Late Permian-Early Triassic Conodont Biofacies and Bioprdvinces in South China and Himalaya1

Abstract On the basis of existing conodont data, the authors have studied the Late Permian-Early Triassic cohodonts of different farms and biofacies in detail Five conodont biofacies are recognised, from shallow to deep water: 1. Hindeodus conodont biofacies 2. Pachycladina-Parachi rognathus conodont biofacies, 3. Gondolella-Hindeodus Conodont biofacies, 4. Gondolella - Neospathodus conodont biofacies, and 5. Xaniognathus cohodont biofacies. Considering the temporal and spatial changes of these conodont biofacies, 3 conodont bioprovinces have been divided. In light of the biofacies changes of each bioprovince, the authors also discuss, in this paper, the regularity of transgression-regression cycles of eastern Tethys and their possible relation to the mass biotic alternation.     

华南和喜马拉雅古,中生代之交牙形石生物相和生物分区

在前人工作的基础上，对华南和喜马拉雅地区二叠世－早三叠世不同生态类型的牙形石进行了详细研究研究，依水体深浅不同，从浅到深识别出５种牙形石生物相即Ｈｉｎｄｅｏｄｕｓ生物相，Ｐｃａｃｈｙｃｌａ，ｄｉｎａ－ｐａｒａｃｈｉｒｏｇｎａｔｈｕｓ生物相，Ｈｉｎｄｅｏｄｕｓ－Ｇｏｎｄｏｌｅｌｌａ生物相Ｇｏｎｄｏｌｅｌｌａ－Ｎｅｏｓｐａｔｈｏｄｕｓ生物相和Ｘａｎｉｏｇｎａｔｈｎｓ生物相。通过这些牙形石生物相在时间和     

Early–Middle Ordovician conodont biofacies on the Yangtze Platform margin,South China: Applications to palaeoenvironment and sea-level changes

9172 Conodonts have been recovered from the uppermost Hunghuayuan Formation and the Zitai Formation at two sections in Shitai County, southern Anhui Province, South China, which was situated close to the margin of the Lower Yangtze Platform during the Early to Middle Ordovician. Systematic and multivariate statistical studies on these conodonts permit recognition of seven conodont biofacies: Tropodus biofacies, Diaphorodus biofacies, Oepikodus biofacies, Baltoniodus biofacies, Paroistodus biofacies, Periodon biofacies and Protopanderodus biofacies. Each biofacies is restricted to a particular lithofacies and stratal position and shows a consistent order and/or position within the succession. Turnover of these conodont biofacies is related to sea-level changes. The transgressive–regressive patterns demonstrated by the conodont biofacies compare closely to published sea level curves for South China, and highlight the utility of conodont biofacies as a means of confirming sedimentological evidence of relative sea-level change.     

安徽巢湖平顶山下三叠统牙形石生物地层序列

简要介绍安徽巢湖平顶山下三叠统牙形石生物地层研究的最新成果.对平顶山西坡剖面112.8 m厚的下三叠统地层所采集的286件牙形石样品, 经处理后获得2 870余枚牙形石样本, 经鉴定共有11属51种.这些属分别是Hindeodus, Isarcicella(?), Neogondolella, Neospathodus, Platyvillosus, Cratognathodus, Parachirognathus, Pachycladina, Ellisonia, Aduncodina和Cornudina.牙形石生物地层从老到新可划分为9个牙形石带: (1)Hindeodus typicalis- Neogondolella planata带; (2)Neogondolella krystyni带; (3)Neospathodus kummeli带; (4)Neospathodus dieneri带; (5)Neospathodus n. sp. C- Neospathodus n.sp. D带; (6) Neospathodus waageni带; (7)Neospathodus n.sp. M带; (8)Neospathodus eotriangularis带; (9) Neospathodus abruptus- Neospathodus homeri带.牙形石的组合面貌基本可与国内外其他地区牙形石带相对比.但其中牙形石(2), (3), (5), (7), (8)带为巢湖地区首次建立.在下三叠统菊石带控制下所建立的连续的牙形石分带对全球的三叠系阶的对比具有十分重要的意义. 关键词: 牙形石; 牙形石生物地层; 下三叠统; 安徽巢湖.      

