中国泥盆纪维管植物的组成与古地理分布

依据泥盆纪早、中、晚3个时期的中国岩相古地理图及中国泥盆纪植物已有的资料,绘制出中国泥盆纪这3个时期的植物古地理分布图。中国泥盆纪海侵的主要方向是从中国西南部的云南地区（早泥盆世）逐渐向长江中游的华中地区（中泥盆世）及长江下游的华东地区（晚泥盆世）发展。中国泥盆纪维管植物在南方的分布是随着海侵,逐渐向华中及华东地区扩展,由早泥盆世西南地区的云南、四川、贵州,以及广西和湖南南部等地区,发展至中泥盆世云南、四川、贵州、湖南的中北部以及湖北的南部地区,在晚泥盆世的分布范围扩大到广东、湖南、湖北、江西、安徽、浙江以及江苏等地区。泥盆纪维管植物在北方的分布范围较小,从中泥盆世才开始出现,主要分布在新疆的北部,晚泥盆世在宁夏等地区也有分布。中国泥盆纪3个时期陆地维管植物各个种类所占的百分含量表明,早期的植物主要以较为原始的莱尼蕨类、工蕨类以及前石松类为主,中期石松类占优势,到了晚期石松类依然占优势,而楔叶类、真蕨类以及前裸子植物开始繁盛。     

贵州都匀下泥盆统蟒山群Zosterophyllum australianum(澳大利亚工蕨)新材料及其古地理意义*

Zosterophyllum(工蕨属)是早期陆生植物的代表性类群之一,全球广泛分布。Z.australianum(澳大利亚工蕨)产自华南和澳大利亚,是东北冈瓦纳古植物地理区系的典型分子。文中描述了产自贵州都匀包阳剖面蟒山群中的Z.australianum新材料。该标本的孢子囊具短柄,呈椭圆形或扇形,宽2.6~4.1 mm,高可达3.9 mm,加厚带宽约0.6 mm,紧密螺旋排列形成孢子囊穗,与云南文山早泥盆世坡松冲植物群中的Z. australiaunum极为相似。依据目前对Z. australianum时代延限的认识,并结合蟒山群其他植物属种的发现(如Adoketophyton subverticillatum),推断该群下段的时代为早泥盆世布拉格期。Z. australianum在蟒山群中的发现,扩展了该植物的地理分布范围,反映出蟒山群中的植物组合与坡松冲植物群存在一定联系。     

贵州都匀下泥盆统蟒山群Zosterophyllum australianum(澳大利亚工蕨)新材料及其古地理意义*

Zosterophyllum(工蕨属)是早期陆生植物的代表性类群之一,全球广泛分布。Z.australianum(澳大利亚工蕨)产自华南和澳大利亚,是东北冈瓦纳古植物地理区系的典型分子。文中描述了产自贵州都匀包阳剖面蟒山群中的Z.australianum新材料。该标本的孢子囊具短柄,呈椭圆形或扇形,宽2.6~4.1 mm,高可达3.9 mm,加厚带宽约0.6 mm,紧密螺旋排列形成孢子囊穗,与云南文山早泥盆世坡松冲植物群中的Z. australiaunum极为相似。依据目前对Z. australianum时代延限的认识,并结合蟒山群其他植物属种的发现(如Adoketophyton subverticillatum),推断该群下段的时代为早泥盆世布拉格期。Z. australianum在蟒山群中的发现,扩展了该植物的地理分布范围,反映出蟒山群中的植物组合与坡松冲植物群存在一定联系。     

中国云南早泥盆世坡松冲植物群——探究早期陆生维管植物演化分异的窗口

简要回顾了目前云南早泥盆世坡松冲植物群的研究，论述了坡松冲植物群的时代、组成和性质。研究表明坡松冲植物群在某些简单的、常见的工蕨类植物和三枝蕨类植物组成的背景下，含有具复杂形态和结构特征的植物类群，显示了较劳亚大陆同期陆生维管植物更进化的营养器官、生殖器官分异的水平。不同古植物地理区的长期隔离（也就是，早泥盆世的东北冈瓦纳古植物地理区和华南亚区）和独特的环境可能是造成众多地方性分子演化分异的原因。坡松冲植物群的研究表明中国华南可能是早期陆生维管植物演化辐射的中心之一。     

