An 11 000-year record of vegetation and environment from Lago di Martignano,Latium, Italy

A pollen diagram from Lago di Martignano, a maar lake in central Italy, provides an 11000-year record of vegetation and environment change. The earliest pollen spectra are dominated by Artemisia and Gramineae, representing late glacial steppe vegetation typical of the Mediterranean region. Broad-leaved forests were established by ca. 11 000 yr BP. Although Quercus initially dominated their canopy, a wide range of other mesophyllous trees were also present. Pollen values for sclerophyllous tree and shrub taxa characteristic of Mediterranean woodlands and scrub are initially low (<10%). After ca. 7000 yr BP, however, they begin to increase and rise to a peak of >40% of total land pollen at ca. 6700 yr BP, with Olea europaea the single most abundant taxon. Human influence upon the vegetation only becomes significant somewhat after this peak, with progressive clearance of woodland and expansion of herbaceous communities. Castanea sativa and luglans regia pollen is recorded consistently from the beginning of the rise in pollen values for taxa characteristic of Mediterranean scrub communities. Pollen values for arable crops increase progressively after ca. 5500 yr BP, following the peak pollen values for taxa characteristic of Mediterranean scrub vegetation. Late glacial and Holocene climate changes have been complex in this region, with the present character of the climate developing only during the last millennium. Rates of change of pollen spectra peak during this period.     

Dynamic early Holocene vegetation development on the Faroe Islands inferred from high-resolution plant macrofossil and pollen data

Vegetation dynamics during the earliest part of the Holocene (11,250-10,250 cal yr BP) have been reconstructed from a lacustrine sequence on Sandoy, the Faroe Islands, using detailed plant macrofossil and pollen evidence. The plant macrofossils suggest the initial vegetation was sparse herb and shrub tundra, with Salix herbacea and open-ground species, followed by the development of a denser and more species-rich arctic heathland after 11,150 cal yr BP. Despite high pollen values for Betula nana, macrofossils are rare. The bulk of the macrofossils recorded are S. herbacea and Empetrum leaves with numerous herb taxa and an abundance of Racomitrium moss. Conditions start to change around 10,800 cal yr BP, with increased catchment erosion and sediment delivery to the lake from ca. 10,600 cal yr BP, and a transition to alternating Cyperaceae and Poaceae communities between ca. 10,450 and 10,250 cal yr BP. This vegetation change, which has been recorded throughout the Faroes, has previously been interpreted as a retrogressive shift from woody shrubs to a herbaceous community. The detailed plant macrofossil data show the shift is the replacement of an Empetrum arctic heathland by grassland and moist sedge communities. These taxa dominate the modern landscape.     

Late quaternary vegetation of southern Isla Grande de Chiloé, Chile

Late-glacial-Holocene forest history of southern Isla Chiloé (latitude 43°10′ S) was reconstructed on the basis of pollen analysis in three profiles (Laguna Soledad, Laguna Chaiguata, Puerto Carmen). Prior to 12,500 yr B.P. pollen records are dominated by plant taxa characteristic of open habitats (Zone I). From 12,500 yr B.P. to the present, tree species predominate in the pollen records (Zones II–V). Between 12,500 and 9500 yr B.P. ombrophyllous taxa (Nothofagus, Podocarpus nubigena. Myrtaceae, Fitzroya/Pilgerodendron, and Drimys) are frequent in all pollen diagrams, suggesting a wetter and colder climate than the present. Between 9000 and 5500 yr B.P. Valdivian forest elements, such as Nothofagus dombeyi type, Weinmannia, and Eucryphia/Caldcluvia, dominate, indicating a period of drier and warmer climate. From 5500 yr B.P. onward, the expansion of mixed North Patagonian-Subantarctic forest elements and the increased frequence of Tepualia suggest increased rainfall and temperatures oscillating around the modern values.The change from open to forest vegetation (ca. 12,500 yr B.P.) probably represents the most pronounced climatic change in the record and can be interpreted as the glacial-postglacial transition in the study area.     

A record of late-Holocene environmental change from southern New England, USA

Analyses of a sediment core from Little Pond, located in the town of Bolton, Massachusetts, provide new insights into the history of environmental and ecological changes in southern New England during the late Holocene. Declines in organic content and peaks in the abundance of Isoetes spores indicate reduced water depth at 2900–2600, 2200–1800, and 1200–800 calibrated years before present (cal yr BP), generally consistent with the timing of dry conditions in records from elsewhere in the northeastern United States. The Little Pond pollen record features little change over the last 3000 yr, indicating that the surrounding vegetation was relatively insensitive to these periods of drought. The 1200–800 cal yr BP dry interval, however, coincides with increased abundance of Castanea pollen, suggesting that the expansion of Castanea in southern New England may have been influenced by late-Holocene climatic variability.     

