Tepsankumpu revisited — pollen evidence of stable Eemian climates in Finnish Lapland

Several till-covered organic deposits, principally lake gyttja, in Finnish Lapland have been correlated with the last (i.e. Eemian) interglacial on the basis of their lithostratigraphic position and pollen stratigraphy. Most of the sequences are short, but together with three longer sequences from Finnish Lapland and one from Swedish Lapland (Leveäniemi) they provide a complete picture of Eemian vegetational and climatic development. The Tepsankumpu site was revisited, and the till-covered thick freshwater gyttja deposit was studied in detail for pollen in order to search for signals of rapid climatic fluctuations postulated for the earlier part of the Eemian on the basis of Greenland ice core studies. The Eemian pollen stratigraphy in Finnish Lapland closely resembles the Holocene pollen stratigraphy of the area. The abundance of spruce and alder pollen suggests, however, more northerly limits for forest vegetation zones during the Eemian than during the Holocene. Oak also grew closer to Lapland, indicating a wanner climate than during the Holocene climatic optimum. The Tepsankumpu pollen stratigraphy indicates climatic stability over the entire time-span it covers, i.e. the major part of the interglacial. This finding is in conflict with results from Greenland GRIP ice core studies and interpretations of some Continental European Eemian pollen diagrams.     

Modern pollen assemblages from surface lake sediments and their environmental implications on the southwestern Tibetan Plateau

The relationships amongst modern pollen assemblages, vegetation, climate and human activity are the basis for reconstructing palaeoenvironmental changes using pollen records. It is important to determine these relationships at regional scales due to the development of vegetation under different climatic conditions and human activities. In this paper, we report on an analysis of modern pollen assemblages of 31 surface lake samples from 31 lakes (one sample per lake) on the southwestern Tibetan Plateau where the knowledge of modern pollen and their relationships with vegetation, climate and human activities is insufficient. The region includes five vegetation zones: sub‐alpine shrub steppe, alpine steppe, alpine meadow and steppe ecotone, mountain desert and alpine desert. The lakes span a wide range of mean annual precipitation (50–500 mm) and mean annual temperature (?8 to 6 °C). Modern pollen assemblages from our samples mainly consist of herb taxa (Artemisia, Cyperaceae, Poaceae, Chenopodiaceae, etc.) and some tree taxa (Pinus, Fagaceae, Alnus, etc.). The results indicate that modern pollen assemblages are able to reflect the main vegetation distribution. Redundancy analysis for the main pollen types and environmental variables shows that precipitation is the leading factor that influences the pollen distribution in the study area with the first axis capturing 13.7% of the variance in the pollen data set. The Artemisia/Chenopodiaceae ratio is valid for separating the desert component (<2) from the steppe and other vegetation zones (>2) but is unable to distinguish moisture variations. The Artemisia/Cyperaceae ratio is able to identify meadows (<1) and steppes (>1) and can be used as a moisture index on the southwestern Tibetan Plateau. Our results show that an appropriate range is needed for a modern pollen data set in order to perform pollen‐based quantitative climate reconstructions in one region. It is essential to perform modern studies before using pollen ratios to reconstruct palaeovegetation and palaeoclimate at a regional scale.     

Multivariate statistical estimates of Holocene vegetation and climate change,forest-tundra transition zone,NWT, Canada

Newly derived fossil pollen data were obtained from four sites along a transect from the boreal forest limit into tundra in the eastern Northwest Territories. Multivariate statistical analyses were employed to interpret the pollen assemblages. Transfer functions were constructed between the pollen data and climatic data, and paleoclimatic estimates were derived. The objective nature of the reconstructions provides an independent verification of the general outlines of the chronology of tree-line movements during the mid- and late-Holocene as established in previous paleosol and pollen studies. Boreal forest extended to approximately 62°N, associated with mean July temperatures 1 to 3°C above modern means, from at least 5500 to 3700 yr B.P. Although a major episode of southward displacement of tree line at about 3700 yr B.P. is recorded, later events are not clearly represented. Considerations of the statistics, the time scales, and the nature of the pollen rain suggest only conservative interpretations of the results are possible. It is suggested that the pollen sites may have been sensitive recorders of regional vegetation change only when they were near the ecotone, corresponding to a climatic threshold.     

