Using packrat middens to assess grazing effects on vegetation change

Research on grazing effects usually compares the same sites through time or grazed and ungrazed sites over the same time period. Both approaches are complicated in arid environments where grazing can have a long undocumented history and landscapes can be spatially heterogenous. This work employs both approaches simultaneously by comparing grazed and ungrazed samples through both time and space using fossil plant macrofossils and pollen from packrat middens. A series of 27 middens, spanning from 995 yr BP to the present, were collected from Glen Canyon in southeastern Utah, USA. These middens detail vegetation change just prior to, and following, the historical introduction of domesticated grazers and also compares assemblages from nearby ungrazable mesas. Pre-grazing middens, and modern middens from ungrazed areas, record more native grasses, native herbs, and native shrubs such as Rhus trilobata, Amelanchier utahensis, and Shepherdia rotundifolia than modern middens from grazed areas. Ordinations demonstrate that site-to-site variability is more important than any temporal changes, making selection of comparable grazed versus ungrazed study treatments difficult. But within similar sites, the changes through time show that grazing lowered the number of taxa recorded, and lessened the pre-existing site differences, homogenizing the resultant plant associations in this desert grassland.     

Vegetation patterns and diversity along an altitudinal and a grazing gradient in the Jabal al Akhdar mountain range of northern Oman

Little is known about the effects of grazing on vegetation composition on the Arabian Peninsula. The aim of this study therefore was to analyse the vegetation response to environmental conditions of open woodlands along an altitudinal and a grazing gradient in the Jabal al Akhdar mountain range of Oman. The species composition, vegetation structure, grazing damage and several environmental variables were investigated for 62 samples using a nested plot design. Classification analysis and a Canonical Variate Analysis (CVA) were used to define vegetation types and to identify underlying environmental gradients. The relationship between environmental variables and diversity was analysed using correlation coefficients and a main-effects ANOVA. The plant species richness followed a unimodal distribution along the altitudinal gradient with the highest number of species at the intermediate altitudinal belt. The cluster analysis led to five vegetation groups: The Sideroxylon mascatense–Dodonaea viscosa group on grazed and the Olea europaea–Fingerhuthia africana group on ungrazed plateau sites at 2000 m a.s.l., the Ziziphus spina-christi–Nerium oleander group at wadi sites and the Moringa peregrina–Pteropyrum scoparium group at 1200 m a.s.l, and the Acacia gerrardii–Leucas inflata group at 1700 m a.s.l. The CVA indicated a clear distinction of the groups obtained by the agglomerative cluster analysis. The landform, altitude and grazing intensity were found to be the most important variables distinguishing between clusters. Overgrazing of the studied rangeland is an increasing environmental problem, whereas the plant composition at ungrazed sites pointed to a relatively fast and high regeneration potential of the local vegetation.     

Assessment of grazing effect on sheep fescue (Festuca valesiaca) dominated steppe rangelands, in the semi-arid Central Anatolian region of Turkey

In the semi-arid steppe rangelands of Central Turkey, Festuca valesiaca and Thymus sipyleus ssp rosulans have become the dominant species on degraded pastures. We hypothesized that decreases in species richness and abundance are correlated with increasing prevalence of these two species. Therefore, our objectives were to determine whether there are patterns in examined vegetation; how dominant species contribute to these patterns; and how patterns differ between grazed and ungrazed vegetation. We determined that protection from grazing increased species richness. Grazing significantly changed composition through decreasing total plant, forb, grass and F. valesiaca covers, while substantially increasing T. sipyleus cover. Topography, soil and grazing appear to impact the dominance of plant communities where F. valesiaca and T. sipyleus prevail. These two dominant species had a significant effect in shaping vegetation patterns. Based on regression analysis, alterations in species richness with changes in cover of forbs and shrubs were evident, and spatial heterogeneity of F. valesiaca and T. sipyleus indicated unstable vegetative patterns in heavily grazed pastures and successional changes in protected pastures. Our study results identify F. valesiaca and T. sipyleus as indicator species of vegetation suppression in condition assessments of degraded steppe rangelands.     

