藏北高原高寒草地围栏禁牧和生长季降水对物种丰富度和多样性的影响(英文)

2009和2010年夏天沿藏北高原高寒草地样带调查了高寒草地生态系统(高寒草甸、高寒草原和荒漠草原)在围栏禁牧和自由放牧管理下的物种丰富度和多样性(Shannon-Wiener指数,Simpson优势度指数和Pielou均匀度指数)。研究结果显示:自2006年起藏北高原围栏禁牧在植被类型和区域尺度上没有显著改变物种丰富度和多样性。物种丰富度和多样性主要受生长季降水驱动,超过87%的变异可由生长季降水来解释。物种丰富度和多样性在自由放牧和围栏禁牧2类样地对生长季降水的响应方式一致。物种丰富度随降水呈指数型增长关系,多样性指数则呈现正线性关系。研究结果预示藏北高原地区生长季降水的变化对于物种丰富度和多样性管理至关重要,在未来高寒草地保护研究中应予以重视。     

Spatial pattern of grassland aboveground biomass and its environmental controls in the Eurasian steppe

Vegetation biomass is an important component of terrestrial ecosystem carbon stocks. Grasslands are one of the most widespread biomes worldwideplaying an important role in global carbon cycling. Thereforestudying spatial patterns of biomass and their correlations to environment in grasslands is fundamental to quantifying terrestrial carbon budgets. The Eurasian steppean important part of global grasslandsis the largest and relatively well preserved grassland in the world. In this studywe analyzed the spatial pattern of aboveground biomass(AGB)and correlations of AGB to its environment in the Eurasian steppe by meta-analysis. AGB data used in this study were derived from the harvesting method and were obtained from three data sources(literatureglobal NPP database at the Oak Ridge National Laboratory Distributed Active Archive Center(ORNL)some data provided by other researchers). Our results demonstrated that:(1) as for the Eurasian steppe overallthe spatial variation in AGB exhibited significant horizontal and vertical zonality. In detailAGB showed an inverted parabola curve with the latitude and with the elevationwhile a parabola curve with the longitude. In additionthe spatial pattern of AGB had marked horizontal zonality in the Black Sea-Kazakhstan steppe subregion and the Mongolian Plateau steppe subregionwhile horizontal and vertical zonality in the Tibetan Plateau alpine steppe subregion.(2) Of the examined environmental variablesthe spatial variation of AGB was related to mean annual precipitation(MAP)mean annual temperature(MAT)mean annual solar radiation(MAR)soil Gravel contentsoil p H and soil organic content(SOC) at the depth of 0–30 cm. NeverthelessMAP dominated spatial patterns of AGB in the Eurasian steppe and its three subregions.(3) A Gaussian function was found between AGB and MAP in the Eurasian steppe overallwhich was primarily determined by unique patterns of grasslands and environment in the Tibetan Plateau. AGB was significantly positively related to MAP in the Black Sea-Kazakhstan steppe subregion(elevation 3000 m)the Mongolian Plateau steppe subregion(elevation 3000 m) and the surface(elevation ≥ 4800 m) of the Tibetan Plateau. Neverthelessthe spatial variation in AGB exhibited a Gaussian function curve with the increasing MAP in the east and southeast margins(elevation 4800 m) of the Tibetan Plateau. This study provided more knowledge of spatial patterns of AGB and their environmental controls in grasslands than previous studies only conducted in local regions like the Inner Mongolian temperate grasslandthe Tibetan Plateau alpine grasslandetc.     

