of the total rainfall during the snow-free (5 July–18 October) measurement periods of 2017 and 2018. The maximum canopy storage estimated from rainfall interception measurements was on average 1.6 ± 0.7 mm, though a higher value was found using the stem compression method (2.2 ± 1.6 mm). Taking the average of the two forest stands studied, evaporation of intercepted water represented 21 ± 8% of evapotranspiration, while the contribution of transpiration and understory evapotranspiration was 36 ± 9% and 18 ± 8%. The observations of each of the evapotranspiration terms underestimated the total evapotranspiration observed, so that 26 ± 12% of it was not attributed. These results highlight the importance to account for the evaporation of rain intercepted by humid boreal forests in hydrological models.     

Canopy rainfall storage capacity as affected by sub‐alpine grassland degradation in the Qinghai–Tibetan Plateau,China

Grassland degradation resulting from global climate change, overgrazing, and rodent damage is expected to influence the magnitude of canopy hydrological fluxes because of reduced vegetation biomass and changed species composition. The objectives of this study were to estimate herbaceous canopy rainfall storage capacity (S) along three different stages of sub‐alpine grassland degradation (non‐degraded, lightly degraded and moderately degraded) in the Qinghai–Tibetan Plateau, China, and relate changes in S to canopy properties. An artificial wetting method and the water budget balance method, using rain simulations, were used for estimating S. Grassland degradation significantly reduced S. In non‐degraded, lightly degraded and moderately degraded grasslands, S estimated using the artificial wetting method were 0.612 ± 0.08 mm, 0.289 ± 0.04 mm, and 0.217 ± 0.01 mm, respectively; S estimated using the water budget balance method were 0.979 ± 0.32 mm, 0.493 ± 0.13 mm, and 0.419 ± 0.09 mm, respectively. These changes could be explained by accompanying changes in above‐ground biomass and leaf area index, as well as changes in species composition. Species‐specific rainfall storage capacity varied by a factor of 2.7 among the investigated species, with graminoids having the lowest values. Leaf area index was more correlated to S than was canopy coverage. Converting fresh weight of non‐leaf tissues into effective leaf area of the corresponding species and then introducing a coefficient of leaf area according to the specific storage capacity of leaves improved the linear relationship between S and leaf area index. Copyright © 2011 John Wiley & Sons, Ltd.     

Transpiration of a Linze jujube orchard in an arid region of China

As an important source of income in the region's economy, the jujube plantations are very common in arid north‐western China, and their planted areas continue to expand. In the central Heihe River Basin of arid north‐western China, Linze jujube (Zizyphus jujuba Mill. var. inermis (Bunge) Rehd.) plantations cover more than 10,000 ha, too. Water use by this species is expected to change or modify catchment hydrological process. To our knowledge, there is no information on the transpiration and canopy conductance of the jujube plantations in arid north‐western China. Therefore, Transpiration and canopy conductance were monitored in a 14‐year‐old Linze jujube orchard. The experiment was carried out in the central Heihe River Basin, near Pingchuan Town (Linze County, Gansu Province, China) during growing season of 2006, from May to the first ten days of October. Eight trees were used to measure sap flow using the heat‐pulse‐velocity method. The orchard was irrigated adequately during the study. Transpiration was estimated from the sap flow measurements. During the experiment, the transpiration rate of the orchard ranged from 0·32 to 1·40 mm per day. Canopy conductance was obtained from estimated daily transpiration and climatic variables measured on a half‐hour basis, and canopy conductance for water vapour transfer was between 1·20 to 82·57 mm s^?1, with a mean of 11·86 ± 6·84 mm s^?1 during the observation period. Air temperature and vapour‐pressure deficit exhibited a linear relationship with sap flow velocity and the relationship between these factors and canopy conductance could be represented by an exponential decay function. Copyright © 2009 John Wiley & Sons, Ltd.     