Physical impact of grazing by sheep on soil parameters in the Nama Karoo subshrub/grass rangeland of South Africa

The direct short-term impact of three rates of stocking (4, 8 and 16 small-stock units [SSU] ha^?1) was quantified in terms of soil characteristics of arid Nama Karoo vegetation (subshrub/grass). Mature Merino wethers grazed in the experimental plots throughout May in 1995 and 1996 (the plots were not subjected to grazing at any other time). Stocking rate proved inversely related to initial infiltration rate. Light trampling (4 SSU ha^?1) loosened the topsoil sufficiently to increase the initial infiltration rate: infiltration capacity of soil in fields stocked at 4 SSU ha^?1 and 16 SSU ha^?1 was 17% higher and 14% lower respectively than that of soil of ungrazed rangeland over the two grazing periods. Increased soil compaction and greater bulk density due to higher stocking rates significant decreased the infiltration rate. Compared to ungrazed rangeland stocking rates of 4, 8 and 16 SSU ha^?1 over the two grazing periods increased bulk density respectively by 2.73%, 6.67% and 8.945% and compaction by 10.90%, 16.78% and 20.90%. No grazing also increased bulk density and soil compaction and decreased infiltration rate. Light stocking (4 SSU ha^?1) influenced all soil parameters most favourably. From a hydrologic point of view, grazing levels and rotation schemes need to be tailored for sustainable utilization of arid subshrub/grass vegetation by livestock.     

Cattle grazing effects on annual plants assemblages in the Central Monte Desert,Argentina

Plant assemblages' dynamics have been studied to evaluate the influence of different environmental factors. The aim of our work was to assess the effect of a disturbance in the form of livestock grazing on annual plants, in a South American desert. We tested the exclusion of cattle grazing by comparing the composition of annual plants in three major vegetation types within a MaB Reserve in the Monte Desert of Argentina, with those of an adjacent grazed field. Sampling was conducted in two consecutive years that differed in precipitation. We established three sampling sites within each vegetation type at the reserve and the grazed field. Transects were set to assess plant cover, abundance, and richness. Theoretically, changes in diversity are explained by changes in one of its components: species richness. Species richness of annual plants was not different between the grazed and ungrazed sites. However, plant cover and diversity were lower in grazed sites, whereas abundance increased. Owing to a strong species-specific effect, we propose that it is evenness the main parameter involved in diversity dynamics in the heterogeneous vegetation mosaic of the Monte desert. Finally, vegetation types (spatial heterogeneity) and rainfall regime (temporal heterogeneity) greatly interacted with grazing effects. We endorse the idea that rapid responses of annual plant assemblages to changes in rainfall conditions coupled with herbivore control, might result in a restoration pathway for degraded arid landscapes.     

Small mammal herbivory: Feedbacks that help maintain desertified ecosystems

We tested the hypothesis that herbivores contribute to feedbacks maintaining arid ecosystems in a degraded state. We studied small mammal herbivory on a subshrub, broom snakeweed (Gutierrezia sarothrae), and perennial grasses at three sites: (1) ungrazed black grama (Bouteloua eriopoda) grassland; (2) grassland degraded by intense short-duration grazing; and (3) mesquite (Prosopis glandulosa) coppice dunes. Snakeweed was browsed by herbivores primarily during dry winter months.The average percent of G. sarothrae standing crop biomass removed by browsing was 9.2 in ungrazed grassland, 7.4 in intensely grazed grassland, and 4.1 in the dunes. In ungrazed grassland, an average of 12% of grass cover was harvested by herbivores; in the intensely grazed plots – 80%. Herbivore exclusion plots showed that jackrabbits (Lepus californicus) were the primary browsers on snakeweed and rodents on grasses and G. sarothrae inflorescences. Rodent removal of G. sarothrae inflorescences allows wind dispersal of seeds in disturbed and desertified areas, thereby increasing abundance of this poisonous shrub. Grass-tiller cutting by rodents provides a strong feedback that may be responsible for keeping the grass cover low on the intensely grazed areas. Jackrabbit pruning has little effect on G. sarothrae abundance at any stage of desertification.     

Diet of a fossorial rodent (Octodontidae), above-ground food availability,and changes related to cattle grazing in the Central Monte (Argentina)

Overgrazing by livestock has caused desertification in the Monte, where ctenomyids and livestock share grasses as main food items. The diet of Ctenomys eremophilus, above-ground food availability and changes related to cattle grazing are analyzed in the arid plain of Mendoza, Argentina. The most available categories were grasses, followed by low shrubs and tall shrubs. Tuco-tucos showed dietary generalism, ate mainly above-ground plant parts, preferred grasses and avoided shrubs at both grazed and ungrazed sites. Plant cover, grass diversity and availability decreased under livestock grazing, which was reflected in the diet by a lower percentage of grasses, a shift toward low shrubs and higher number of frequently used resources. Tuco-tucos in the grazed paddock compensated for lower consumption of vegetative plant parts by increasing the use of Prosopis flexuosa pods stored inside burrows. Moreover, greater dietary variation among individuals suggests foraging restricted to the items closest to burrow holes. These feeding tactics would allow them to reduce above-ground foraging as a response to high raptor predation risk due to increased bare soil. The plant recovery detected during the rest period, favoured by moderate stocking rate and rotational grazing system, would allow coexistence of tuco-tucos and cattle.     