An 8000-year record of vegetation, climate, and human disturbance from the Sierra de Apaneca, El Salvador

An ∼8000-cal-yr stratigraphic record of vegetation change from the Sierra de Apaneca, El Salvador, documents a mid-Holocene warm phase, followed by late Holocene cooling. Pollen evidence reveals that during the mid-Holocene (∼8000-5500 cal yr B.P.) lowland tropical plant taxa were growing at elevations ∼200-250 m higher than at present, suggesting conditions about 1.0°C warmer than those prevailing today. Cloud forest genera (Liquidambar, Juglans, Alnus, Ulmus) were also more abundant in the mid-Holocene, indicating greater cloud cover during the dry season. A gradual cooling and drying trend began by ∼5500 cal yr B.P., culminating in the modern forest composition by ∼3500 cal yr B.P. A rise in pollen from weedy plant taxa associated with agriculture occurred ∼5000 cal yr B.P., and pollen from Zea first appeared in the record at ∼4440 cal yr B.P. Human impacts on local vegetation remained high throughout the late Holocene, but decreased abruptly following the Tierra Blanca Joven (TBJ) eruption of Volcán Ilopango at ∼1520 cal yr B.P. The past 1500 years are marked by higher lake levels and periodic depositions of exogenous inorganic sediments, perhaps indicating increased climatic variability.     

Climatic instability in west equatorial Africa during the Mid- and Late Holocene

Millennial-scale climatic variations have punctuated the Holocene characterised by abrupt changes from warm to cool or wetter to drier conditions. Amongst these climatic events, there is increased evidence for an abrupt multicentennial shift of climatic conditions around 3.8/3.7 kyr BP (4.1 cal. kyr BP) in mid- to low-latitude regions which had a profound impact on landscape and population migration. In the Mediterranean region, subtropical, tropical and equatorial Africa, a number of continental proxies (lake-levels, pollen sequences, stable isotopes) record this abrupt change towards drier conditions. However, regionalism in climatic conditions is reflected in the vegetation records, possibly in relation to orographic conditions and the influence of sea-surface conditions. Hitherto there have been very few marine sequences that record this particular climatic shift at high-resolution. We present here new data from the Congo deep-sea fan containing integrated marine and terrestrial proxies. Around 5–4 cal. kyr BP, shifts in surface conditions off the Congo River mouth are observed, with possible establishment of seasonal coastal upwelling, and lower sea-surface temperatures. In parallel, pollen data indicate fluctuations of herbaceous, afromontane taxa and charred grass cuticles, suggesting more open vegetation in the lowland regions and an increase in cloud forest and/or afromontane vegetation at higher altitudes within the Congolese region.     

Fire and Vegetation History from the Coastal Rain Forest of the Western Oregon Coast Range

High-resolution charcoal and pollen analyses were used to reconstruct a 4600-yr-long history of fire and vegetation near Taylor Lake in the wettest forests of coastal Oregon. Today, fires in these forests are rare because the season of ignition does not coincide with months of dry fuels. From ca. 4600 to 2700 cal yr B.P. fire episodes occurred at intervals of 140±30 yr while forest vegetation was dominated by disturbance-adapted taxa such as Alnus rubra. From ca. 2700 cal yr B.P. to the present, fire episodes have become less common, occurring at intervals of 240±30 yr, and fire-sensitive forest taxa, such as Tsuga heterophylla and Picea sitchensis, have become more prominent. Fire occurrence during the mid-Holocene was similar to that of the more xeric forests in the eastern Coast Range and suggests that summer drought was widespread. After ca. 2700 cal yr B.P., a decrease in fire episode frequency suggests that cooler conditions and possibly increased summer fog allowed the establishment of present-day Picea sitchensis forests within the watershed. These results provide evidence that fire has been an important disturbance agent in the Coast Range of Oregon, and variations in fire frequency and climate have led to the establishment of present-day forests.     