Late Quaternary vegetational and climatic change in the Popayán region,southern Colombian Andes

Late Pleistocene and Holocene vegetational and climatic change have been studied palynologically at a site at 1750 m elevation in the subandean vegetation belt near Popayán, in the southern Colombian Andes. Time control on the pollen record is based on six AMS ¹⁴C ages, ranging from possibly Middle Pleniglacial time (around 50000 yr BP) to 1092 ± 44 yr BP. Because of the presence of two hiatuses only the Middle Pleniglacial and Late Holocene periods (the last 2300 yr BP) are represented. Pollen data indicate the presence of closed subandean forest during glacial time. Changes in the contribution of pollen originating from the uppermost and lowermost subandean forest belts, changes in the contribution of a number of other subandean forest taxa, and changes in species composition between the three pollen zones, suggest that the climate during the Middle Pleniglacial was markedly colder, and perhaps also wetter, than during the Late Holocene. Pollen assemblages from the Late Holocene indicate that the landscape has been affected by deforestation and agriculture since at least 2300 yr BP, but that human impact decreased in the last 780 yr BP. © 1998 John Wiley & Sons, Ltd.     

Late Wisconsin vegetational and climatic history at Kylen Lake, northeastern Minnesota

Kylen Lake, located within the Toimi drumlin field, is critically positioned in relation to Late Wisconsin glacial advances, for it lies between the areas covered by the Superior and St. Louis glacial lobes between 12,000 and 16,000 yr B.P. The pollen and plant-macrofossil record suggests the presence of open species-rich “tundra barrens” from 13,600 to 15,850 yr B.P. Small changes in percentages of Artemisia pollen between 14,300 and 13,600 yr B.P. appear to be artifacts of pollen-percentage data. Shrub-tundra with dwarf birch, willow, and Rhododendron lapponicum developed between 13,600 and 12,000 yr B.P. Black and white spruce and tamarack then expanded to form a vegetation not dissimilar to that of the modern forest-tundra ecotone of northern Canada. At 10,700 B.P. spruce and jack pine increased to form a mosaic dominated by jack pine and white spruce on dry sites and black spruce, tamarack, and deciduous trees such as elm and ash on moist fertile sites. At 9250 yr B.P. red pine and paper birch became dominant to form a vegetation that may have resembled the dry northern forests of Wisconsin today. The diagram terminates at 8410 ± 85 yr B.P. Climatic interpretation of this vegetational succession suggests a progressive increase in temperature since 14,300 yr B.P. This unidirectional trend in climate contrasts with the glacial history of the area. Hypotheses are presented to explain this lack of correspondence between pollen stratigraphy and glacial history. The preferred hypothesis is that the ice-margin fluctuations were controlled primarily by changes in winter snow accumulation in the source area of the glacier, whereas the vegetation and hence the pollen stratigraphy were controlled by climatic changes in front of the ice margin.     

Late Weichselian and early Flandrian vegetational history of Varanger peninsula, northeast Norway

Relative a absolute (pollen concentration) diagrams are presented from Bergebyvatnet, Holmfjellvatnet and Stjernevatnet on Varanger peninsula. All three sites are outside the younger Dryas (Main substage) moraines and the pollen assemblage zones are correlated biostratigraphically with chronozones from Allerød to Middle Flandrian. Radiocarbon dates from Bergebyvatnet appear to have been affected by hard water error, but dates from the other two sites agree will with the inferred chronostratigraphy. Pollen diagrams from Varanger peninsula suggest broadly similar vegetational histories, the longest record beign that from Østcrvatnet (H. C. Prentice 1981, Boreas , Vol. 10, pp. 53–70). Open tundra-like conditions prevailed throughout the Late Weichselian, with Salix dominance interrupted by unstable vegetation with abundant Artemisia during the Older and Younger Dryas zones. Major vegetational and floristic changes began just before 10,000 B.P., the rapid scquence from herb pollen flora was rich and varied, including a mixture of floristic clements similar to that found during the Late Weichselian in southern Scandinavia. Basiphilous herbs were particularly abundant at Østervatnet and Bergebyvtnet. Betula nand and species of Ericales became locallydominant just before the full establishment of B. pubescens , which rapidly spread beyond its present limit. Later immigrants included Alnus incana; Juniperus communis ; and Pinus sylvestris , which reached the south western part.     