Changes in soil nutrients, vegetation structure and herbaceous biomass in response to grazing in a semi-arid savanna of Ethiopia

The effect of grazing was studied on vegetation structure, herbaceous biomass, basal and bare ground covers, together with soil nutrient concentrations in two locations in an Ethiopian semi-arid savanna. The lightly grazed sites had significantly higher herbaceous diversity, total abundance, basal cover and aboveground biomass, and a lower percentage of bare ground compared with the heavy grazed sites. Grazing pressure had no effect on the density and number of woody species as well as on the proportion of encroaching woody species. The light grazing sites had higher organic carbon, phosphorus and exchangeable bases, and therefore a higher pH and higher electrical conductance, indicating an improved soil nutrient status compared with heavy grazing sites, mainly attributed to the higher basal cover and standing biomass at light grazed sites, and the export of nutrients through grazing and dung collection from the heavily grazed sites. There were significantly higher soil nutrients, species diversity, aboveground biomass and basal cover in the light grazing sites compared with heavy grazing sites. We concluded that changes in herbaceous vegetation, standing biomass and soil compositions are caused by interactions between grazing, soil and vegetation, and these interactions determine the transitions of semi-arid savannas.     

Effects of fire, grazing, and the presence of shrubs on Chihuahuan desert grasslands

Responses of herbaceous and suffrutescent species to fire, grazing, and presence of Prosopis glandulosa were examined in a Chihuahuan desert grassland in south-central New Mexico. Treatments were assigned randomly to eight 12×8 m plots within each of two blocks. Following fires in June 1995, unfenced plots were exposed to livestock grazing over 4 years. Plots were established that either included or excluded P. glandulosa. Perennial grass cover, primarilyBouteloua eriopoda , decreased by 13% in burned plots but increased 5% in unburned areas. Conversely, perennial forb cover was 4% greater after fire. Perennial grass frequency decreased 30% more and perennial forb frequency increased 10% more following burning. Further, increases in evenness after fire resulted in a 225% increase in species diversity. Grazing also resulted in a decrease in perennial grass cover while frequency decreased 22% more in grazed than ungrazed plots. Only frequency and not cover of perennial forbs and annual grasses increased more following grazing. Presence of P. glandulosa had no differential effect on responses of non-shrub species. Fires were conducted during near drought conditions while grazing occurred during years of precipitation equivalent to the long-term average. Precipitation immediately following fire may be critical for recovery of B. eriopoda -dominated desert grasslands; relationships between fire and post-fire precipitation patterns require future investigation.     

Grazing itineraries and forage selection of goats in the Al Jabal al Akhdar mountain range of northern Oman

In view of the repeatedly reported overstocking of the high-altitude pastures on Al Jabal al Akhdar, northern Oman, plant species abundance, cover and frequency, and herbaceous mass yield were studied in ungrazed versus heavily grazed areas of this mountain range. In addition, plant species selection by goats along a gradient of 1000–2000 m and spatial extent of pasture areas were investigated after abundant rainfall and a subsequent 6-months dry spell by means of manual observation and GPS/GIS tools.The substantially higher species diversity and herbaceous mass yield in the ungrazed area illustrate the production potential of these mountain pastures or, respectively, the biodiversity and productivity loss resulting from continuous grazing. The concentration of goats' selection on only a dozen herbaceous and ligneous species favours pasture encroachment with poisonous shrubs such as Nerium mascatense in the lower and Dodonaea viscosa in the higher altitudes. Given the spatially limited extent of pasture areas, these are exposed to high stocking rates. Therefore, grazing and feeding schemes need to be developed which reduce livestock pressure on the pastures, taking into account local property rights, herding skills and the recovery potential of the vegetation, which heavily depends on unpredictable rainfall events.     