Separating climatic and possible human impacts in the early Holocene: biotic response around the time of the 8200 cal. yr BP event

The early Holocene is characterised by rapid climate change events, which in the North Atlantic region are often associated with changes in thermohaline circulation. Superimposed on this in northwest Europe is localised evidence for human impact on the landscape, although separating climatic and anthropogenic mechanisms for environmental change is often difficult. Biotic and sedimentological evidence from a lacustrine sequence from the Inner Hebrides, Scotland, shows a considerable reduction in inferred local woodland centred upon 8250 cal. yr BP. These data correlate precisely with a distinctive rise in the charcoal:pollen ratio and hence suggest a possible Mesolithic human impact upon the vegetation around this time. A quantitative temperature reconstruction from chironomid analyses from the same sequence, supported by sedimentological data, indicates that the fall in arboreal pollen taxa occurred as climate warmed significantly during the early Holocene. This warming was followed by a significant cold event, with mean July temperatures reduced by 2°C, that lasted for at least 320 years ca. 7790–7470 cal. yr BP. Woodland recovered during this phase suggesting that the vegetation during the 8250 cal. yr BP interval was likely to have been responding to human activity, and not climate, and hence it is possible at specific sites to separate the influence of these key drivers of environmental change. Copyright © 2006 John Wiley & Sons, Ltd.     

Pollen Evidence of Changing Holocene Monsoon Climate in Sichuan Province, China

Pollen evidence from Lake Shayema, Mianning County, was obtained to examine postglacial vegetation and climatic change in southwestern Sichuan, China. The sclerophyllous character of the region's warm temperate vegetation today is a reflection of extreme drought in spring before the onset of the Asian monsoons. The pollen record displays several changes in the vegetation over the last 11,000 yr. From 11,000 to 9100 yr B.P., cold-tolerant species, such as Abies , Betula, and deciduous oaks, dominated the vegetation. Between 9100 and 7800 yr B.P., the abundance of deciduous oaks decreased and evergreen oaks increased, as did Tsuga and mesic deciduous species. This change suggests a warming climate with increased precipitation. From 7800 to 4000 yr B.P., sclerophyllous species increased at the expense of mesic deciduous species, an indication that precipitation was becoming more seasonal. Except for increased disturbance starting ca. 1000 yr B.P., the predominance of sclerophyllous vegetation continued until today. The pollen results are compatible with proposed global circulation hypotheses of a strengthened monsoon system during the early to mid Holocene.     

Expansion of Magellanic Moorland during the late Pleistocene: Palynological evidence from northern Isla de Chiloé, Chile

The late Quaternary vegetation of northern Isla de Chiloé is inferred from palynological analysis of a section in the Río Negro drainage (42°03′S, 73°50′W). At ca. 30,500 yr B.P., maxima of Astelia and Donatia occurred, suggesting wetland development. From that time until ca. 27,000 yr B.P., steppe indicators such as Compositae/Gramineae dominated, suggesting drier conditions. After 27,000 yr B.P., the moorland shrub Dacrydium gradually increased, reaching a maximum by 18,000 yr B.P. At this time Astelia increased again, suggesting development of cushion bog during cold and wet conditions. The glacial-postglacial transition is characterized by a marked change from peaty sediments to clays, a decrease in the cushion bog flora, and the prevalence of Gramineae/ Compositae and swamp taxa. This vegetation prevailed until ca.7000 yr B.P. when forest taxa became dominant. The floristic pattern inferred from the pollen spectra of the Rio Negro section suggests that the late Pleistocene vegetation of Chiloé resembled modern Magellanic Moorland vegetation (52°–56°lat S). Based on climatic conditions presently associated with Magellanic Moorland, its occurrence in Chiloé at low elevations during the late Pleistocene implies a decrease in average temperature of at least 4°C and an increase in annual precipitation of at least 1500 mm.     

Late Quaternary vegetational and climatic change in the Popayán region,southern Colombian Andes

Late Pleistocene and Holocene vegetational and climatic change have been studied palynologically at a site at 1750 m elevation in the subandean vegetation belt near Popayán, in the southern Colombian Andes. Time control on the pollen record is based on six AMS ¹⁴C ages, ranging from possibly Middle Pleniglacial time (around 50000 yr BP) to 1092 ± 44 yr BP. Because of the presence of two hiatuses only the Middle Pleniglacial and Late Holocene periods (the last 2300 yr BP) are represented. Pollen data indicate the presence of closed subandean forest during glacial time. Changes in the contribution of pollen originating from the uppermost and lowermost subandean forest belts, changes in the contribution of a number of other subandean forest taxa, and changes in species composition between the three pollen zones, suggest that the climate during the Middle Pleniglacial was markedly colder, and perhaps also wetter, than during the Late Holocene. Pollen assemblages from the Late Holocene indicate that the landscape has been affected by deforestation and agriculture since at least 2300 yr BP, but that human impact decreased in the last 780 yr BP. © 1998 John Wiley & Sons, Ltd.     