Latest Devonian to earliest Carboniferous conodont and carbon isotope stratigraphy of a shallow‐water sequence in South China

To improve regional and intercontinental correlation of the uppermost Devonian–lowermost Carboniferous, we examined the conodont faunas and carbon isotopic records of the Tangbagou Formation in the Qilinzhai section, southern Guizhou, South China. The Tangbagou Formation is a succession of mixed carbonate–siliciclastic rocks that accumulated on a shallow‐water platform under normal marine conditions. Seven conodont zones for shallow‐water biofacies in South China, the Cl. gilwernensis–Cl. unicornis Zone, the Po. spicatus Zone, the Si. homosimplex Zone, the Si. sinensis Zone, the Si. eurylobata Zone, the Ps. multistriatus Zone and the Po. co. porcatus Zone in ascending order, are recognized in the Tangbagou Formation. Although apparently limited in its value for global correlation, this conodont zonation is more applicable to shallow‐water biofacies in South China. Carbonate samples have yielded carbon isotopic signatures consistent with those recorded in Euroamerica sections, in particular showing four distinct characteristics: (1) the peak values of Hangenberg Carbon Isotope Excursion (HICE) during the latest Devonian, (2) a minor positive shift (P1) in the Si. homosimplex Zone during the early Tournaisian, (3) a second minor positive shift (P2) in the Si. sinensis Zone and (4) the middle Tournaisian Carbon Isotope Excursion (TICE) in the middle part of the Tangbagou Formation. The similarity in peak values (~5.5‰) and magnitude of TICE for the Qilinzhai and Belgian sections indicates that the Euro‐asia δ¹³C_carb trends may reflect the changes in global mean ocean δ¹³C_DIC, rather than having been overprinted by local carbon cycling. Integration of conodont biostratigraphy and δ¹³C stratigraphy provides a powerful tool for stratigraphic correlation. Copyright © 2015 John Wiley & Sons, Ltd.     

Upper Famennian and Lower Tournaisian sections of the Moravian Karst (Moravo‐Silesian Zone,Czech Republic): a proposed key area for correlation of the conodont and foraminiferal zonations

Foraminiferal and conodont faunas at the Devonian–Carboniferous (D–C) boundary in the southern part of the Moravian Karst (Czech Republic) were studied in different facies of the basin slope. The joint presence of foraminifers and conodonts in calciturbidites along with a positive δ¹³C excursion of the Hangenberg anoxic event enabled the high‐resolution calibration of the late Famennian–early Tournaisian interval (Upper expansa–crenulata conodont zones). The conodont stratigraphic and biofacies succession reveals a strong correlation with other European areas. The Siphonodella sulcata morphotype (close to Group 1 sensu Kaiser and Corradini and “nov. gen. nov. sp. 1” sensu Tragelehn) enters prior to the Hangenberg Event, which resembles Upper and Uppermost Famennian conodont successions from Franconia, Bavaria and Morocco. The diversification of the early siphonodellids takes place after the Hangenberg Event and after the protognathodid radiation. In terms of foraminiferal biostratigraphy, the D–C boundary interval is characterized by the first appearance datum (FAD) of Tournayellina pseudobeata close below the D–C boundary followed by a sequence of Tournaisian bioevents, where apart from the last appearance datums (LADs) of quasiendothyrs, the FADs of the Neoseptaglomospiranella species and chernyshinellids play an important role in a similar manner as in Eastern Europe. The correlation of these bioevents elsewhere is often hindered by glacioeustatically‐driven unconformities and widespread occurrences of unfavourable facies for plurilocular foraminifers (Malevka beds and Bisphaera beds). Copyright © 2013 John Wiley & Sons, Ltd.     