Effect of grazing intensity on evapotranspiration in the semiarid grasslands of Inner Mongolia,China

The eddy covariance technique was used to measure evapotranspiration (ET) at four different grazing intensity sites to investigate the grazing effects on ET in the semiarid steppe ecosystems of Inner Mongolia. By reducing available energy, and decreasing soil water content (SWC), grazing decreased ET on a seasonal scale compared with the site ungrazed since 1979 (UG79). The most important climatic factor controlling ET on daily scale shifted from SWC to Net radiation (R_n) when grazing intensity increased. SWC, R_n and air temperature (or vapor pressure deficit) can explain 59%–71% of the variation in daily ET. On the other hand, leaf area index (LAI) affected ET slightly at UG79 under the commonly limited soil water conditions. Even no effect of LAI at the heavily grazed site was detected. This suggests that the direct effect of grazing reducing LAI on ET is not significant in this semiarid steppe ecosystem. Soil evaporation compensates for most of the loss in transpiration due to reduced LAI.     

Grazing effects on forb diversity and abundance in a honey mesquite parkland

Livestock grazing is the most common use of rangelands, and overgrazing by domestic livestock has stirred much debate concerning the continued use of public rangelands for livestock production. With the increase of wildlife-related enterprises on south Texas rangelands, there are little data available on the compatibility of livestock and wildlife management and in particular the effects of livestock grazing on plant communities in the semi-arid portions of the western South Texas Plains. The objective of this study was to compare forb diversity and density on rangelands subjected to various grazing under a high-intensity, low-frequency grazing system and nongrazed rangelands under controlled conditions. Four rangeland sites that were moderately grazed during the dormant season and 4 rangeland sites receiving long-term deferment from grazing were selected on the Chaparral Wildlife Management Area, Dimmit and La Salle Counties, Texas. Forb density was estimated with 20×50 cm² quadrats during early spring 2000 and 2001. Forb diversity was greatest on grazed sites. Important annuals such as hooker plantain (Plantago hookeriana) and Rough buttonweed (Diodia teres) were more prevalent on grazed sites. Lazy daisy (Aphanostephus spp.) was more frequently encountered on nongrazed sites. Golden dalea (Dalea aurea), a beneficial perennial, was more common on grazed sites. Long-term deferment from livestock grazing appears detrimental to forb communities. A high-intensity, low-frequency grazing system utilizing moderate stocking rates employed during the dormant season is recommended for enhancing forb diversity and abundance.     

Effect of sheep grazing on rangeland plant communities: Case study of landscape depressions within Syrian arid steppes

The arid rangelands of Syria cover over half of the nation's landmass. Punctuating this landscape are broad, dry basins, or wadis, and gentle landscape depressions that exhibit localized elevated vegetation productivity and unique edaphic and hydrologic properties. Historically, continuous heavy grazing and aggressive agricultural activities resulted in excessive ecological degradation within these sensitive environments. Information is needed to determine the influence of livestock grazing on plant communities in landscape depressions and the impact that this has on ecosystem resilience. The purpose of this research is to evaluate the effect of short-term sheep grazing on vegetation characteristics and plant community structure within depressions, and to provide recommendations for improved grazing management. Study plots were randomly located within paired topographic depressions located in northwestern Syria. Vegetation samples were collected along transects including plant biomass, plant density, herbaceous cover, and species diversity. In grazed plots, plant biomass was 49 g DM/m² compared to 234.4 g DM/m² in protected plots. Average plant density was 65 plants/m² in grazed plots compared to 1013 plants/m² in protected plots (P = 0.001). Herbaceous cover was 175% higher on protected sites compared to grazed plots. Average diversity (Shannon–Wiener index value) was 0.8 in grazed plots compared to 2.3 in protected plots. These results suggest that plant community structure will be impacted from short-term grazing and that a site's ability to positively respond to disturbance over time may be limited. We conclude that carefully planned grazing management should result in greater plant productivity and diversity.     