Multifunctionality and Thresholds of Alpine Grassland on the Tibetan Plateau

Ecosystems can simultaneously provide multiple functions and services. Knowledge on the combinations of such multi-dimensional functions is critical for accurately assessing the carrying capacity and implementing sustainable management. However, accurately quantify the multifunctionality of ecosystems remains challenging due to the dependence and close association among individual functions. Here, we quantified spatial patterns in the multifunctionality of alpine grassland on the Tibetan Plateau by integrating four important individual functions based on data collected from a field survey and remote sensing NDVI. After mapping the spatial pattern of multifunctionality, we extracted multifunctionality values across four types of grassland along the northern Tibet Plateau transect. Effects of climate and grazing intensity on the multifunctionality were differentiated. Our results showed that the highest values of multifunctionality occurred in the alpine meadow. Low values of multifunctionality were comparable in different types of grassland. Annual precipitation explained the large variation of multifunctionality across the different types of grassland in the transect, which showed a significantly positive effect on the multifunctionality. Grazing intensity further explained the rest of the variation in the multifunctionality (residuals), which showed a shift from neutral or positive to negative effects on multifunctionality across the different types of grassland. The consistently rapid declines of belowground biomass, SOC, and species richness resulted in the collapse of the multifunctionality as bare ground cover amounted to 75%, which corresponded to a multifunctionality value of 0.233. Our results are the first to show the spatial pattern of grassland multifunctionality. The rapid decline of the multifunctionality suggests that a collapse in the multifunctionality can occur after the vegetation cover decreases to 25%, which is also accompanied by rapid losses of species and other individual functions. Our results are expected to provide evidence and direction for the sustainable development of alpine grassland and restoration management.     

雅鲁藏布江源头区的植被及其地理分布特征

雅鲁藏布江源头区是国家级重要生态功能区,该区域自然背景资料极为缺乏。2002—06和2002—11,结合遥感影像数据,对源头区主要河谷典型地理环境位点植被进行了2次地面踏勘。结果表明：源头区主要植被类型有高寒草原、高寒草甸、高寒灌丛以及高寒垫状植物和流石坡植物。高寒草原类型主要有紫花针茅(Stipa pur purea)草原、青藏苔草(Carex moorcroftii)草原、固沙草(Orinus thoroldii)草原、藏白蒿(Artemisia younghusbandii)草原、藏沙蒿(Artemisia weiibyi)草原。高寒草甸主要类型有高山嵩草(Kobresia pygmaea)草甸、藏北嵩草(Kobresia littledalei)、三角草(Trikeraia hookeri)草甸。高寒灌丛的主要建群种有小叶金露梅(Potentilla parvifolia)、金露梅(Potentilla fruticosa)和变色锦鸡儿(Caragana versicolor)。在雪线附近有由多种高寒植物组成的垫状植物群落和流石坡稀疏植物。对群落的物种组成,分布区的土壤、水分等生态要素以及植被地理格局进行了概括性描述。     

青藏高原高寒草甸生态系统CO2通量研究进展

高寒草甸是广布于青藏高原的主要植被类型,它是青藏高原大气与地面之间生物地球化学循环的重要构成部分,在区域碳平衡中起着极为重要的作用。基于对青藏高原主要高寒草甸生态系统类型CO2通量研究方面的综述,系统分析了高寒草甸生态系统CO2通量日、季、年等不同时间尺度的变化特征以及温度、光合有效辐射、降水等主要环境因子对高寒草甸生态系统CO2通量的影响;同时,结合其他地区草地生态系统,就青藏高原三种典型高寒草甸生态系统类型源汇效应和Q10值进行了比较;最后,结合青藏高原高寒草甸生态系统CO2通量研究的现实与需要,提出了当前存在一些不确定性和有待深入研究的问题。     

青藏高原植被垂直带与气候因子的空间关系

集成了青藏高原气候区149个山地植被垂直带数据,利用国家基本气象台站自建站以来到2001年的地面观测日气象数据,计算了地面的温暖(WI)、寒冷(CI)、湿润(MI)、吉良龙夫(Kira)干湿指数、干燥度(Idm)等水热指数,运用GIS的空间分析模块,模拟了青藏高原水热条件的空间分布形势,探讨山地植被垂直带谱分布规律与制约因子的定量指标.结果表明:在高原的东北部、西北边缘,以荒漠、荒漠草原、山地森林、山地草原、灌丛、草甸为组合的半干旱、干旱结构向高原腹地以高寒草原、高山草甸、荒漠带组合的高寒干旱带谱结构的变化;东南、南部边缘,以温暖湿润为特征的以森林带为优势带谱组合结构逐渐向寒冷的高原中心变化;高原的地势效应,致使的水热形势旱现从中央向边缘变化的趋势是致使青藏高原植被垂直带谱分布的重要原因.     

