Late-quaternary vegetational dynamics and community stability reconsidered

Defining the spatial and temporal limits of vegetational processes such as migration and invasion of established communities is a prerequisite to evaluating the degree of stability in plant communities through the late Quaternary. The interpretation of changes in boundaries of major vegetation types over the past 20,000 yr offers a complementary view to that provided by migration maps for particular plant taxa. North of approximately 43°N in eastern North America, continual vegetational disequilibrium has resulted from climatic change, soil development, and species migrations during postglacial times. Between 33° and 39°N, stable full-glacial vegetation was replaced by a relatively unstable vegetation during late-glacial climatic amelioration; stable interglacial vegetation developed there after about 9000 yr B.P. Late-Quaternary vegetation has been in dynamic equilibrium, with a relatively constant flora, south of 33°N on upland interfluves along the northern Gulf Coastal Plain, peninsular Florida, and west-central Mexico.     

A palaeoecological study of the mites (Acari) from a Lateglacial deposit at Usselo (The Netherlands)

The distribution of the remains of mites in the Lateglacial (Late Weichselian) type section at Usselo (The Netherlands) is presented. The division of the sequence into (sub-)zones, based upon combined palaeo-botanical and palaeoentomological studies, is shown, as well as the approximate age based on ¹⁴C determinations. The present habitats of the species are in agreement with the reconstruction of the local vegetational succession Four species of mites, three oribatids and one uropodid, have not previously been recorded as subfossils.     

Modern pollen assemblages and vegetational history of the moraines of the Klutlan Glacier and its surroundings,Yukon Territory,Canada

Modern pollen assemblages have been studied from surficial lake muds and moss polsters collected from five vegetated ice-cored moraines of the Klutlan Glacier. The youngest vegetated moraine (K-II) is characterized by high pollen values for Salix and Hedysarum, K-III by high Salix and Shepherdia canadensis and low Hedysarum and Picea, K-IV by high Betula, Salix, and Shepherdia, and K-V and the Harris Creek moraine (HCM) by high Picea. Variations are summarized by canonical variates analysis. A percentage pollen diagram from Gull Lake on the upland east of the glacier records vegetational development since the deposition of the White River volcanic ash 1220 ¹⁴C yr ago. An initial species-rich treeless vegetation was replaced by birch-alder-willow shrub-tundra, and this by open Picea glauca forest similar to present vegetation around the lake. Sites on HCM show two basic stratigraphies. Triangle Lake reflects vegetational succession from Salix-Shepherdia canadensis scrub similar to that on K-III today, through open Picea woodland of K-IV type, to closed Picea forests of K-V and HCM. Heart Lake and Cotton Pond reflect vegetational development following melting of ice underlying the spruce forests of HCM. These two types are summarized by positioning the fossil spectra on the first two canonical variate axes of the modern surface spectra.     

The present flora and vegetation of the moraines of the Klutlan Glacier,Yukon Territory,Canada: A study in plant succession

The flora and vegetation of six ice-cored moraines of the Klutlan Glacier were analyzed in 65 plots by European plant-sociological techniques. The age of each plot was estimated from annual growth rings of shrubs or trees in the plots. Nine major vegetation types are distinguished: Crepis nana, Dryas drummondii, Hedysarum mackenzii, Hedysarum-Salix, Salix-Shepherdia canadensis, Picea-Salix, Picea-Arctostaphylos, Picea-Ledum, and Picea-Rhytidium. These contain plants aged 2–6, 9–23, 10–20, 24–30, 32–58, 58–80, 96–178, 177–240, and >163- >339 yr, respectively. Six other vegetation types are described from windthrow areas, drainage channels, volcanic tephra slopes, lake margins, fens, and drained lakes. The major vegetation types reflect a vegetational succession related to moraine age and stability, with the Crepis nana type as the pioneer vegetation developing through the other vegetation types to the Picea-Rhytidium type on the oldest moraines. Changes in species diversity and soil development, particularly humus accumulation, parallel the vegetational succession. This succession differs from patterns of revegetation of deglaciated landscapes in Alaska and British Columbia today and in Minnesota in late-Wisconsin times because of differences in climate, plant migration, and local ecology.     