华南下/中奥陶统界线附近笔石相的分异与全球下/中奥陶统界线生物标志的选择

通过对宜昌黄花场(扬子地台中部)大湾组剖面和重庆城口(扬子地台北部)的湄潭组剖面笔石动物群特征的分析,结合江西崇义地区(盆地相)、江南过渡带笔石动物群的研究成果,讨论了华南下／中奥陶统界线附近笔石相分异的现象,并探讨了下／中奥陶统界线附近笔石动物和牙形石的相互关系。认为宜昌黄花场剖面完全具备建立全球下／中奥陶统界线层型剖面和点(GSSP)的条件,牙形石B.triangularis的首次出现(FAD)是良好的界线生物标志。     

浙江长兴地区二叠纪与三叠纪之交牙形石动物群及地层意义

本文把采自华南浙江长兴煤山二叠系与三叠系界线剖面的牙形石动物群从老至新分为三个带:1.Gondolella subcarinata-G·wangi组合带;2.G.changxingensis-G.deflecta组合带;3.Anchignathodus parvus带。将煤山剖面的二叠系与三叠系界线划在2、3组合带之间。并初步探讨了长兴地区长兴期至早格里斯巴赫期的牙形石分异度与沉积环境变迁。     

新疆阿图什温古尔地区二叠纪牙形石动物群

新疆阿图什的温古尔－巴音库鲁提一带二叠系由于化石稀少而且保存不好，地层确切时代仍存在疑问，该区温古尔剖面二叠纪重要牙形石化石Neostreptognathodus pequopensis Behnken,Streptognathodus nashuiensis Zhong et Zhang等的发现为这套地层确切时代的确定提供了重要依据。通过牙形石生物相分析，推断温古尔地区中二叠世早期可能为盆地相较深水沉积环境。     

牙形石动物分类归属研究新进展—牙形石是最早的脊椎动物

在简要回顾牙形石动物分类位置研究历史的基础上，主要介绍了近十年来这一问题的最新研究成果。英国古生物学家们通过牙形石动物的软体解剖学研究，发现眼睛、脊索、神经索、肌肉组织、尾鳍等软组织；牙形石分子硬体组织学研究揭示了细胞状骨骼、球状钙化软骨、类珐琅质及牙质等硬组织。新的证据表明牙形石动物是脊椎动物，从而将脊椎动物的历史向前推移了４０～５０Ｍａ。     

Subdivision of the Lochkovian Stage based on conodont faunas from the stratotype area (Prague Synform,Czech Republic)

Relatively rich conodont faunas from sections in the Prague Synform (Barrandian area, Czech Republic) include a number of indexes and other important guide conodonts that can be correlated with other regions, especially with Nevada and the Spanish Central Pyrenees. The collation and detailed correlation of conodont data from the Lochkovian in two parallel sections in the Požáry quarries, together with biostratigraphic control of additional data from several (incomplete) sections with changing facies development, is the basis for a new detailed regional biozonal scale for the Lochkovian in the Prague Synform. The new subdivision follows, with modification, the global threefold conodont subdivision of the Lochkovian. Data from the Prague Synform enable further detailed subdivision of the lower, middle and upper Lochkovian into small‐scale units. The conodont distribution shows a large proportional discrepancy between the late Lochkovian elsewhere; the conodont record in the latest Lochkovian in the Prague Synform area, which appears to be rather restricted and requires further discussion. Copyright © 2012 John Wiley & Sons, Ltd.     

Guadalupian （Middle Permian） Conodont Faunas at Shangsi Section, Northeast Sichuan Province

This study recovered the Guadalupian conodont faunas from Shangsi (上寺) Section in Northeast Sichuan (四川). Four genera and nine species were identified, and three conodont zones were recognized and established; they include Jinogondolella nankingensis Zone, J. aserrata Zone, and J. postserrata Zone. The Roadian and Wordian boundary is set in the interval 5 m from the top of Bed 86 by the first appearance of the conodont J. aserrata. The Wordian and Capitanian boundary is set in the interval 2.3 m from the top of the Bed 95 by the first appearance of J. postserrata. Most of the specimens demonstrated low color alteration index (CAI) as 1.5-3, indicating that most part of the Maokou (茅口) Formation may have the suitable thermal conditions for the formation of oil source rocks.     