Stress resistance strategy in an arid land shrub: interactions between developmental instability and fractal dimension

This paper investigates allocation of energy to mechanisms that generate and preserve architectural forms (i.e. developmental stability, complexity of branching patterns) and productivity (growth and reproduction) in response to environmental disturbances (i.e. grazing and resource availability). The statistical error in translational symmetry was used to detect random intra-individual variability during development. This can be thought of as a measure of developmental instability caused by stress. Additionally, we use changes in fractal complexity and shoot distribution of branch structures as an alternate indicator of stress. These methods were applied to Anthyllis cytisoides L., a semi-arid environment shrub, to ascertain the effect of grazing and slope exposure on developmental traits in a 2×2 factorial design. The results show that A. cytisoides maintains developmental stability at the expense of productivity. Anthyllis cytisoides was developmentally more stable when grazed and when on south-facing, as opposed to north-facing slopes. On the contrary, shoot length, leaf area, fractal dimension and reproductive-to-vegetative allocation ratio were larger in north- than in south-facing slopes. As a consequence, under extreme xeric conditions, shrub mortality increased in north-facing slopes, especially when not grazed. The removal of transpiring area and the reduction of plant competition favoured developmental stability and survival in grazed plants. Differences between grazed and ungrazed plants were most evident in more mesic (north-facing) areas.     

Desert grassland canopy arthropod species richness: temporal patterns and effects of intense, short-duration livestock grazing

Arthropods living in the canopies of two woody shrub species (a sub-shrub (Gutierrezia sarothrae) and a large shrub (Prosopis glandulosa)) and perennial grasses plus associated herbaceous species, were sampled on 18, 0.5 hectare plots in a Chihuahuan Desert grassland for five consecutive years. Mesquite shrubs were removed from nine plots, six plots were grazed by yearling cattle in August and six plots were grazed in February for the last 3 years of the 5 year study. Arthropod species richness ranged between 154 and 353 on grasses, from 120 to 266 on G. sarothrae, and from 69 to 116 on P. glandulosa. There was a significant relationship between the number of families of insects on grass and G. sarothrae and growing season rainfall but species richness was not a function of growing season rainfall on any of the plants. Several of the arthropod families that were the most species rich in this grassland were found on all of the plants sampled, i.e. Salticid spiders, Bruchid and Curculionid beetles, Cicadellid and Psyllid homopterans, and ants (Formicidae). There were more species rich families that were shared by grasses and the sub-shrub G. sarothrae than with mesquite. The absence of a relationship between growing season rainfall and species richness was attributed to variation in life history characteristics of arthropods and to the non-linear responses of annual and perennial desert grassland plants to rainfall.There were no significant differences in insect family or species richness on any of the plant types as a result of removal of mesquite (P. glandulosa) from selected plots. Intense, short duration (24 h) grazing by livestock during in late summer resulted in reduced species richness in the grass-herb vegetation layer but had no effect on insect species richness on snakeweed or mesquite shrubs. Livestock grazing in winter had no effect on insect species richness on any of the vegetation sampled.     

沙质草地放牧和封育下物种丰富度和生物量关系的季节变化

研究了科尔沁沙质草地放牧和封育下植被盖度、密度、丰富度与地上生物量关系的季节变化。结果表明:放牧和封育草地4月的植被盖度、丰富度和生物量均低于生长旺季8月,而4月植物密度要高于8月;封育草地生长季盖度高于放牧草地,6月和8月生物量高于放牧草地, 4月和6月植物密度则低于放牧草地。封育草地8月物种丰富度与生物量呈极显著的负线性关系（p<0.01）;放牧草地植物密度与生物量4月表现出极显著的正线性关系（p<0.01）、8月为极显著的负线性关系（p<0.01）。放牧草地优势植物糙隐子草4月的密度和生物量呈极显著的正线性关系（p<0.01）、8月为极显著的负线性关系（p<0.01）;黄蒿和狗尾草8月的密度和生物量均呈极显著的负线性关系（p<0.01）;猪毛菜4月和6月的密度和生物量呈极显著的正线性关系（p<0.01）。封育草地达乌里胡枝子、猪毛菜8月的植物密度和生物量呈极显著的正线性关系（p<0.01）。持续放牧和季节变化及其二者的交互作用对植被盖度、密度和生物量均有重要的影响（p<0.05）;放牧通过影响草地生长季中的优势植物密度变化和生物量积累,引起了草地植物密度和生物量关系的季节变化,也导致生长季物种丰富度和生物量无显著关系;封育草地植物的竞争导致了物种丰富度和生物量关系的季节变化。     

Is the analysis of plant community structure better than common species-diversity indices for assessing the effects of livestock grazing on a Mediterranean arid ecosystem?