1982~2013年青藏高原高寒草地覆盖变化及与气候之间的关系

利用GIMMS NDVI数据和地面气象站台观测数据,对青藏高原1982~2013年高寒草地覆盖时空变化及其对气象因素的响应进行研究,结果表明：青藏高原高寒草地生长季NDVI表现为从东南到西北逐渐减少的趋势,近32 a来,整个高原草地生长季NDVI呈上升趋势,增加速率为0.000 3/a (p<0.05);高寒草地生长季NDVI年际变化具有空间异质性,整体为增加趋势,呈增加趋势的面积约占研究区域面积的75.3%,其中显著增加的占26.0% （p<0.05）,类型主要为分布在青藏高原东北部地区的高寒草甸;比例为4.7%,草地类型主要为高寒草原,主要分布在高原西部地区;基于生态地理分区的分析显示,青藏高原草地与降水、温度的相关关系具有明显的空间差异,高寒草地生长季NDVI均值与降水呈显著正相关,对降水的滞后效应显著;高原东北部温度较高,热量条件较好,降水为高寒草地生长季NDVI变化的主导因子;东中部地区降水充沛,温度则为高寒草地生长的制约因子;南部地区降水和温度都较适宜,均与高寒草地生长季NDVI相关性显著(p< 0.05),共同作用于草地的生长;中部和西部地区,气候因子与高寒草地生长季NDVI关系均不显著。     

Cryosphere evapotranspiration in the Tibetan Plateau: A review

Land surface actual evapotranspiration is an important process that influences the Earth's energy and water cycles and determines the water and heat transfer in the soil-vegetation-atmosphere system. Meanwhile, the cryosphere's hydrological process is receiving extensive attention, and its water problem needs to be understood from multiple perspectives. As the main part of the Chinese cryosphere, the Tibetan Plateau faces significant climate and environmental change. There are active interaction and pronounced feedback between the environment and ET_a in the cryosphere. This article mainly focuses on the research progress of ET_a in the Tibetan Plateau. It first reviews the ET_a process, characteristics, and impact factors of typical underlying surfaces in the Tibetan Plateau (alpine meadows, alpine steppes, alpine wetlands, alpine forests, lakes). Then it compares the temporal and spatial variations of ET_a at different scales. In addition, considering the current greening of cryosphere vegetation due to climate change, it discusses the relationship between vegetation greening and transpiration to help clarify how vegetation activities are related to the regional water cycle and surface energy budget.     

Estimation of the Tibetan Wild Ass Population in Gaize County of Chang Tang Plateau based on the Belt Transect Method and Random Forest Model

As a typical representative of the herbivorous wild animals in Chang Tang Plateau, the number of Tibetan wild asses has increased significantly in recent years. Clarifying the distribution, population, and size of its habitats is conducive to formulating the protection plan for wild animals and managing the conflict between people and wild animals in Chang Tang Plateau. Based on the distribution probability of Tibetan wild ass habitats and environmental factors, the number of Tibetan wild asses in Gaize County of Chang Tang Plateau was calculated by using the belt transect method and random forest model, and due to the uncertainty of the calculations, the results were corrected and analyzed. The results show that the living environment of Tibetan wild asses in Gaize County of Chang Tang Plateau is at 4400-4600 m above sea level, 350-400 m away from the river, and the average temperature in the warmest season is 10-12 ℃. The vegetation types of habitats are generally temperate tufted dwarf grass, dwarf semi shrub desert grassland, alpine grass, Carex grassland, alpine cushion dwarf semi shrub desert, among others. On the basis of studying the environmental preferences of Tibetan wild asses, the random forest model was corrected by using the data of the second scientific survey sample line of Qinghai-Tibet Plateau for three years. The approximate number of Tibetan wild asses in each of the different areas of Gaize County was obtained. The number of Tibetan wild asses in Gaize Town, Xianqian Township, Gumu Township, Chabu Township, Mami Township, Wuma Township and Dongcuo Township is 855, 3458, 2358, 1453, 743, 943 and 647, respectively. By studying the environmental preferences of Tibetan wild asses and analyzing the results of the belt transect survey, the random forest model can accurately estimate the number of Tibetan wild asses in Gaize County of Chang Tang Plateau.     