Two Tephra Layers Bracketing Late Holocene Paleoecological Changes in Northern Germany

Paleoecological records from two Holocene peat bogs in northern Germany are linked by two microscopic volcanic ash layers, correlated by petrology and geochemistry to explosive volcanism on Iceland. The younger “Microlite tephra” cannot be correlated to any known eruption, while the older tephra layer is identified as a deposit of the Hekla 3 eruption. The tephra layers are dated by an age–depth regression of accelerator mass spectrometry ¹⁴C ages that have been calibrated and combined in probability distributions. This procedure gives an age of 730–664 cal yr B.C. for the “Microlite tephra” event and 1087–1006 cal yr B.C. for the Hekla 3 event. Accordingly, the tephra layers were deposited during the late Bronze Age. At this time, human settlement slowly increased pressure on the environment, as indicated by changes in woodland pollen composition at the two bogs. The tephra-marker horizons further show that the palynologically defined transition from the Subboreal to the Subatlantic Period is synchronous in the investigated area. However, the macroscopic visible marker in peat, the change from fibrous to sapric peat, the “Schwarztorf-Weißtorf-Kontakt,” is asynchronous. Bog vegetation did not immediately react in unison to a climatic change at this pollen zone boundary; instead, the timing of vegetation change depended on the location within the bog.     

Rapid vegetation change during the early Holocene in the Faroe Islands detected in terrestrial and aquatic ecosystems

High‐resolution pollen, plant macrofossil and sedimentary analyses from early Holocene lacustrine sediments on the Faroe Islands have detected a significant vegetation perturbation suggesting a rapid change in climate between ca. 10 380 cal. yr BP and the Saksunarvatn ash (10 240±60 cal. yr BP). This episode may be synchronous with the decline in δ¹⁸O values in the Greenland ice‐cores. It also correlates with a short, cold event detected in marine cores from the North Atlantic that has been ascribed to a weakening of thermohaline circulation associated with the sudden drainage of Lake Agassiz into the northwest Atlantic, or, alternatively, a period with distinctly decreased solar forcing. The vegetation sequence begins at ca. 10 500 cal. yr BP with a succession from tundra to shrub‐tundra and increasing lake productivity. Rapid population increases of aquatic plants suggest high summer temperatures between 10 450 and 10 380 cal. yr BP. High pollen percentages, concentrations and influx of Betula, Juniperus and Salix together with macrofossil leaves indicate shrub growth around the site during the initial phases of vegetation colonisation. Unstable conditions followed ca. 10 380 cal. yr BP that changed both the upland vegetation and the aquatic plant communities. A decrease in percentage values of shrub pollen is recorded, with replacement of both aquatics and herbaceous plants by pioneer plant communities. An increase in total pollen accumulation rates not seen in the concentration data suggests increased sediment delivery. The catchment changes are consistent with less seasonal, moister conditions. Subsequent climatic amelioration reinitiated a warmth‐driven succession and catchment stabilisation, but retained high precipitation levels influencing the composition of the post‐event communities. Copyright © 2003 John Wiley & Sons, Ltd.     

Holocene insect fossils from two sites at Ennadai Lake, Keewatin, Northwest Territories, Canada

Ennadai Lake, in the forest-tundra ecotonal region of Keewatin, Northwest Territories, Canada, has been the subject of several paleoecological investigations (palynology, plant macrofossils, fossil soils). This study concerns Holocene insect fossils at Ennadai, a new approach in a region shown to be sensitive to climatic change. The Ennadai I site yielded 53 taxa, representing 13 families of Coleoptera and 7 families of other insects and arachnids, including abundant ants. These fossils range in age from about 6300 to 630 yr B.P. The Ennadai II site produced fossils of 58 taxa, including 13 beetle families and 15 families of other arthropods, ranging in age from 4700 to 870 yr B.P. The insect evidence suggests the presence of trees in the Ennadai region from 6000 to 2200 yr B.P. A conifer pollen decline from 4800 to 4500 yr B.P. at Ennadai has previously been interpreted as an opening up or retreat of forest in response to climatic cooling, but the insect fossils reveal the continued presence of trees during this interval. Both insect assemblages suggest trends of forest retreat and tundra expansion between about 2200 and 1500 yr B.P., presumably due to climatic cooling, with a return of woodland by about 1000 yr B.P.     