Modern pollen spectra from lake sediments in Finland and Finnmark, north Norway

Absolute (APF) and relative surface pollen spectra from 67 moderate-sized lakes in Finland and Finnmark are presented as pollen diagrams and isopoll maps and analysed by multidimensional scaling. The results are applicable to problems of regional vegetational history through comparisons of pollen spectra and the use of R-values. It is shown that relative spectra can discriminate the major vegetational regions. APFs give better resolution among northern forest regions and low APFs characterize tundra. Gradients of tree-pollen percentages in south and central Finland are related quantitatively to the major forest gradients, which are azonal, mainly related to soils, and affected but not obscured by human influence. Forest clearance is reflected in frequencies of various non-tree pollen types. Non-tree pollen and spores also provide vegetational and climatic information in the far north.     

阳原西堡剖面的孢粉分析

通过对阳原西堡剖面的孢粉分析,得到16000年以来该地区植被与环境发展的历史。约16000~11000aBP期间,植被是以蒿和藜为主的干旱草原景观,气候寒冷而干燥;约11000~9000aBP期间,植被有一定的增长,气候变得温凉稍湿;约9000~4800aBP期间,气温上升湿度下降,植被呈现疏林草原景观;距今4800aBP以来,植被虽然仍以草原为主,但低花粉浓度指示环境条件又趋于恶劣,气候进一步向暖干方向发展。      

Palynological Evidence for Climatic Change and Human Activity during the Holocene on Minorca (Balearic Islands)

Four pollen diagrams from Minorca (Balearic Islands) have been correlated with other previously studied sequences from Majorca and Minorca to define a Holocene landscape sequence for the region from 8000 yr B.P. to the present. The lower part of the pollen diagrams reflects a climatic phase with more rain and less-marked seasonality than today. Significant quantities ofCorylus, Buxus,and mesophilous taxa are found. In the middle part, between 5000 and 4000 yr B.P., a strong change is recorded in composition and structure of the vegetational landscape, with vegetation appearing that was adapted to Mediterranean conditions. This episode coincided with the first human colonization of the island and also with a widespread climatic change in the western Mediterranean region. The change in taxa was complex and some sclerophyllous taxa suchOleaplayed an important role in the transformation of the landscape physiognomy from the mid-Holocene until the present. Although human activities have removed much of the Mediterranean vegetation on the Balearic Islands, it seems clear that the changes have been brought about, in part, by increasing dryness.     

The dynamics of chironomid assemblages and vegetation during the Late Quaternary at Laguna Facil, Chonos Archipelago, southern Chile

We compare high-resolution pollen and chironomid records from the last 15,000 yr in Laguna Facil, southern Chile. Major vegetation and chironomid changes are recorded between ca 14,900 and 14,700 cal. yr BP. During the Lateglacial, changes in the chironomid stratigraphy lag behind changes in the pollen stratigraphy suggesting that the chironomids are responding to changes in the tree canopy or in soil chemistry brought about by vegetational development. At about 7200 cal. yr BP there is a change in the chironomid stratigraphy in advance of changes in the vegetation. This suggests that the response is to regional climatic change. The relatively close correlation of the chironomid and pollen stratigraphies with changes in charcoal concentrations also implicates the importance of fire and/or vulcanism in influencing the dynamics of forest and limnological systems. There is no clear evidence of cooling during the Younger Dryas chronozone in Laguna Facil.     

川北若尔盖高原玛曲-红原一带表土中孢粉组合的特征

通过对四川若尔盖高原玛曲—红原一带20个表土孢粉样的分析,研究了该区域表土孢粉组合与现代植被的关系。结果显示,玛曲—红原一带表土的孢粉组合基本反映了现代植被的整体特征,与现代植被在植被类型、植被带的空间排列、主要建群种属和优势种属等方面基本一致。依据孢粉的百分含量,可以划分为亚高山灌丛植被区,亚高山灌丛、亚高山草甸植被区和高寒草甸、草原植被区3个植被带。表土花粉的代表性因种属的不同而各有差异,松属和菊科花粉具超代表性,云杉属花粉的代表性较好,禾本科和莎草科具低代表性,而桦木属则不具有超代表性,可能与桦木属花粉不易搬运有关。3个植被区中的表土孢粉的种类、比例与现代植被均存在一定的差别,这可能与植被退化有关。     