Grazing in arid Mediterranean ecosystems brings about changes in species diversity, but the best way to measure such changes is unclear. In this study we compared various methods in order to identify indicators that might be useful for the management and conservation of grazed arid Mediterranean ecosystems.Changes in community structure and composition were compared along a previously studied grazing gradient in Cabo de Gata-Nijar Natural Park. Plant diversity was measured by calculating diversity (Shannon information index), evenness (Pielou index) and richness (species number). Rarefaction curves were used to measure plant species richness and the results were compared with traditional richness values. Community structure was measured as the percentage of bare ground and the coverage of overall perennial grasses, Stipa tenacissima L., perennial shrubs and annual plants. Our results showed that the proportion of bare ground, the cover of perennial shrubs and the relative abundance of S. tenacissima are good indicators of grazing effects on vegetation. Overall plant community structure was more sensitive to the grazing gradient than were the diversity indices. Finally, the adequacy of standardized methods for comparing species richness (i.e. rarefaction curves) along a grazing gradient are discussed.     

Grazing effects on patchy dryland vegetation in northern Patagonia

In this study the spatial patterns and dynamics of vegetation patches along a grazing gradient in the steppe ofLarrea divaricataandStipaspp. in NE Patagonia (Argentina) are described. A general effect of grazing is the reduction of total plant cover resulting from the decrease in cover of perennial grasses (Stipa speciosa, Poa ligularis, Stipa tenuis) and some tall shrubs (Chuquiraga hystrix, Bougainvillea spinosa, Lycium chilense). Dwarf shrubs (Nassauvia fueguianaandJunellia seriphioides) increase their cover under medium and/or high grazing pressures. Plant species are spatially grouped into patches which alternate with areas of bare soil. Eleven types of vegetation patch differing in the dominant plant functional type or species, floristic richness and size were identified with different relative frequency along the grazing gradient. Based on these results, it is postulated that grazing forces the replacement of large patches dominated by tall shrubs with high species richness, byLarrea divaricatapatches or small dwarf shrub patches with low species richness and the extinction of grass patches. This results from: (1) disruption of local balances of species deletions and additions; (2) fragmentation of large patches; and (3) formation of new vegetation patches. These changes lead to differing plant spatial organization and heterogeneity along the grazing gradient which may be described by characteristic arrays of vegetation patches.     

Effects of seasonal grazing,drought, fire,and carbon enrichment on soil microarthropods in a desert grassland

This study was designed to test hypotheses about the combined effects of short-term, seasonal grazing with seasonal drought, fire, and carbon enrichment on soil microarthropod communities in a Chihuahuan Desert grassland. The study was conducted in eighteen 0.5 ha plots following three consecutive years of treatment: six plots intensively grazed in summer, six in winter, and six not grazed. There was no difference in perennial grass cover on the summer-grazed and winter-grazed plots. Intensive seasonal grazing had no effect on the abundance and community composition of soil microarthropods. Within each plot there were six subplots: summer rain-out, winter rain-out, burned, glucose amendment, rain-out control and burn-glucose control. Fire and carbon enrichment had no significant effect on soil microarthropod abundance or community composition. The average number of microarthropods ranged from 8915 ± 1422 m⁻² in the ungrazed, unburned plots to 7175 ± 1232 m⁻² in the winter-grazed, unburned plots. Microarthropod densities in the glucose-amended plots were 8917 ± 4902 m⁻² in the winter-grazed plots and 10,731 ± 863 m⁻² in the glucose-amended, summer-grazed subplots.The prostigamatid mite, Tydeus sp., was the most abundant microarthropod taxon in all treatment plots.     

Camel grazing affects species diversity and community structure in the deserts of the UAE

Camel grazing plays a crucial role in the desert ecosystems of the UAE. In this study, we compare areas grazed by small antelope (Al Maha Resort – the AMR) with areas grazed by both camels and small antelope (Dubai Desert Conservation Reserve – the DDCR). A total of 126 plots were selected during the growing season 2006/07 on three soil substrates: gravel plains, sand flats and sand dunes. In each plot, several vegetation parameters were assessed: density, frequency, percent cover and diversity indices. The replacement of camels with wild antelope has significantly increased the number of species on gravel plains, vegetation density on sand dunes and diversity indices on both sand flats and sand dunes, but significantly decreased plant cover on sand flats and sand dunes. The increase in species diversity in the AMR was attributed to moderate grazing by antelope. Replacement of camels by antelope in the AMR has resulted in change in plant community composition of the three substrate types. Species recovered after protection from camel grazing are palatable, especially for camels, except Heliotropium kotschyi and Aerva javanica. The absence of most of the palatable species from the DDCR was attributed to both selective foraging and overgrazing by camels.