Analysis of coprolites from the extinct mountain goat Myotragus balearicus

Humans colonized the Balearic Islands 5–4 ka ago. They arrived in a uniquely adapted ecosystem with the Balearic mountain goat Myotragus balearicus (Bovidae, Antilopinae, Caprini) as the only large mammal. This mammal went extinct rapidly after human arrival. Several hypotheses have been proposed to explain the extinction of M. balearicus. For the present study ancient DNA analysis (Sanger sequencing, Roche-454, Ion Torrent), and pollen and macrofossil analyses were performed on preserved coprolites from M. balearicus, providing information on its diet and paleo-environment. The information retrieved shows that M. balearicus was heavily dependent on the Balearic box species Buxus balearica during at least part of the year, and that it was most probably a browser. Hindcast ecological niche modelling of B. balearica shows that local distribution of this plant species was affected by climate changes. This suggests that the extinction of M. balearicus can be related to the decline and regional extinction of a plant species that formed a major component of its diet. The vegetation change is thought to be caused by increased aridity occurring throughout the Mediterranean. Previous hypotheses relating the extinction of M. balearicus directly to the arrival of humans on the islands must therefore be adjusted.     

Reconstruction of climate and vegetation changes of Lake Bayanchagan (Inner Mongolia): Holocene variability of the East Asian monsoon

A high-resolution pollen and Pediastrum record, spanning 12,500 yr, is presented for Lake Bayanchagan (115.21°E, 41.65°N, and 1355 m a.s.l.), southern Inner Mongolia. Individual pollen taxa (PT-MAT) and the PFT affinity scores (PFT-MAT) were used for quantitative climatic reconstruction from pollen and algal data. Both techniques indicate that a cold and dry climate, similar to that of today, prevailed before 10,500 cal yr B.P. The wettest climate occurred between 10,500 and 6500 cal yr B.P., at which time annual precipitation was up to 30–60% higher than today. The early Holocene increases in temperature and precipitation occurred simultaneously, but mid-Holocene cooling started at approximately 8000 cal yr B.P., 1500 yr earlier than the drying. Vegetation reconstruction was based on the objective assignment of pollen taxa to the plant functional type. The results suggest that this region was dominated by steppe vegetation throughout the Holocene, except for the period 9200 to 6700 cal yr B.P., when forest patches were relatively common. Inner Mongolia is situated at the limit of the present East Asian monsoon and patterns of vegetation and climate changes in that region during the Holocene probably reflect fluctuations in the monsoon's response to solar insolation variations. The early to middle Holocene monsoon undoubtedly extended to more northern latitudes than at present.     

Seasonality change and the third millennium BP rainforest crisis in southern Cameroon (Central Africa)

The third millennium BP crisis of the central African rainforest is not sufficiently understood. The low resolution of most pollen profiles and a large plateau of the calibration curve aggravate the exact dating of the event, and its causal climatic parameters are debated. We present a high-resolution pollen profile from the swamp site Nyabessan in the southern Cameroonian rainforest, covering the period 3100-2300 cal yr BP. Between 3100 and 2500 cal yr BP, the climate was favourable for a regional evergreen forest with Caesalpiniaceae and Lophira and a local Raphia swamp forest. Around 2500/2400 cal yr BP, a significant decrease of mature forest and swamp forest taxa and an increase of pioneers indicate that the rain forest was seriously disturbed and replaced by secondary formations. The dominance of Trema orientalis, a pioneer well adapted to seasonal desiccation, points to a much more accentuated seasonality after 2500 cal yr BP, which seems to be linked to a southwards shift of the ITCZ during the northern hemisphere winter months. We propose that the rain forest crisis between 2500 and 2200 cal BP created favourable conditions for farming and paved the way for a major expansion of Bantu speaking populations.     