Water flux measurement and prediction in young cashew trees using sap flow data

Measurements of sap flow, meteorological parameters, soil water content and tension were made for 4 months in a young cashew (Anacardium occidentale L.) plantation during the 2002 rainy season in Ejura, Ghana. This experiment was part of a sustainable water management project in West Africa. The Granier system was used to measure half‐hourly whole‐tree sap flow. Weather variables were observed with an automatic weather station, whereas soil moisture and tension were measured with a Delta‐T profile probe and tensiometers respectively. Clearness index (CI), a measure of the sky condition, was significantly correlated with tree transpiration (r² = 0·73) and potential evaporation (r² = 0·86). Both diurnal and daily stomata conductance were poorly correlated with the climatic variables. Estimated daily canopy conductance g_c ranged from 4·0 to 21·2 mm s⁻¹, with a mean value of 8·0 ± 3·3 mm s⁻¹. Water flux variation was related to a range of environmental variables: soil water content, air temperature, solar radiation, relative humidity and vapour pressure deficit. Linear and non‐linear regression models, as well as a modified Priestley–Taylor formula, were fitted with transpiration, and the well‐correlated variables, using half‐hourly measurements. Measured and predicted transpiration using these regression models were in good agreement, with r² ranging from 0·71 to 0·84. The computed measure of accuracy δ indicated that a non‐linear model is better than its corresponding linear one. Furthermore, solar radiation, CI, clouds and rain were found to influence tree water flux. Copyright © 2005 John Wiley & Sons, Ltd.     

Influence of forest and shrub canopies on precipitation partitioning and isotopic signatures

Over a 4‐month summer period, we monitored how forest (Pinus sylvestris ) and heather moorland (Calluna spp. and Erica spp.) vegetation canopies altered the volume and isotopic composition of net precipitation (NP) in a southern boreal landscape in northern Scotland. During that summer period, interception losses were relatively high and higher under forests compared to moorland (46% of gross rainfall [GR] compared with 35%, respectively). Throughfall (TF) volumes exhibited marked spatial variability in forests, depending upon local canopy density, but were more evenly distributed under heather moorland. In the forest stands, stemflow was a relatively small canopy flow path accounting for only 0.9–1.6% of NP and only substantial in larger events. Overall, the isotopic composition of NP was not markedly affected by canopy interactions; temporal variation of stable water isotopes in TF closely corresponded to that of GR with differences of TF‐GR being ?0.52‰ for δ²H and ?0.14‰ for δ¹⁸O for forests and 0.29‰ for δ²H and ?0.04‰ for δ¹⁸O for heather moorland. These differences were close to, or within, analytical precision of isotope determination, though the greater differences under forest were statistically significant. Evidence for evaporative fractionation was generally restricted to low rainfall volumes in low intensity events, though at times, subtle effects of liquid–vapour moisture exchange and/or selective transmission though canopies were evident. Fractionation and other effects were more evident in stemflow but only marked in smaller events. The study confirmed earlier work that increased forest cover in the Scottish Highlands will likely cause an increase in interception and green water fluxes at the expenses of blue water fluxes to streams. However, the low‐energy, humid environment means that isotopic changes during such interactions will only have a minor overall effect on the isotopic composition of NP.     

Lumped parameter sensitivity analysis of a distributed hydrological model within tropical and temperate catchments

Parameter sensitivity of the Distributed Hydrology‐Soil‐Vegetation Model (DHSVM) was studied in two contrasting environments: (1) Pang Khum Experimental Watershed (PKEW) in tropical northern Thailand; and (2) Cedar River basin (CRB) in Washington State of the temperate US Pacific Northwest. The analysis shows that for both basins, the most sensitive soil parameters were porosity, lateral saturated hydraulic conductivity, and the exponential decrease rate of lateral saturated hydraulic conductivity with soil depth. The most sensitive vegetation parameters were leaf area index, vegetation height, vapour pressure deficit, minimum stomatal resistance (for both grassland and forest scenarios), hemisphere fractional coverage, overstory fractional coverage, and trunk space (for the forest scenario only). Parameter sensitivity was basin‐specific, with the humid, temperate CRB being more influenced by vegetation parameters, while tropical PKEW was more influenced by soil properties. Increases and decreases in parameter values resulted in opposite and unequal changes in bias and root mean square error (RMSE), indicating the non‐linearity of physical process represented in the hydrological model. Copyright © 2011 John Wiley & Sons, Ltd.     