Late Wisconsin vegetational and climatic history at Kylen Lake, northeastern Minnesota

Kylen Lake, located within the Toimi drumlin field, is critically positioned in relation to Late Wisconsin glacial advances, for it lies between the areas covered by the Superior and St. Louis glacial lobes between 12,000 and 16,000 yr B.P. The pollen and plant-macrofossil record suggests the presence of open species-rich “tundra barrens” from 13,600 to 15,850 yr B.P. Small changes in percentages of Artemisia pollen between 14,300 and 13,600 yr B.P. appear to be artifacts of pollen-percentage data. Shrub-tundra with dwarf birch, willow, and Rhododendron lapponicum developed between 13,600 and 12,000 yr B.P. Black and white spruce and tamarack then expanded to form a vegetation not dissimilar to that of the modern forest-tundra ecotone of northern Canada. At 10,700 B.P. spruce and jack pine increased to form a mosaic dominated by jack pine and white spruce on dry sites and black spruce, tamarack, and deciduous trees such as elm and ash on moist fertile sites. At 9250 yr B.P. red pine and paper birch became dominant to form a vegetation that may have resembled the dry northern forests of Wisconsin today. The diagram terminates at 8410 ± 85 yr B.P. Climatic interpretation of this vegetational succession suggests a progressive increase in temperature since 14,300 yr B.P. This unidirectional trend in climate contrasts with the glacial history of the area. Hypotheses are presented to explain this lack of correspondence between pollen stratigraphy and glacial history. The preferred hypothesis is that the ice-margin fluctuations were controlled primarily by changes in winter snow accumulation in the source area of the glacier, whereas the vegetation and hence the pollen stratigraphy were controlled by climatic changes in front of the ice margin.     

Late Holocene vegetational history,human impact and pollen representativity variations in northern Cumbria,England

The comparison of pollen diagrams and their inferred vegetational histories are an important component of palaeoecological research. Radiocarbon-dated pollen profiles from three cores taken from two adjacent mires located in northern Cumbria, Bolton Fell Moss and Walton Moss, have been used to reconstruct the Late Holocene vegetation history between the Bronze Age and the present day. The profiles have been interpreted in the light of available archaeological and historical records and, although the pollen records are broadly similar, there are some notable differences between them, particularly during Iron Age and medieval times. Dissimilarities between the diagrams are explored numerically, and the statistical and palynological results are discussed in relation to pollen representativity. The results suggest that it may be advantageous to construct more than one pollen diagram from a mire, or even adjacent mires, as extra-local pollen may be a more important part of the pollen rain than previously envisaged. © 1998 John Wiley & Sons, Ltd.     

鄂尔多斯高原植被生态分区及其水文地质意义

保护生态环境是鄂尔多斯高原地下水可持续开发利用中亟待解决的问题。在查清植被生态群落分布特征的基础上,通过分析植被演替的自然驱动因素,将鄂尔多斯高原的植被生态划分为基岩台地植被生态水文地质区、沙盖基岩植被生态水文地质区、沙地植被生态水文地质区和滩地植被生态水文地质区。结果表明,气象和地形地貌控制着区内植被生态的总体分布格局,包气带和地下水因素控制着植被生态的演替,沙地植被生态和滩地植被生态的生长发育与地下水关系密切,是地下水开发时保护植被生态的靶区。     

Differentiation of MIS 9 and MIS 11 in the continental record: vegetational,faunal, aminostratigraphic and sea-level evidence from coastal sites in Essex,UK