基于植被盖度和高度的不同退化程度高寒草地地上生物量估算

由于气候变化和不合理的人类活动，20世纪80年代以来青藏高原高寒草地发生严重退化。地上生物量是评价草地退化的直观指标。通常采用植被盖度和高度来估算草地地上生物量，但草地退化后，植被盖度和高度与地上生物量之间的关系是否会发生变化目前还不清楚，这影响着退化草地生物量估算的精度。通过多元回归分析研究了青藏高原中部和东北部高寒草甸、高寒草原在不同退化程度下植被盖度和高度与地上生物量的关系。结果表明：（1）高寒草甸与高寒草原地上生物量整体上及不同退化阶段都没有显著差异（P>0.05）。（2）随着退化程度的加剧植被盖度和高度对地上生物量的影响也发生改变，体现在未退化阶段地上生物量主要受植被高度影响，退化后主要受植被盖度影响。（3）无论是高寒草甸还是高寒草原分退化程度的回归模型估算结果都较不分退化程度模型估算的生物量更接近实测值。我们建议在退化高寒草地研究中采用盖度和高度估算生物量时，根据退化阶段采用不同的估算模型。     

Pollen-based reconstructions of Holocene vegetation and climatic change of Tibetan Plateau

A synthesis of Holocene pollen records from the Tibetan Plateau shows the history of vegetation and climatic changes during the Holocene. Palynological evidences from 24 cores/sections have been compiled and show that the vegetation shifted from subalpine/alpine conifer forest to subalpine/alpine evergreen sclerophyllous forest in the southeastern part of the plateau; from alpine steppe to alpine desert in the central, western and northern part; and from alpine meadow to alpine steppe in the eastern and southern plateau regions during the Holocene. These records show that increases in precipitation began about 9 ka from the southeast, and a wide ranging level of increased humidity developed over the entire of the plateau around 8-7 ka, followed by aridity from 6 ka and a continuous drying over the plateau after 4-3 ka. The changes in Holocene climates of the plateau can be interpreted qualitatively as a response to orbital forcing and its secondary effects on the Indian Monsoon which expanded northwards     

青藏高原植被覆盖对水热条件年内变化的响应及其空间特征

利用1982-2000年NOAA/AVHRR卫星的NDVI数据(时间分辨率旬,空间分辨率8 km×8 km),结合同时期的气温和降水资料,基于时滞互相关方法和GIS工具,分析了青藏高原植被覆盖对水、热条件年内变化的时滞响应及其空间特征。结果如下:①除高寒荒漠、森林外,青藏高原植被NDVI与同期旬均温和旬降水相关性均呈高度正相关。其中,中等覆盖度的植被受水、热影响表现更为强烈。②青藏高原植被NDVI对气温和降水有滞后效应,且滞后水平存在空间差异,高原北部(柴达木盆地、昆仑山北冀)和高原南部植被对降水、和温度的响应比较迟缓,而高原中、东部地区植被对温度和降水的响应比较敏感。③不同植被类型对水热条件的响应程度也存在差异,由高到低依次是草甸、草原、灌丛、高寒垫状植被、荒漠,最后是森林。     