Simulation of willow short‐rotation forest evaporation using a modified Shuttleworth–Wallace approach

Evaporation from a willow short‐rotation forest was analysed using a modified version of the Shuttleworth–Wallace model. The main modification consisted of a two‐layer soil module, which enabled soil surface resistance to be calculated as a function of the wetness of the top soil. Introduction of the threshold value of the leaf area index when scaling up from the leaf to the canopy resistance resulted in improvement to the simulated evaporation. The analysis was concentrated mainly on the 1988 season (May–October) when total evaporation was measured by the energy balance/Bowen ratio method throughout the growing season, covering all stages of canopy development. At the beginning of the 1994 season, soil evaporation were also measured with a ventilated chamber system. The general seasonal dynamics of the evaporation were fairly well simulated with the model. The largest deviation between measured and simulated evaporation occurred in June, when the model underestimated evaporation by about 1 mm day^?1. The model underestimated also in May but not as much as in June. In September and October the performance of the model was very good. For 130 days of the period May–October the cumulated measured evaporation was 364 mm and the simulated evaporation for the same days was 362 mm. It should be pointed out that this result was obtained without calibrating the model against the measured evaporation. The total simulated evaporation for the season was 450 mm with transpiration constituting 298 mm (66%), soil evaporation 102 mm (23%) and interception evaporation 50 mm (11%). The sensitivity analysis showed, in general, that simulated evaporation was most sensitive to changes in resistances when the leaf area index was smallest, i.e. under non‐closed canopy conditions. Changes in stomatal resistance, which is one of the most sensitive parameters, with associated changes in canopy transpiration, resulted in a negative feedback effect on soil evaporation. This reduced the total evaporation's sensitivity to stomatal resistance. This type of interaction between canopy and soil or undergrowth fluxes has been observed in other studies as well. Copyright © 2001 John Wiley & Sons, Ltd.     

Effect of succession gaps on the understory water‐holding capacity in an over‐mature alpine forest at the upper reaches of the Yangtze River

The water‐holding capacity (WHC) of the understory in the headwater regions of major rivers plays an important role in both the capacity of the forest water reservoir and water quality and quantity in the butted rivers. Although forest gaps could regulate water‐holding patterns in the understory by redistributing coarse woody debris (CWD), fine woody debris (FWD), non‐woody debris (NWD) and understory vegetation, little information is available on the effects of forest gaps on understory WHC. Therefore, we investigated the WHCs of CWD, FWD, NWD, herbaceous vegetation, mosses, epiphytes (including fern and lichen growing on the surface of logs) and soils from the gap centre to the adjacent closed canopy in an alpine forest at the upper reaches of the Yangtze River. The total WHC of the alpine forest understory components was approximately 300 mm. Soil layer had the largest contribution to the total understory WHC (90%), and among the aboveground components, CWD and mosses contributed 5% and 4% to the aboveground WHC, respectively. With the exception of that of the herbaceous layer, the WHC of the forest floor increased from the gap centre to the closed canopy. Although mosses had the lowest biomass allocation on the alpine forest floor, the water‐holding ratio (k) of mosses reached 485%. In conclusion, biomass is the parameter that most strongly and positively correlated with the WHC of the alpine forest understory, and forest gap formation decreases the understory WHC of alpine forest resulting from a decrease in organic soils, CWDs and mosses. Copyright © 2015 John Wiley & Sons, Ltd. Highlights

• The effects of gaps on the understory WHC were examined in an alpine forest.
• Gaps decreased the understory WHC by decreasing the amounts of the larger WHC components.
• The contribution of CWD and mosses to the aboveground WHC was large.
• The WHC of dead debris was higher than that of the vegetation.