论利用牙形石色变度值研究碳酸盐岩低级─极低级变质作用

牙形石色变度──ＣＡＩ值是一项有效的古地温指标，它不但可以帮助确定低级－极低级变质碳酸盐岩的最高变质温度，而且可以判别变质类型。本文在介绍牙形石ＣＡＩ值研究现状及低级－极低级变质作用研究现状的基础上，对运用ＣＡＩ值进行低级－极低级碳酸盐岩的研究进行了综述及探讨。     

全球奥陶系底界的“金钉子”问题及我国特马豆克阶(Tremadocian)的划分与对比

对全球寒武系与奥陶系界线层型剖面和点位(GSSP)——西纽芬兰绿岬(Green Point)剖面和我国吉林白山(原浑江)大阳岔小洋桥原全球寒武系与奥陶系界线候选剖面的对比研究表明,绿岬"金钉子"剖面所指定的界线生物标志——波动古大西洋牙形石(Iapetognathus fluctivagus)并不存在于所指定的界线生物层和点位上,而且在分布和分类上是均存在争议的物种,更遗憾的是,所展示的大部分牙形石和笔石标本图片以及碳氧同位素异常数据也并非来自该"金钉子"剖面,因而完全违背了选择金钉子剖面的原则和要求,有必要进行重新的评估。小洋桥寒武系与奥陶系界线剖面交通方便、环境优美,寒武系与奥陶系界线地层系由一套深水且未变质的黑色、黄绿色页岩与灰色薄层瘤状灰岩所构成的韵律沉积组成,并保存了完整的具有广泛代表性牙形石和笔石序列,在详细研究的34 m界线间隔中,自下而上保存了完整的Cambrostodus、Codylodus proavus、C.intermedius、C.lindstromi和C.angulatus等5牙形石生物带,在C.intermedius带上部至C.angulatus带之间,还依次出现3层笔石,归属于重新厘定的Rhabdinopora parabola和Anisograptus matanensis等2个笔石带,此外还有大量三叶虫和介形类化石共生,结合高精度同位素地球化学的研究,笔者等建议,以全球广泛分布的牙形石C.intermedius的首现,取代Iapetognathus fluctivagus,作为寒武系与奥陶系界线划分对比的标志,其层位与原来所指定的界线层几乎一致或接近,界线之上所记录的最大碳同位素异常和首次出现的最早的浮游正笔石——R.parabola(含R.praeparabola),可作为该界线划分的辅助标志。     

浙江长兴煤山二叠-三叠系界线层牙形石序列及其全球对比

在详细研究全球二叠－三叠系界线层候选剖面───中国浙江长兴煤山剖面Ｐ／Ｔ界线层牙形石动物群序列基础上，自下而上识别出４个牙形石带：（１）ＣｌａｒｋｉｎａｃｈａｎｇｘｉｎｇｅｎｓｉｓＣｌａｒｋｉｎａｄｅｆｌｅｃｔａ带；（２）Ｉｓａｒｃｉｃｅｌｌａｐａｒｖａ带；（３）Ｉｓａｒｃｉｃｅｌｌａｉｓａｒｃｉｃａ带；（４）ＣｌａｒｋｉｎａｃａｒｉｎａｔａＣｌａｒｋｉｎａｐｌａｎａｔａ带，并在第一带中划分出３个动物群，从下向上是：ＣｌａｒｋｉｎａｃｈａｎｇｚｉｎｇｅｎｓｉｓＣｌａｒｋｉｎａｄｅｆｌｅｃｔａ－Ｃｌａｒｋｉｎａｓｕｂｃａｒｉｎａｔａ动物群，Ｈｉｎｄｅｏｄｕｓｌａｔｉｄｅｎｔａｔｕｓ－Ｃｌａｒｋｉｎａｍｅｉｓｈａｎｅｎｓｉｓｓｐ．ｎｏｖ．动物群和Ｈｉｎｄｅｏｄｕｓｔｙｐｉｃａｌｉｓ动物群，煤山剖面二叠、三叠系界线层牙形石序列的建立、完善，对确立该剖面在全球二叠、三在系界线高精度对比中的标准地位具重要意义。     