Forest–savanna–morichal dynamics in relation to fire and human occupation in the southern Gran Sabana (SE Venezuela) during the last millennia

The southern Gran Sabana (SE Venezuela) holds a particular type of neotropical savanna characterized by the local occurrence of morichales (Mauritia palm swamps), in a climate apparently more suitable for rain forests. We present a paleoecological analysis of the last millennia of Lake Chonita (4°39′N–61°0′W, 884 m elevation), based on biological and physico-chemical proxies. Savannas dominated the region during the last millennia, but a significant vegetation replacement occurred in recent times. The site was covered by a treeless savanna with nearby rainforests from 3640 to 2180 cal yr BP. Water levels were higher than today until about 2800 cal yr BP. Forests retreated since about 2180 cal yr BP onwards, likely influenced by a higher fire incidence that facilitated a dramatic expansion of morichales. The simultaneous appearance of charcoal particles and Mauritia pollen around 2000 cal yr BP supports the potential pyrophilous nature of this palm and the importance of fire for its recent expansion. The whole picture suggests human settlements similar to today – in which fire is an essential element – since around 2000 yr ago. Therefore, present-day southern Gran Sabana landscapes seem to have been the result of the synergy between biogeographical, climatic and anthropogenic factors, mostly fire.     

A Late Holocene Paleoecological Record from Torrey Pines State Reserve, California

Paleoenvironments of the Torrey Pines State Reserve were reconstructed from a 3600-yr core from Los Peñasquitos Lagoon using fossil pollen, spores, charcoal, chemical stratigraphy, particle size, and magnetic susceptibility. Late Holocene sediments were radiocarbon dated, while the historical sediments were dated using sediment chemistry, fossil pollen, and historical records. At 3600 yr B.P., the estuary was a brackish-water lagoon. By 2800 yr B.P., Poaceae (grass) pollen increased to high levels, suggesting that the rising level of the core site led to its colonization by Spartina foliosa (cord-grass), the lowest-elevation plant type within regional estuaries. An increase in pollen and spores of moisture-dependent species suggests a climate with more available moisture after 2600 yr B.P. This change is similar to that found 280 km to the north at 3250 yr B.P., implying that regional climate changes were time-transgressive from north to south. Increased postsettlement sediment input resulted from nineteenth-century land disturbances caused by grazing and fire. Sedimentation rates increased further in the twentieth century due to closure of the estuarine mouth. The endemic Pinus torreyana (Torrey pine) was present at the site throughout this 3600-yr interval but was less numerous prior to 2100 yr B.P. This history may have contributed to the low genetic diversity of this species.     

Pollen‐based biome reconstructions for Colombia at 3000, 6000, 9000, 12 000, 15 000 and 18 000 14C yr ago: Late Quaternary tropical vegetation dynamics

Colombian biomes are reconstructed at 45 sites from the modern period extending to the Last Glacial Maximum (LGM). The basis for our reconstruction is pollen data assigned to plant functional types and biomes at six 3000‐yr intervals. A reconstruction of modern biomes is used to check the treatment of the modern pollen data set against a map of potential vegetation. This allows the biomes reconstructed at past periods to be assessed relative to the modern situation. This process also provides a check on the a priori assignment of pollen taxa to plant functional types and biomes. For the majority of the sites, the pollen data accurately reflect the potential vegetation, even though much of the original vegetation has been transformed by agricultural practices. At 18 000 ¹⁴C yr BP, a generally cool and dry environment is reflected in biome, assignments of cold mixed forests, cool evergreen forests and cool grassland–shrub; the latter extending to lower altitudes than presently recorded. This signal is strongly recorded at 15 000 and 12 000 ¹⁴C yr BP, the vegetation at these times also reflecting a relatively cool and dry environment. At 9000 ¹⁴C yr BP there is a shift to biomes thought to result from slightly cooler environmental conditions. This trend is reversed by 6000 ¹⁴C yr BP; most sites, within a range of different environmental settings, recording a shift to more xeric biome types. There is an expansion of steppe and cool mixed‐forest biomes, replacing tropical dry forest and cool grassland–shrub biomes, respectively. These changes in biome assignments from the modern situation can be interpreted as a biotic response to mid‐Holocene climatic aridity. At 3000 ¹⁴C yr BP the shift is mainly to biomes characteristic of slightly more mesic environmental conditions. There are a number of sites that do not change biome assignment relative to the modern reconstruction, although the affinities that these sites have to a specific biome do change. These ‘anomalies’ are interpreted on a site‐by‐site basis. Spatially constant, but differential response of the vegetation to climatic shifts are related to changes in moisture sources and the importance of edaphic controls on the vegetation. The Late Quaternary reconstruction of large‐scale vegetation dynamics in Colombia allows an understanding of the environmental controls on these to be developed. In particular, shifts in the character of the main climatic systems that influence Colombian vegetation are described. Copyright © 2002 John Wiley & Sons, Ltd.