Changing patterns in the Holocene pollen record of northeastern North America: A mapped summary

By mapping the data from 62 radiocarbon-dated pollen diagrams, this paper illustrates the Holocene history of four major vegetational regions in northeastern North America. Isopoll maps, difference maps, and isochrone maps are used in order to examine the changing patterns within the data set and to study broad-scale and long-term vegetational dynamics. Isopoll maps show the distributions of spruce (Picea), pine (Pinus), oak (Quercus), herb (nonarboreal pollen groups excluding Cyperaceae), and birch + maple + beech + hemlock (Betula, Acer, Fagus, Tsuga) pollen at specified times from 11,000 BP to present. Difference maps were constructed by subtracting successive isopoll maps and illustrate the changing patterns of pollen abundances from one time to the next. The isochrone maps portray the movement of ecotones and range limits by showing their positions at a sequence of times during the Holocene. After 11,000 BP, the broad region over which spruce pollen had dominated progressively shrank as the boreal forest zone was compressed between the retreating ice margin and the rapidly westward and northward expanding region where pine was the predominant pollen type. Simultaneously, the oak-pollen-dominated deciduous forest moved up from the south and the prairie expanded eastward. By 7000 BP, the prairie had attained its maximum eastward extent with the period of its most rapid expansion evident between 10,000 and 9000 BP. Many of the trends of the early Holocene were reversed after 7000 BP with the prairie retreating westward and the boreal and other zones edging southward. In the last 500 years, man's impact on the vegetation is clearly visible, especially in the greatly expanded region dominated by herb pollen. The large scale changes before 7000 BP probably reflect shifts in the macroclimatic patterns that were themselves being modified by the retreat and disintegration of the Laurentide ice sheet. Subsequent changes in the pollen and vegetation were less dramatic than those of the early Holocene.     

Late-quaternary vegetational dynamics and community stability reconsidered

Defining the spatial and temporal limits of vegetational processes such as migration and invasion of established communities is a prerequisite to evaluating the degree of stability in plant communities through the late Quaternary. The interpretation of changes in boundaries of major vegetation types over the past 20,000 yr offers a complementary view to that provided by migration maps for particular plant taxa. North of approximately 43°N in eastern North America, continual vegetational disequilibrium has resulted from climatic change, soil development, and species migrations during postglacial times. Between 33° and 39°N, stable full-glacial vegetation was replaced by a relatively unstable vegetation during late-glacial climatic amelioration; stable interglacial vegetation developed there after about 9000 yr B.P. Late-Quaternary vegetation has been in dynamic equilibrium, with a relatively constant flora, south of 33°N on upland interfluves along the northern Gulf Coastal Plain, peninsular Florida, and west-central Mexico.     

Vegetation and associated environments during the past 14,000 years near Moulton Pond,Maine

Pollen influx and percentage diagrams were prepared from an 11.4 m core from Moulton Pond, Maine. The pond basin was deglaciated about 14,000 y. a., after which it was located on an island in a sea of subarctic character until about 12,400 y. a. when the surrounding area emerged from the sea. The terrestrial vegetation was tundra until about 10,000 y. a. A change in the tundra vegetation is synchronous with the emergence from the sea, but synchroneity with the Pineo Ridge glacial readvance, which reached its maximum 50 km to the east of the pond about 12,700 y. a., is also possible because of imprecision in the dating. Comparisons of the Moulton Pond results with late-glacial pollen sequences elsewhere in eastern United States and adjacent Canada reveal a lack of synchroneity in vegetational changes casting doubt on claims of major broad-scale climatic shifts over the entire area.The tundra period at Moulton Pond ended with a transition of a few hundred years to partly open, relatively xeric forests of low diversity dominated by white pine, oak, and birch trees. There was no intervening boreal forest. In the postglacial period the vegetation was continually changing, including in the early portion a series of immigrations of temperate tree taxa which later became important in the forests. The transient nature of these assemblages is further indicated by their differences from the closest modern analogs. From about 7100 y. a. until settlement by Europeans 200 y. a., the forests were closed. A major decline of conifers centering about 4700 y. a. was followed by maxima of mesic hardwoods about a thousand years later. In the most recent 2000 yr, the pollen record suggests greater environmental severity, evidenced by increasing spruce. But for the entire postglacial period, the closest modern vegetational analogs are all in the conifer-hardwood region. Much of the postglacial pollen sequence is inexplicable in climatic terms, as evidenced by nonsynchronous behavior of hemlock and beech.The pollen influx diagram is useful for distinguishing tundra from forest, but for the postglacial period it is difficult to interpret. Pollen influx data are strongly affected by shifts in the pattern of sedimentation in lakes. We propose that such shifts account for the major changes in influx in mid- and late-postglacial time at Moulton Pond and at Rogers Lake, Connecticut. This complicates the interpretation of influx data which otherwise are superior to percentage data.     