欧亚大陆草原地上生物量的空间格局及其与环境因子的关系

本文以欧亚大陆草原为研究对象,收集和整合分析了该区域采用收获法实测的地上生物量数据,探讨了地上生物量的空间分布规律,地上生物量空间格局与环境因子之间的关系。主要结论为：① 欧亚大陆草原地上生物量的空间格局具有复杂的水平和垂直地带性分布规律,具体表现为：随着纬度的升高,海拔的升高,地上生物量都呈开口向下的抛物线变化趋势;随着经度的增大,呈开口向上的抛物线变化趋势。② 欧亚大陆草原地上生物量的空间分布虽然与年平均温度、年总太阳辐射、表层土壤（0~30 cm）的砾石含量、pH和有机碳含量有一定的相关关系,但是,主要受到年总降水量空间变异的影响。③ 欧亚大陆草原地上生物量的空间格局与年总降水量空间变异之间呈现为高斯函数关系,这主要是由青藏高原独特的草原分布与环境因素空间格局特征决定的。     

青藏高原中东部植被覆盖对水热条件的响应研究

植被覆盖的变化常是自然因子和人类活动的综合作用,分析植被对水热条件的响应关系有助于认识人类活动在地表植被变化中的作用程度。本文旨在结合1982~2000年地面气象观测资料和NOAA卫星的AVHRR 植被指数（8km）,对气象站点分布相对密集的青藏高原中东部的NDVI（归一化植被指数）空间变化同水热条件的响应关系进行分析。通过水热有关指标的趋势面模拟、植被类型比较和样带分析,表明：在青藏高原中东部地区,水热条件组合较好（如常绿针叶林）或较差（如荒漠半荒漠）的区域,多年平均的NDVI旬值同水热条件的相关性不强;而范围广阔的水热条件组合中等区域（如高山草甸/草原）同水热条件相关性很高;青藏高原周边区域植被对水热条件相对不敏感,而高原主体部分植被覆盖同水热的相关性则很高（0.75以上）;此外,海拔对热量条件影响很大,进而影响植被覆盖。     

ON THE LAYER-ZONE OF ALPINE MEADOW ON THEQINGHAI-TIBETAN PLATEAU

RuleofzonalityisoneofthedassicaltheonesinGeOgraphy.Thedifferentiationofzonalityfromaz0nalityindicatesthedevel0pmentofpeople'Scoguhveabilitiestonaturallaw.Longitudinal,latitudinalandaltitudinalzonalitiesbelongt0idealconcePts,however,theywerefavorablet0theformationoftheconcePtofthIeedimensi0nalzonality.UPt0n0wtherehavebeenmanydifferentideasonzonality.ForexamPle,inanarr0wsense,zonality0ulyreferst0thelatitudinalzonality,namely,thequatityofheatorairtemPeraturegraduallyChangs,whichresultsinthezo…     

NPP vulnerability of the potential vegetation of China to climate change in the past and future

Using the Integrated Biosphere Simulator, a dynamic vegetation model, this study initially simulated the net primary productivity (NPP) dynamics of China’s potential vegetation in the past 55 years (1961–2015) and in the future 35 years (2016–2050). Then, taking the NPP of the potential vegetation in average climate conditions during 1986–2005 as the basis for evaluation, this study examined whether the potential vegetation adapts to climate change or not. Meanwhile, the degree of inadaptability was evaluated. Finally, the NPP vulnerability of the potential vegetation was evaluated by synthesizing the frequency and degrees of inadaptability to climate change. In the past 55 years, the NPP of desert ecosystems in the south of the Tianshan Mountains and grassland ecosystems in the north of China and in western Tibetan Plateau was prone to the effect of climate change. The NPP of most forest ecosystems was not prone to the influence of climate change. The low NPP vulnerability to climate change of the evergreen broad-leaved and coniferous forests was observed. Furthermore, the NPP of the desert ecosystems in the north of the Tianshan Mountains and grassland ecosystems in the central and eastern Tibetan Plateau also had low vulnerability to climate change. In the next 35 years, the NPP vulnerability to climate change would reduce the forest–steppe in the Songliao Plain, the deciduous broad-leaved forests in the warm temperate zone, and the alpine steppe in the central and western Tibetan Plateau. The NPP vulnerability would significantly increase of the temperate desert in the Junggar Basin and the alpine desert in the Kunlun Mountains. The NPP vulnerability of the subtropical evergreen broad-leaved forests would also increase. The area of the regions with increased vulnerability would account for 27.5% of China.     