鄂西地区上二叠统的牙形石及其分带意义

湖北恩施天桥剖面发育有完好的二叠系大隆组下段硅质岩层。对其下伏吴家坪组灰岩及上覆大隆组上段灰岩中牙形石的研究表明,吴家坪组灰岩自下而上可分为Clarkina postbitteri postbitteri带、Clarkina dukouensis带、Clarkina asymmetrica带以及Clarkina guangyuanensis—Clarkina transcaucasica带,属吴家坪阶中下部牙形石带;大隆组上段灰岩可划分出两个牙形石带,即Clarkina subcarinata—Clarkina wangi带和Clarkina changxingensis changxingensis—Clarkina deflecta带,为长兴阶底部牙形石带。认为鄂西地区大隆组下段硅质岩的时代应归属于二叠纪吴家坪中期至长兴早期。     

荆州地区晚三叠世—中侏罗世地层及沉积环境

晚三叠世—中侏罗世沉积地层特征表明，该时期荆州地区为前陆盆地构造环境，形成于秦岭造山带南部前陆地带，沉积特征及上下接触关系表明其为Ⅰ级构造层。上三叠统和中下侏罗统具有相反的粒序特征和不同的建造特征，分属于两个Ⅱ级层序，分别由３个和７个Ⅲ级层序构成。综合研究结果表明，荆州前陆盆地经历了三个演化阶段，即挠曲变形阶段（Ｔ３ｊ）—粘弹性流变阶段（Ｔ３ｗ）—后期前陆盆地演化阶段（Ｊ１－２），不同演化阶段前陆盆地沉降中心及前隆具纵向迁移的特征，同时，沉积盆地规模及拗陷幅度亦显示规律性变化。     

石炭系全球界线层型研究进展

早石炭世晚期开始的全球大规模构造运动和冈瓦纳大陆的冰川,为石炭系的地层划分和对比带来了困难。通过国际石炭系分会的努力,近年来石炭系各阶的界线层型研究取得了显著的进展。维宪阶已有了古生物学界线定义,即有孔虫Eoparastaffella simplex的首现,其余各阶也有了意向性的界线定义,它们均以牙形刺作为主导化石类群。其中,谢尔普霍夫阶以L ochriea ziegleri的首现为底界;莫斯科阶以Idiognathoides postsulcatus或Declinognathodus donetzianus的首现为底界;卡西莫夫阶的底界为Idiognathodus sagittalis的首现层位;格舍尔阶的底界是Idiognathodus simulator的首现层位     

二次离子质谱微区原位牙形石氧同位素分析及其在古海表水温记录中的应用

利用生物成因磷灰石的氧同位素信息能够重建古海水温度,尤其是牙形石,结合其在生物地层学上的研究,它的数据解释具有明确的古环境意义。牙形石微小的个体一直是其精确地球化学分析的限制因素。作为对牙形石微区原位氧同位素测试方法的探索研究,本文利用Cameca IMS--1280 型二次离子质谱仪对牙形石进行了微区原位氧同位素组成测试,通过分析得到可靠的古表层海水温度 ( SST) 记录,表明从晚二叠世到早三叠世SST 存在明显的上升过程。相比常规Ag3PO4 实验方法,牙形石微区原位氧同位素分析方法更加快速,而且能够有效避免牙形石易受到污染的部位,获得更为可靠的牙形石氧同位素组成的测定结果。     

桂西下三叠统牙形石序列的新认识

桂西下三叠统为海相沉积,根据其岩性、厚度、化石类型等诸方面的明显差异可以分为两种类型,即作登型和太平型。本文详细研究了桂西两种类型下三叠统的牙形石,认为不同类型的下三叠统具有不同的牙形石序列。在前人工作基础上,对桂西下三叠统牙形石序列予以重新厘定。补充了作登型下三叠统的Neogondolella carinta带;指出作登型下三叠统不宜建立Platyvillosus costatus带;在太平型下三叠统Dienerian阶—Spathian阶中新建了Pachvcladina erromera带,Platyvillosus costatus带,Pachycladina obliqua-Parachirognathus delicatulus带和Neospathodus triangularis带;指出了每一个牙形石带的性质并与作登型下三叠统牙形石序列进行了对比。本文还从事件地层学和生物地层学两个角度,对桂西二叠一三叠系界线提出了新的认识。