内蒙古宁城县道虎沟叶肢介化石群的时代

内蒙古赤峰宁城县道虎沟含丰富的叶肢介壳瓣及许多珍罕的软体化石 ,计有 1属 4种 :Euestheria ziliu-jingensis Chen,Euestheria haifanggouensis Chen,Euestheria jingyuanensis Chen,Euestheria luanpingensis Shen etNiu。这些分子都是我国中侏罗世早期自流井真叶肢介 (Euestheria ziliujingensis)动物群的重要成员 ,以个体较小、壳瓣圆或椭圆形、生长带具小网状装饰、种类单调、密集群生为特点 ,主要出现于中侏罗世早中期 (Bajocian- MiddleBothonia)。特别是 Euestheria luanpingensis Shen et Niu及其软体化石在道虎沟的发现 ,证明其层位应与冀北滦平九龙山组相当。道虎沟叶肢介群与中侏罗世晚期的假线叶肢介 (Pseudograpta)动物群的面貌相差甚远 ,与比其层位更高的大北沟组的尼斯脱叶肢介 (N estoria)动物群及义县组的东方叶肢介 (Eosestheria)动物群迥然不同 .     

New Fossils of Eoptychopteridae (Diptera) from the Middle Jurassic of Northeastern China

Three new species of the extinct genus of Eoptychopterina from the Eoptychopteridae     

A New Axymyiid Genus with Two New Species from the Middle Jurassic of China (Diptera: Nematocera: Axymyiidae)

The Axymyiidae is one of the small families of the suborder Nematocera within the Diptera.Up to date,three genera and seven species of extant axymyiids have been described from the Holarctic Region,including Canada,China,Hungary,Japan,Russia,and USA,with three genera and three species of fossil taxa described from the Middle Jurassic of Daohugou,northeastern China.A new genus Raraxymyia gen.nov.and two new species,R.parallela and R.proxima gen.et sp.nov.from the Middle Jurassic Jiulongshan Formation of Daohugou Village,Inner Mongolia,China,are herein described based on their well-preserved wings and body characters.A revised key to the genera of axymyiids is provided.     

A New Fossil Scorpionfly (Insecta: Mecoptera: Holcorpidae) with Extremely Elongate Male Genitalia from Northeastern China

Since Holcorpa maculosa was first described in 1878, it has attracted lots of attention due to its extremely elongate abdominal segments from the sixth to the eighth. Fossil records of family Holcorpidae are very rare, comprising two species in one genus represented by three fossil specimens, resulting in limited knowledge on the structure and function of holcorpid’s unique male organ, as well as its evolutionary relationship with other scorpionflies. Herein we describe a new genus with a new species, Conicholcorpa stigmosa gen. et sp. nov., of the Holcorpidae from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Inner Mongolia, China. This is the first holcorpid documented from the Daohugou locality, albeit two previously reported species with extremely elongate male organs but deemed as Family Incertae sedis. C. stigmosa gen. et sp. nov. is the earliest holcorpid hitherto, extending its existence from the Early Eocene to the Middle Jurassic by 114 million years and providing one more early case of presenting exaggerated male body parts for sexual display and/or selection. Our analysis supports the sister relationship between Holcorpidae and Orthophlebiidae. Our results not only enhance our knowledge of the morphology of Holcorpidae but also elucidate the relationship of known species of this family.     

New Species of Cimbrophlebiidae (Insecta: Mecoptera) from the Middle Jurassic of Northeastern China

Four new species of three genera in the family Cimbrophlebiidae, Telobittacus decorus sp. nov., Bellicimbrophlebia heteroneura sp. nov., Cimbrophlebia amoena sp. nov., and Cimbrophlebia gracilenta sp. nov., are described and illustrated based on eleven well-preserved fossil specimens. These specimens were collected from the latest Middle Jurassic Jiulongshan Formation of Daohugou, Inner Mongolia, China. This is the first record of the genus Cimbrophlebia in the Jurassic extending its age from the Lower Cretaceous to the Middle Jurassic. Due to same venational characters, we propose that an undescribed fossil specimen of Cimbrophlebiidae from the Jurassic of Germany is attributed to C. gracilenta sp. nov. Our new findings demonstrate an evolutionary trend of venational change from 185 to 50 Ma that the level of the bifurcation of Rs is gradually moving toward the base of wing in reference to the bifurcation of M.     