Lake Neor reveals how mountain vegetation responded to 7000 years of hydroclimate variability in northwestern Iran

Palynological and geochemical analyses provide valuable information about modern and past climatic regimes and vegetation. The impact of climate and humans on past vegetation in the semi-arid areas of northwestern Iran has received increased interest in the wake of warming temperatures in the Middle East. Palynological and down-core XRF elemental abundances from a peat core from Lake Neor enabled a reconstruction of vegetational changes of the past 7000 years over the highlands of northwestern Iran. Periods of increased arboreal pollen (AP) types and high (Artemisia + Poaceae)/Chenopodiaceae ratios along with low titanium abundances, high percentages of total organic carbon, more negative δD values, and higher carbon accumulation rates suggest a relatively wet climate. These conditions have persisted during the periods 6700–6200, 5200–4450 and 3200–2200 cal a bp. The overall low AP values, substantial rise of Chenopodiaceae, high Ti abundances and low values of palaeo-redox proxies, are all evidences of a drier climate, as has been reconstructed for the periods 6200–5200 and 4030–3150 cal a bp and the last 2200 years. An important feature of the last centuries is the increase of anthropogenic and pastoral indicator pollen types. Our results may provide basic data to predict future trends in vegetation dynamics under future climate change in western Asia.     

Early human impact in the forest ecotone of southern High Asia (Hindu Kush, Himalaya)

The vegetation of the treeline ecotone of the southern declivity of arid High Asia (Hindu Kush, northern areas of Pakistan; Himalaya, northern central Nepal) is dominated by hedgehog-like open dwarf shrublands of thorny cushions. Since climatically sensitive ecotones are always also sensitive to human impact, the question arises whether the current lack of forests is a result of the Subboreal climate decline or of human impact. Due to inadequate knowledge of the pollen flora and of ecological indicator values of the plants, pollen analyses in High Asia have mainly been limited to the regional verification of globally known climatic impulses. However, the role of human impact on regional vegetation patterns has been widely neglected. We postulate that today's open dwarf shrublands replace woodlands and forests. Isolated vigorous juniper trees and successful reforestation appear to confirm our hypothesis. An abrupt decline of Pinus forests before 5700 and 5400 ka cal yr BP can be demonstrated. As the first indicator pollen of human impact appeared at both sites synchronous with the forest pollen decline, we infer human impact to be a more decisive cause for this environment change superimposing the effects of a climatic deterioration. The forests were displaced by open dwarf shrublands.     

Paleoecology and high-resolution paleohydrology of a kettle peatland in upper Michigan

We investigated the developmental and hydrological history of a Sphagnum-dominated, kettle peatland in Upper Michigan using testate amoebae, plant macrofossils, and pollen. Our primary objective was to determine if the paleohydrological record of the peatland represents a record of past climate variability at subcentennial to millennial time scales. To assess the role of millennial-scale climate variability on peatland paleohydrology, we compared the timing of peatland and upland vegetation changes. To investigate the role of higher-frequency climate variability on peatland paleohydrology, we used testate amoebae to reconstruct a high-resolution, hydrologic history of the peatland for the past 5100 years, and compared this record to other regional records of paleoclimate and vegetation. Comparisons revealed coherent patterns of hydrological, vegetational, and climatic changes, suggesting that peatland paleohydrology responded to climate variability at millennial to sub-centennial time scales. Although ombrotrophic peatlands have been the focus of most high-resolution peatland paleoclimate research, paleohydrological records from Sphagnum-dominated, closed-basin peatlands record high-frequency and low-magnitude climatic changes and thus represent a significant source of unexplored paleoclimate data.     