Multidisciplinary investigations of the vegetational, faunal and sea-level history inferred from the infills of buried channels on the coast of eastern Essex have a direct bearing on the differentiation of MIS 11 and MIS 9 in continental records. New data are presented from Cudmore Grove, an important site on Mersea Island that can be linked to the terrace sequence of the River Thames. The vegetational history has been reconstructed from a pollen sequence covering much of the interglacial represented. The temperate nature of the climate is apparent from a range of fossil groups, including plant remains, vertebrates (especially the rich herpetofauna), molluscs and beetles, which all have strong thermophilous components. The beetle data have been used to derive a Mutual Climatic Range reconstruction, suggesting that mean July temperatures were about 2 °C warmer than modern values for southeast England, whereas mean January temperatures may have been slightly colder. The sea-level history has been reconstructed from the molluscs, ostracods and especially the diatoms, which indicate that the marine transgression occurred considerably earlier in the interglacial cycle than at the neighbouring Hoxnian site at Clacton. There are a number of palynological similarities between the sequence at Cudmore Grove and Clacton, especially the presence of Abies and the occurrence of Azolla filiculoides megaspores. Moreover, both sites have yielded Palaeolithic archaeology, indeed the latter is the type site of the Clactonian (flake-and-core) industry. However, the sites can be differentiated on the basis of mammalian biostratigraphy, new aminostratigraphic data, as well as the differences in the sea-level history. The combined evidence suggests that the infill of the channel at Cudmore Grove accumulated during MIS 9, whereas the deposits at Clacton formed during MIS 11. The infill of a much later channel, yielding non-marine molluscs and vertebrates including Hippopotamus, appears to have formed during the Ipswichian (MIS 5e). This evidence is compared with other important sites of late Middle Pleistocene age in Britain and elsewhere on the continent and the importance of a multidisciplinary approach is stressed.     

Late Weichselian and early Flandrian vegetational history of Varanger peninsula, northeast Norway

Relative a absolute (pollen concentration) diagrams are presented from Bergebyvatnet, Holmfjellvatnet and Stjernevatnet on Varanger peninsula. All three sites are outside the younger Dryas (Main substage) moraines and the pollen assemblage zones are correlated biostratigraphically with chronozones from Allerød to Middle Flandrian. Radiocarbon dates from Bergebyvatnet appear to have been affected by hard water error, but dates from the other two sites agree will with the inferred chronostratigraphy. Pollen diagrams from Varanger peninsula suggest broadly similar vegetational histories, the longest record beign that from Østcrvatnet (H. C. Prentice 1981, Boreas , Vol. 10, pp. 53–70). Open tundra-like conditions prevailed throughout the Late Weichselian, with Salix dominance interrupted by unstable vegetation with abundant Artemisia during the Older and Younger Dryas zones. Major vegetational and floristic changes began just before 10,000 B.P., the rapid scquence from herb pollen flora was rich and varied, including a mixture of floristic clements similar to that found during the Late Weichselian in southern Scandinavia. Basiphilous herbs were particularly abundant at Østervatnet and Bergebyvtnet. Betula nand and species of Ericales became locallydominant just before the full establishment of B. pubescens , which rapidly spread beyond its present limit. Later immigrants included Alnus incana; Juniperus communis ; and Pinus sylvestris , which reached the south western part.     

Relative timing of last glacial maximum and late-glacial events in the central tropical Andes

Whether or not tropical climate fluctuated in synchrony with global events during the Late Pleistocene is a key problem in climate research. However, the timing of past climate changes in the tropics remains controversial, with a number of recent studies reporting that tropical ice age climate is out of phase with global events. Here, we present geomorphic evidence and an in-situ cosmogenic ³He surface-exposure chronology from Nevado Coropuna, southern Peru, showing that glaciers underwent at least two significant advances during the Late Pleistocene prior to Holocene warming. Comparison of our glacial-geomorphic map at Nevado Coropuna to mid-latitude reconstructions yields a striking similarity between Last Glacial Maximum (LGM) and Late-Glacial sequences in tropical and temperate regions.Exposure ages constraining the maximum and end of the older advance at Nevado Coropuna range between 24.5 and 25.3 ka, and between 16.7 and 21.1 ka, respectively, depending on the cosmogenic production rate scaling model used. Similarly, the mean age of the younger event ranges from 10 to 13 ka. This implies that (1) the LGM and the onset of deglaciation in southern Peru occurred no earlier than at higher latitudes and (2) that a significant Late-Glacial event occurred, most likely prior to the Holocene, coherent with the glacial record from mid and high latitudes. The time elapsed between the end of the LGM and the Late-Glacial event at Nevado Coropuna is independent of scaling model and matches the period between the LGM termination and Late-Glacial reversal in classic mid-latitude records, suggesting that these events in both tropical and temperate regions were in phase.     