A new species of Eomeropidae (Insecta: Mecoptera) from the Middle Jurassic of China

The Eomeropidae is a species-poor family with only 11 known fossil species from the Early Jurassic to Palaeogene. A new species of fossil eomeropid, Tsuchingothauma gongi sp. nov., is described based on a well-preserved wing from the Middle Jurassic Daohugou deposits of Inner Mongolia, China. Our new species is distinguished from the type species T. shihi mainly in having very numerous crossveins and cells; more longitudinal veins: 13 and 11 terminal branches in radial sector and medial field, CuA with 3 terminal branches and A with 7 terminal branches.     

燕辽生物群叶肢介动物群组成和演化*

中国北方著名中、晚侏罗世的燕辽生物群含有丰富的叶肢介动物群。早期燕辽生物群也称为道虎沟生物群,主要产出层位为海房沟组,含有三饰叶肢介动物群,主要分子包括: 海房沟三饰叶肢介(Triglypta haifanggouensis)、滦平三饰叶肢介(T.luanpingensis)和平泉三饰叶肢介(T. pingquanensis)。晚期燕辽生物群又可称为玲珑塔生物群,主要产出层位为髫髻山组,含有丰富的叶肢介化石,主要分子包括: 建昌三饰叶肢介(Triglypta jiancangensis)、玲珑塔辽西叶肢介(Liaoxiestheria linglongtaensis)、大西山玲珑塔叶肢介(Linglongtaestheria daxishanensis)和青龙玲珑塔叶肢介(Linglongtaestheria qinglongensis)。文中总结了北方雕饰叶肢介(Aquilonoglypta)、柴达木叶肢介(Qaidamestheria)和针孔叶肢介(Punctatestheria)的最新研究成果,支持将柴达木叶肢介、辽西叶肢介和玲珑塔叶肢介划归北方雕饰叶肢介科。同时,三饰叶肢介是由针孔叶肢介演化而来,与柴达木叶肢介并无直接演化关系。     

New Isophlebioid Dragonflies from the Middle Jurassic of Inner Mongolia, China (Insecta: Odonata: Isophlebioptera: Campterophlebiidae)

Three new species of fossil dragonflies assigned to Sinokaratawia Nel, Huang and Lin in family Campterophlebiidae, i.e.S, daohugouica sp. nov., S. magica sp. nov. and S. gloriosa sp. nov., and new materials of male S. prokopi Nel, Huang and Lin, 2007 are described from the Middle Jurassic of Daohugou, Inner Mongolia, China. An emended diagnosis of genus Sinokaratawia was proposed.     

New Fossils of Eoptychopteridae (Diptera)from the Middle Jurassic of Northeastern China

Three new species of the extinct genus of Eoptychopterina from the Eoptychopteridae family, Eoptychopterina antica sp. nov., Eoptychopterina adnexa sp. nov., and Eoptychopterina mediata sp. nov., are described and illustrated. These three new species are established based on fossil specimens with bodies and complete wings. All were collected from the Middle Jurassic Jiulongshan Formation of Daohugou in eastern Inner Mongolia, China. Based on the new materials, the name of two species in Eoptychopterina from China--Eoptychopterina elenae Ren and Krzeminski and Eoptychopterina gigantea Zhang--is sysnonymum junius.     

Current Knowledge of Mesozoic Coleoptera from Daohugou and Liaoning (Northeast China)

<正>The present paper is devoted to an overview on fossil Coleoptera studied from Inner Mongolia, Daohugou(Middle Jurassic,Jiulongshan Formation)and Liaoning(Upper Jurassic-Lower Cretaceous,Yixian Formation)deposited in Chinese collections.As a result,species of the tribe Sperchopsini and Hydrophilini from Hydrophilidae,families and subfamilies Silphidae,Syndesinae from Lucanidae,Pleocomidae,Trogidae, Trogissitidae,Pyrochroidae,Diaperinae from Tenebrionidae,and Cerambycidae were first registered in the Mesozoic and some families were defined as new.It was shown that many superfamilies represented in the Recent Fauna were formed within the Middle Jurassic and Lower Cretaceous.The materials examined confirm the hypothesis that Cucujiformian beetles are a younger group than other infraordera of Polyphaga(Staphyliniformia and Elateriformia)and,therefore,they appeared in the fossil record only in the late Mesozoic.It was shown and confirmed that most superfamilies appeared in the fossil records before Cucujoidea.The synonymy of Notocupes Ponomarenko,1964;Sinocupes Lin,1976,syn.nov.;Amblomma Tan,Ren et Liu 2005,syn.nov.;Euryomma Tan, Ren et Shih,2006,syn.nov.,non Stein,1899 and Ovatocupes Tan et Ren,2006,syn.nov.;synonymy of Tetraphalerus Waterhouse,1901 and Odontomma Tan,Ren et Ge 2006,syn.nov.;and synonymy of Priacmopsis Ponomarenko,1966 and Latocupes Tan et Ren,2006,syn.nov.are proposed.Sinorhombocoleus papposus Tan et Ren,2009 is transferred from the family Rhombocoleidae to Schizophoridae.Cervicatinius complanus Tan,Ren et Shih,2007 and Forticatinius elegans Tan,Ren et Shih,2007 are transferred from the family Catiniidae(suborder Archostemata)to superfamily Cleroidea(suborder Polyphaga:first among the family Peltidae and second as a closely related group to the latter family).The family Parandrexidae is transferred from the superfamily Cucujoidea to Cleroidea.The ecological circumstances of the past ecosystems and hypotheses of historical development of the order Coleoptera are discussed.The age of faunas examined is considered.The list of the taxa described from Daohugou and Liaoning is compiled.     