A chronostratigraphic comparison of Late-Weichselian and Late-Devensian subdivisions, illustrated by two radiocarbon-dated profiles from western Britain

¹⁴C-dated percentage and absolute late-glacial pollen diagrams from (1) Blelham Bog, north-west England and (2) Cam Loch, north-west Scotland arc subdivided into chronozones Bølling, Older Dryas, Allerød, and Younger Dryas as proposed for the Late-Weichselian sequence by Mangerud et al., Boreas 3 (1974), with boundaries defined in conventional ¹⁴C years. It is shown how these chronozone boundaries coincide in the two British pollen diagrams with boundaries between pollen assemblage zones which are interpreted as the results of environmental (climatic) changes. Differences between the pollen zones found in western Britain and those of the classic South Scandinavian profiles, which are interpreted as the consequence of regional differentiation in the vegetation of north-west Europe circa 14,500 to 10,000 years ago, pose serious problems in the correlation of Late-Weichselian and Late-Devensian subdivisions on the basis of pollen assemblage zones, but a chronostratigraphical classification makes it possible to compare the Late-Devensian profiles from western Britain with the Late-Weichselian subdivisions of Mangerud et al., in which 'chronozone boundaries seem to be climatically conditioned within southern Scandinavia'.     

石羊河流域表土孢粉与植被的关系

通过对石羊河流域不同植被带76个表土样的孢粉分析, 研究了该流域各植被带的表土孢粉组合及主要植物花粉与其植物比例的关系. 结果显示: 石羊河流域各科属孢粉与植物之间的关系复杂, 同一科属孢粉在不同植被带中的代表性也各有差异, 所以某科属孢粉的代表性问题需要在不同植被带中分别讨论. 森林带的云杉属花粉和流域自身的大气环流状况是影响流域表土孢粉组合及各科属花孢粉与植物关系的主要因素.     

Pollen‐based biome reconstructions for Colombia at 3000, 6000, 9000, 12 000, 15 000 and 18 000 14C yr ago: Late Quaternary tropical vegetation dynamics

Colombian biomes are reconstructed at 45 sites from the modern period extending to the Last Glacial Maximum (LGM). The basis for our reconstruction is pollen data assigned to plant functional types and biomes at six 3000‐yr intervals. A reconstruction of modern biomes is used to check the treatment of the modern pollen data set against a map of potential vegetation. This allows the biomes reconstructed at past periods to be assessed relative to the modern situation. This process also provides a check on the a priori assignment of pollen taxa to plant functional types and biomes. For the majority of the sites, the pollen data accurately reflect the potential vegetation, even though much of the original vegetation has been transformed by agricultural practices. At 18 000 ¹⁴C yr BP, a generally cool and dry environment is reflected in biome, assignments of cold mixed forests, cool evergreen forests and cool grassland–shrub; the latter extending to lower altitudes than presently recorded. This signal is strongly recorded at 15 000 and 12 000 ¹⁴C yr BP, the vegetation at these times also reflecting a relatively cool and dry environment. At 9000 ¹⁴C yr BP there is a shift to biomes thought to result from slightly cooler environmental conditions. This trend is reversed by 6000 ¹⁴C yr BP; most sites, within a range of different environmental settings, recording a shift to more xeric biome types. There is an expansion of steppe and cool mixed‐forest biomes, replacing tropical dry forest and cool grassland–shrub biomes, respectively. These changes in biome assignments from the modern situation can be interpreted as a biotic response to mid‐Holocene climatic aridity. At 3000 ¹⁴C yr BP the shift is mainly to biomes characteristic of slightly more mesic environmental conditions. There are a number of sites that do not change biome assignment relative to the modern reconstruction, although the affinities that these sites have to a specific biome do change. These ‘anomalies’ are interpreted on a site‐by‐site basis. Spatially constant, but differential response of the vegetation to climatic shifts are related to changes in moisture sources and the importance of edaphic controls on the vegetation. The Late Quaternary reconstruction of large‐scale vegetation dynamics in Colombia allows an understanding of the environmental controls on these to be developed. In particular, shifts in the character of the main climatic systems that influence Colombian vegetation are described. Copyright © 2002 John Wiley & Sons, Ltd.