Reconstruction of Late-Glacial lake-level fluctuations in The Netherlands based on palaeobotanical analyses, geochemical results and pollen-density data

The Late-Glacial infills of three lake successions in The Netherlands have been analysed for their palaeobotanical content (pollen, macroremains) as well as for their physical and chemical characteristics. The macro-botanical analysis in association with the pollen analysis permitted the reconstruction of relative lake level fluctuations for each lake. There appeared to be good agreement between the three lake successions. Supporting evidence for the presence of alternating dry and wet phases was found in the geochemical results. The termination of the Bölling biozone coincides with a lowering of the lake levels that persisted throughout the Older Dryas. Water levels restore at the beginning of the Allerod biozone. Several fluctuations are recognized during the Allerod. The end of the Alleröd is characterized by a second major lowering in the lake levels, leading to an interruption in the sedimentation in cores from the fringe zone. Water depth increases at the beginning of the Younger Dryas biozone. A third major minimum in the lake levels is reached towards the end of the Younger Dryas biozone, probably explaining the recurrent hiatus at the transition to the Holocene record. Pollen concentration data in association with chronostratigraphical data provided the construction of sedimentation rate curves. Minima in these sedimentation rate curves appeared to correlate with periods of low lake level. The loss on ignition curve in association with the sedimentation rate data allowed an approximation of the organic production of the lakes involved. It revealed that the darker appearance of the Allerödin the lithologycan be ascribed mainly to a retardationin the sedimentation rate and not to a major increase in the organic production of the lakes.     

Late Quaternary vegetational and climatic change in the Popayán region,southern Colombian Andes

Late Pleistocene and Holocene vegetational and climatic change have been studied palynologically at a site at 1750 m elevation in the subandean vegetation belt near Popayán, in the southern Colombian Andes. Time control on the pollen record is based on six AMS ¹⁴C ages, ranging from possibly Middle Pleniglacial time (around 50000 yr BP) to 1092 ± 44 yr BP. Because of the presence of two hiatuses only the Middle Pleniglacial and Late Holocene periods (the last 2300 yr BP) are represented. Pollen data indicate the presence of closed subandean forest during glacial time. Changes in the contribution of pollen originating from the uppermost and lowermost subandean forest belts, changes in the contribution of a number of other subandean forest taxa, and changes in species composition between the three pollen zones, suggest that the climate during the Middle Pleniglacial was markedly colder, and perhaps also wetter, than during the Late Holocene. Pollen assemblages from the Late Holocene indicate that the landscape has been affected by deforestation and agriculture since at least 2300 yr BP, but that human impact decreased in the last 780 yr BP. © 1998 John Wiley & Sons, Ltd.     

Roof Shale Flora of Coal Seam 6 from the Asselian（Lower Permian）Taiyuan Formation of the Wuda Coalfield, Inner Mongolia and Its Ecostratigraphic Significance

Roof shale floras help understand the transition of vegetational landscapes from a peat–forming environment to a clastic one, but are seldom investigated in the Cathaysian province(typical present day China and East Asia). Here the roof shale flora of Coal Seam 6 from the lower Permian Taiyuan Formation of the Wuda Coalfield, Inner Mongolia, is systematically described and its ecostratigraphic and phytogeographic implications are discussed. The flora is composed of seven plant groups, including Lepidodendrales, Medullosales, Gigantopteridales, Peltaspermales, Noeggerathiales, Cordaitales and Cycadales. Many of these taxa are also documented in Euramerica, and the floral composition indicates a more intimate relationship between Cathaysia and Euramerica during the Cisuralian than previously thought. However, there are few genera and species in common with those of the underlying peat–forming flora. Moreover, the flora is hardly comparable with the commonly known flora of the Taiyuan Formation. Such assemblage differences are very likely due to changes of taphonomic and environmental phases, rather than evolutionary floral succession. For a full and more precise understanding of the floral composition, succession and the floristic discrepancy in different depositional environments, an ecostratigraphic investigation on the Late Palaeozoic of Cathaysia is advocated.     