Early Evolution of Apocrita(Insecta,Hymenoptera)as Indicated by New Findings in the Middle Jurassic of Daohugou,Northeast China

<正>The classification and phytogeny of the basal Vespina(=Orussoidea+Apocrita)are reconsidered based primarily on rich and well preserved material from the Middle Jurassic of Daohugou in Inner Mongolia,China. Comparatively smooth morphological transitions are traced from a Xiphydriidae-like ancestor toward Orussoidea via the Jurassic family Karatavitidae,and through Karatavitidae and the Jurassic and Early Cretaceous family Ephialtitidae independently to Stephanidae,to Evanioidea,and,via the extinct Jurassic Kuafuidae fam.nov.to the remaining Apocrita.New hypothesis is proposed concerning development of the characteristic wasp-waist of Apocrita,which is supposed to appear independently and in different ways in Evanioidea and in the rest of Apocrita.As a result,six infraorders are proposed for the suborder Vespina with the following taxonomic structure:infraorder Orussomorpha including the only superfamily Orussoidea(Karatavitidae+Paroryssidae + Orussidae),infraorder Stephanomorpha with the only superfamily Stephanoidea(Ephialtitidae+Stephanidae), infraorder Evaniomorpha with the only superfamily Evanioidea of traditional composition,infraorder Ceraphronomorpha with the superfamilies Ceraphronoidea s.str.and monotypical Megalyroidea and Trigonaloidea,and the infraorders Proctotrupomorpha,Ichneumonomorpha,and Vespomorpha of traditional composition.The family Kuafuidae is unplaced to infraorder because it is putatively paraphyletic with respect to Ceraphronomorpha,Proctotrupomorpha,Ichneumonomorpha and Vespomorpha.Described as new are Karatavites junfengi sp.nov,Praeratavites wuhuaensis sp.nov.,P.perspicuus sp.nov.,Postxiphydria daohugouensis gen.et sp.nov.,P.ningchengensis gen.et sp.nov.,Postxiphydroides strenuus gen.et sp.nov.,Praeratavitoides amabilis gen.et sp.nov.,Proapocrtius densipediculus sp.nov.,P.sculptus sp.nov.,P.longantennatus sp.nov., P.formosus sp.nov.,P.atropus sp.nov.,P.elegans sp.nov.,Stephanogaster pristinus sp.nov.,Asiephialtites lini sp. nov.,Praeproapocritus vulgatus gen.et sp.nov.,Sinaulacogastrinus solidus gen.et sp.nov.,Sinevania speciosa gen. et sp.nov.,Eonevania robusta gen.et sp.nov.,Kuafua polyneura gen.et sp.nov.(all from the Middle Jurassic of Daohugou),as well as Kuafuidae fam.nov.including Kuafua gen.nov.as well as Arthrogaster Rasnitsyn,1975,and Leptogastrella Rasnitsyn,1975 from the Upper Jurassic of Karatau in Kazakhstan.     

New Fossil Palaeontinids from the Middle Jurassic of Daohugou,Inner Mongolia,China (Insecta,Hemiptera)

Two new genera and species of fossil Palaeontinidae are described from Daohugou Village, Inner Mongolia, China: Cladocossus undulatus gen. et sp. nov. and Cricocossus paradoxus gen. et sp. nov. Both new genera are described based on well-preserved forewings. This discovery confirms the high diversity of palaeontinids during the Middle Jurassic. Both specimens have interesting modal structures which are new to Palaeontinidae: M five-branched and M3 with two branches. Based on this character, wing structural characteristic is discussed.     