Variations in the Drought Severity Index in Response to Climate Change on the Tibetan Plateau

Quantifying the relationship between the drought severity index and climate factors is crucial for predicting drought risk in situations characterized by climate change. However, variations in drought risk are not readily discernible under conditions of climate change, and this is particularly the case on the Tibetan Plateau. This study examines the correlations between the annual drought severity index (DSI) and 14 climate factors (including temperature, precipitation, humidity, wind speed, and hours of sunshine factors), on the Tibetan Plateau from 2000 to 2011. Spatial average DSI increased with precipitation and minimum relative humidity, while it decreased as the hours of sunshine increased. The correlation between DSI and climate factors varied with vegetation types. In alpine meadows, the correlation of the spatial DSI average with the percentage of sunshine and hours of sunshine (P<0.001) was higher compared to that in alpine steppes (P<0.05). Similarly, average vapor pressure and minimum relative humidity had significant positive effects on spatial DSI in alpine meadows, but had insignificant effects in alpine steppes. The magnitude of DSI change correlated negatively with temperature, precipitation, and vapor pressure, and positively with wind speed and sunshine. This demonstrates that the correlation between drought and climate change on the Tibetan Plateau is dependent on the type of ecosystem.     

基于改进土壤冻融水循环的Biome-BGC模型估算青藏高原草地NPP

Biome-BGC模型被广泛用于估算植被净初级生产力（Net Primary Productivity, NPP）,但是该模型未考虑冻土区土壤冻融水循环过程对植被生长的影响。本文基于Biome-BGC模型,改进冻土区活动层土壤冻融水循环,估算了2000—2018年青藏高原高寒草地NPP。通过比较原模型和改进后的模型,并对NPP模拟结果的时空特征进行了分析,结果表明：① 增加冻融循环提高了NPP估算精度,青藏高原草地NPP均值由114.68 gC/(m²·a)提高到128.02 gC/(m²·a)。② 原模型和改进后NPP的空间分布差异较大,时间变化趋势差异不明显。③ 青藏高原草地NPP总量为253.83 TgC/a,呈东南向西北递减的空间格局,年均增速为0.21gC/(m²·a)（P=0.023）,显著增加的占17.85%,主要分布在羌塘高寒草原地带的大部分地区和藏南山地灌木草原地带的西部。④ 该冻融水循环改进方法简单可靠,具有在其他多年冻土区推广的价值。     

Aboveground biomass in Tibetan grasslands

This study investigated spatial patterns and environmental controls of aboveground biomass (AGB) in alpine grasslands on the Tibetan Plateau by integrating AGB data collected from 135 sites during 2001–2004 and concurrent enhanced vegetation index derived from MODIS data sets. The AGB was estimated at 68.8 g m^?2, with a larger value (90.8 g m^?2) in alpine meadow than in alpine steppe (50.1 g m^?2). It increased with growing season precipitation (GSP), but did not show a significant overall trend with growing season temperature (GST) although it was negatively correlated with GST at dry environments (<200 mm of GSP). Soil texture also influenced AGB, but the effect was coupled with precipitation; increased silt content caused a decrease of AGB at small GSP, and generated a meaningful increase under humid conditions. The correlation between AGB and sand content indicated an opposite trend with that between AGB and silt content. An analysis of general linear model depicted that precipitation, temperature, and soil texture together explained 54.2% of total variance in AGB. Our results suggest that moisture availability is a critical control of plant production, but temperature and soil texture also affect vegetation growth in high-altitude regions.