Modern vegetational distribution and pollen dispersal study within Gangotri glacier valley, Garhwal Himalaya

Modern pollen-vegetation relationship has been analyzed in respect to vegetational distribution and pollen dispersal from sub-alpine and alpine ecological regimes within the higher reaches of the Gangotri glacier valley. The results in general cope with the local ecological conditions of the sites within the valley. Above the tree line limits, the pollen frequency of arboreal taxa is found lower than those of non-arboreal taxa. In contrast the pine-birch forest area at Chirbasa represented the frequency of arboreal pollen (AP) more than that of non-arboreal pollen (NAP) due to over representation of Pinus wallichiana (conifer). Pollen grains of extra-local arboreal elements, mostly conifers have also been recorded here with fair amount of temperate broad-leaved arboreal taxa viz. Quercus, Alnus, Corylus, Carpinus, Ulmus, Juglans etc. that have been transported by the upthermic winds from their growing limits at lower altitudes to the study sites at higher altitudes. Non-arboreal taxa viz. marshy/aquatic, steppe and of other herbaceaous taxa, represents the existing ecological regimes of their respective sites within the valley. Differential pollen preservation has been observed between the moss cushion and sediment samples from the same site that may be due to the direct exposure of moss cushions to air, thus showing more concentration of wind pollinated pollen-spores as compared to surface sediments.     

Last glacial fire regime variability in western France inferred from microcharcoal preserved in core MD04-2845, Bay of Biscay

High resolution multiproxy analysis (microcharcoal, pollen, organic carbon, Neogloboquadrina pachyderma (s), ice rafted debris) of the deep-sea record MD04-2845 (Bay of Biscay) provides new insights for understanding mechanisms of fire regime variability of the last glacial period in western France. Fire regime of western France closely follows Dansgaard–Oeschger climatic variability and presents the same pattern than that of southwestern Iberia, namely low fire regime associated with open vegetation during stadials including Heinrich events, and high fire regime associated with open forest during interstadials. This supports a regional climatic control on fire regime for western Europe through fuel availability for the last glacial period. Additionally, each of Heinrich events 6, 5 and 4 is characterised by three episodes of fire regime, with a high regime bracketed by lower fire regime episodes, related to vegetational succession and complex environmental condition changes.     

Occurrence of larch (Larix) in Fennoscandia during the Eemian interglacial and the Brørup interstadial according to pollen analytical data

The occurrence of pollen and macrofossils of larch in Eemian deposits in northern Finland indicates that this species must have grown in the area during the last interglacial. Lark spread to Finland from the east, its date of arrival being deducible from the general vegetational succession. It probably did not grow in central or southern Finland during the interglacial, but is thought to have extended fairly far south in Sweden and Noway along the Fennoscandian mountain range. The Lark pollen found at the upper boundary of the interglacial deposits at Margreteberg and Stenberget in southern Sweden may suggest that it did reach southern Sweden by the very end of the Eemian, but it cannot be said for certain whethcr this pollen represents an influx of Larix from the north or from Central Europe.     

Surge moraines of the Klutlan Glacier,Yukon Territory,Canada: Origin,wastage, vegetation succession,lake development,and application to the late-glacial of Minnesota

The Klutlan Glacier in the St. Elias Mountains of the Yukon Territory has surged repeatedly during the last few hundred years, and its drift-covered stagnant ice provides an analog for the downwastage, landform development, vegetational succession, and lake formation on Late Wisconsin moraines of Minnesota. Melting of the buried ice caused collapse of the drift mantle and the formation of lakes, which become filled with sediment that slumps in from receding ice walls. Topographic reversals are common, as the sediment cover of drained lakes inhibits local under-melting, and collapse occurs elsewhere. As the drift mantle thickens the land surface becomes stabilized and pioneer herbs are succeeded by shrubs and then by white spruce. The oldest moraines (600–1200 yr old) have a multiple-generation spruce forest, yet melting of buried ice still locally forms young lakes. Cores of organic sediment from the oldest lakes contain a stratigraphic sequence of pollen, diatoms and cladocerans that record the early stages in lake and landscape succession.     