A New Cockroach Genus of the Family Fuziidae from Northeastern China (Insecta: Blattida)

A new genus and two new species of fossil Fuziidae are described and illustrated: Parvifuzia gen. nov., P. marsa sp. nov. and P. brava sp. nov.. Both species were collected from the Middle Jurassic Jiulongshan, Formation of Daohugou, Inner Mongolia of China. In conjunction with Fuzia Vr?ansky, Liang et Ren, 2009, the present findings allow us to demonstrate that members of the family were morphologically diverse, with body length ranging from medium (17 mm) in Fuzia to small (9.7 mm) in the present genus. Wing venation of F. dadao is complicated, but simple in the present species, and apex of male’s cerci are sharp and forceps-like in F. dadao, but strongly curved inward and round in the new genus.     

New Species of Cretolimonia and Mesotipula (Diptera: Limoniidae) from the Middle Jurassic of Northeastern China

Two new species, Cretolimonia excelsa sp. nov. and Mesotipula gloriosa sp. nov., in Limoniidae of Diptera are described and illustrated, based on three fossil specimens with bodies and complete wings. These specimens were collected from the latest Middle Jurassic Jiulongshan Formation at Daohugou, Inner Mongolia, China. These are the first fossil records of these two genera from China.     

A New Elaterid from the Middle Jurassic Daohugou Biota (Coleoptera: Elateridae: Protagrypninae)

A new genus and species of fossil elaterid namely Clavelater ningchengensis gen. et sp. nov. assigned to the subfamily Protagrypninae based on one specimen from the Middle Jurassic Jiulongshan Formation near Daohugou Village, Ningcheng County, Inner Mongolia, northeastern China is described and illustrated. This specimen displays a pair of large antennae that easily differs from those of most Mesozoic elaterids. There are around 36 species within 20 genera referred to elateroids described from the Early Jurassic to Miocene of China, with another three species within three genera reviewed, but the taxonomic assignment of at least seven species within three genera is doubted. The systematic position of fossil elateriform beetles reported from China is reviewed and the evolution of Mesozoic elateroids briefly discussed.     

Radiometric Dating of Ignimbrite from Inner Mongolia Provides no Indication of a Post-Middle Jurassic Age for the Daohugou Beds

1 Introduction and Background Along with stunning paleontological discoveries from the world-renowned fossil beds in western Liaoning Province of China, recent findings from the lacustrine deposits exposed at the Daohugou village (Fig. 1) near Ningcheng, Inner Mongolia, include superbly preserved pterosaur, salamander, insect, and plant fossils (Ji and Yuan, 2002; Wang et al., 2002; Gao and Shubin, 2003; Zheng et al., 2003; Rasnitsyn and Zhang, 2004). The Daohugou fossil beds consist of…     

A new caraboid larva from the Middle Jurassic of China (Insecta: Coleoptera)

A new species of caraboid larva (Coleoptera, Adephaga), Carabilarva gongi sp. nov., is described based on a well-preserved specimen from the Middle Jurassic Daohugou deposits (in the upper part of Jiulongshan Formation) of Inner Mongolia, China. It is the fourth caraboid larva from the Mesozoic, and differs from the other three Mesozoic species mainly in having a longer body with shorter mandibles. Scanning electron microscopy (SEM) and energy dispersive X-ray spectroscopy (EDS) show that this Daohugou fossil is preserved as carbonaceous compressions that lost the micro-structure, such as macrochaetae.     

New Genus and Species of Rhyacophilidae (Insecta: Trichoptera) from the Middle Jurassic of China

One new genus with a new fossil species,Declinimodus setulosus gen.et sp.nov.which was found in the Middle Jurassic Jiulongshan Formation of Daohugou,Inner Mongolia,China,are described and illustrated.The new species is assigned to the Rhyacophilidae according to buttonshaped of the 2nd segment of maxillary palp,and the forked R1 （in the forewing,located near apex）.These new discovery are the earliest fossil records of Rhyacophilidae,which providing important evident for the study of the origin and early evolution of Trichoptera.     

A New Family of Moths from the Middle Jurassic (Insecta: Lepidoptera)

<正>Three lepidopteran species,from the Middle Jurassic Daohugou beds(inner Mongolia,China),are described in a new family,Mesokristenseniidae,and new genus,Mesokristensenia,which could represent the sister group of the Micropterigidae.Mesokristensenia differs from all extant Lepidoptera,but one genus(Agathiphaga, Agathiphagidae),in retaining four median veins in the forewing,a plesiomorphy also present in many Trichoptera. Evidence for placing Mesokristensenia in the Lepidoptera includes four traits,notably a previously unrecorded autapomorphy of this insect order:beyond stem M1+2,vein M1 is bent and connected to cross-vein r-m(in both wing pairs).Among 24 characters taken into account to assess the systematic position of Mesokristensenia,12 are considered informative for a cladistic analysis involving this fossil taxon and the four suborders recognized in present-day Lepidoptera(Zeugloptera,Aglossata,Heterobathmiina,and Glossata).