Pollen indicators of land-use change in southern Connecticut

Changes in fossil pollen assemblages from a 2-m core from Linsley Pond, North Branford, Connecticut, are compared with historically documented land use changes in the lake watershed. Dating with ²¹⁰Pb and ¹⁴C reveals two sedimentation rate changes in the core which are associated with the arrival of European farmers; the building of cabins and suburban housing subdivisions on the lake shore. At European settlement in 1700 Ambrosia and Rumex pollen first appear, Gramineae-type pollen increases, and Tsuga decreases. Just before the beginning of agricultural disturbance Fagus pollen declines. The chestnut blight of 1913 causes a reduction of Castanea pollen and a subsequent vegetational succession through Betula to Quercus. The sedimentation rate determined by the chestnut blight horizon is consistent with the rate deduced from ²¹⁰Pb analysis.     

Paleoecology of Middle Pennsylvanian-age peat-swamp plants in Herrin coal,Kentucky, U.S.A.

To develop a method for quantifying the vegetation of Pennsylvania-age coal beds, of four coal-ball (permineralized peat) profiles and four coal column samples from the Herrin coal bed (Kentucky No. 11) Carbondale Formation in western Kentucky were compared. An estimated 89.5% of the coal can be identified botanically. Compaction ratios for individual tissues were estimated using point counts of organic matter in coal balls. The estimated abundances of major plant groups (lycopods, ferns, sphenopsids, and pteridosperms) in coal balls differ by less than 10% compared to coal after accounting for differential compaction of plant tissues. Standard deviations in taxonomic and maceral composition among coal columns are generally less than 2%.Consistent differences in botanical composition were found between benches showing that the method is consistent when applied to sufficient thicknesses of coal. It was not possible to make fine-scale correlations within the coal bed using the vegetational data; either the flora varied considerably from place to place or the method of quantification is unreliable for small increments of coal (5 cm or less).In the coal, pteridosperm abundance is positively correlated with underlying shale partings. This correlation suggests that pteridosperms are favored either by higher nutrient levels or disturbance.In the third of four benches in the Herrin coal bed, a succession from Sigillaria-containing zones to zones dominated by Lepidophloios hallii is interpreted as a shift towards wetter conditions. In the other benches, the main factors controlling the taxonomic composition appear to have been the relative abundance of nutrients and/or the frequency of disturbance as indicated by the relative abundance of partings.Criteria for distinguishing between domed and planar swamps are discussed. These include: distribution of partings, type of plant succession, and changes in plant diversity, average plant size, preservational quality and sporinite content. The infrequency of partings in bench C suggests a peat dome developed while the peat of that bench was accumulating but other evidence either fails to support the development of a peat dome or is ambiguous. The maceral composition resembles those of other Carboniferous coals which are thought to have formed from planar peat swamps.Formation of fusain bands appears to be associated with processes occurring above the peat surface, such as burning or prolonged oxidative exposure. Oxidation of accumulated peat is unlikely because fusain bands rarely include more than a single plant.     

Late-glacial vegetational and climatic history of the Allegheny Plateau and the Till Plains of Ohio and Indiana, U.S.A.

Shane, Linda C. K. 1987 03 01: Late-glacial vegetational and climatic history of the Allegheny Plateau and the Till Plains of Ohio and Indiana, U.S.A. Boreas , Vol. 16, pp. 1–20. Oslo. ISSN 0300–9483.
Pollen evidence from the Allegheny Plateau and the Till Plains south of the Great Lakes shows marked post-glacial vegetation gradients. C. 15,500–11,000 B.P .: On the Plateau, spruce forest was rapidly established, persisted for 2,000 years, then began a gradual change to deciduous-conifer forest. On the Till Plains, open spruce forest tundra closed slowly over 1,000 years, declined rapidly c . 13,500 B.P., and a deciduous open woodland developed. C. 11,000–10,300 B.P .: On the Till Plains, a brief cooling is recorded by increases in the abundance of spruce and fir, contemporaneous with the European Younger Dryas. No clear change is seen on the Plateau. 10,300–4,000 B.P .: Warming and/or drying occurred in both areas, as hemlock and jack/red pine trees immigrated, followed by white pine. Conifers disappeared from the Till Plains by 9,800 B.P., but pine and hemlock trees may have persisted on the Plateau. After 10,000 B.P. mixed deciduous forest was established across the entire region. Between 8,000 and 4,000 B.P., further warming/drying is indicated on the Till Plains with development of open oak forest and lake shallowing, and on the Plateau by a minor increase in herbs, lake shallowing, and reduction in pine.