内蒙古乌达地区早二叠世植物群和古生态研究

研究的植物化石采自内蒙古乌达地区早二叠世地层,共计11属20种,分属于石松纲、楔叶纲、瓢叶目、真蕨纲、种子蕨纲和苏铁纲。研究结果表明,植物群含有许多华夏植物群分子和地方性分子,为典型的华夏植物群。根据植物群分子的总体组成,植物群的时代为早二叠世,相当于阿瑟尔期(Asselian)、萨克马尔期(Sakmarian)、亚丁斯克期(Artinskian)和空谷期(Kungurian)。在研究植物群的基础上,对植物群的古生态进行了研究,建立了早二叠世Lepidodendron posthumii Paratingia datongensis植物群落。这一植物群落在二叠纪主要分布在沼泽和低地环境,以大量出现的蕨类和种子蕨分子为特征。植物群落为原地保存,借助于化石资料,植物的生活型得以重建,包括乔木、树蕨、灌木和草本。植物群落指示早二叠世本区为温暖潮湿的气候条件。     

中国斯蒂芬期植物群的基本特征

重新定义后的中国斯蒂芬期植物群所对应的岩石地层单元只限于太原组下部晋祠段。该植物群以石松纲、楔叶纲、真蕨纲和种子蕨纲为主，也有一定数量的瓢叶纲和科达纲植物分子。尽管中国斯蒂芬期植物群与欧美同期植物群有一定共性，但二者差别仍很明显。     

新疆早侏罗世植物群特征

本文根据新疆早侏罗世各组中所产的植物化石,提出了疆新早侏罗世植物群分属于毛羽叶—锥叶蕨植物群与锥叶蕨—拟刺葵植物群的过渡植物群和锥叶蕨—拟刺葵植物群.过渡植物群中苏铁类比较丰富.锥叶蕨—拟刺葵植物群以真蕨类和银杏类为主,真蕨类中锥叶蕨,托第蕨,枝脉蕨最为发育.两个植物群按照不同层位均可分出上下两个组合.本文还探讨了八道湾组时代,认为无疑属早侏罗世,塔里奇克组植物组合也应属早侏罗世(塔里奇克组的地质时代为T_(?)—J_1).早侏罗世早期新疆气侯曾一度炎热,向后转为湿暖潮湿.     

中国滇东南早泥盆世坡松冲组的一个石松植物新属种——辐射正理木(Zhenglia radiata gen.etsp.nov.)及其演化意义

本文描述了一个产自中国滇东南文山地区坡松冲组(早泥盆世布拉格期)的新植物属、种——辐射正理木(Zhenglia radiatagen.etsp.nov.)。这是一个小型草本石松植物。气生枝分化出营养轴和生殖轴,茎轴上叶迹的排列显示出似鳞木的叶序。叶迹可能具有叶舌穴,小型叶由突出和下延的基部和一个全缘的长叶片所组成。生殖轴由螺旋排列的孢子叶组成,它们形成一个紧密的似孢子叶球的结构。每个孢子叶具有侧向的延展和远端的叶片。长卵形的孢子囊近轴面着生在孢子叶上。我们将这个新植物置于原始鳞木目中。考虑到形态的相似性,特别是未分叉的小型叶具有似…     

晚古生代中国大陆与欧美地区植物群之间的差异及其原因

晚古生代同处于赤道或低纬度地带的中国大陆植物群与欧美地区同期植物群之间总存在一定的差异。中泥盆世－早石炭世维宪期,中国大陆以石松类的大量繁育和高度分异为显著特色,而同期的欧美地区植物群以古蕨类和前裸子植物为其主体;纳缪尔期－二叠纪,大量东亚型鳞木类植物及许多地方性植物属相继在中国大陆出现,使它和欧美植物群之间差距进一步拉大。     

新疆吐哈盆地中侏罗世植物化石

研究记录了新疆吐哈盆地三道岭煤田西山窑组的 4 3属 86种植物化石。该植物化石组合以真蕨和银杏类最为繁盛 ,苏铁类和松柏类较发育 ,楔叶类较少 ,另有少量苔藓类和石松类。真蕨类又以蚌壳类占优势 ,出现大量中侏罗世分子 ,属于中国北方温带植物地理区 Coniopteris—Phoenicopsis植物群的典型组合 ,表明其时代为中侏罗世阿连期至巴柔期     

A New Lycopsid, Zhenglia radiata gen. et sp. nov., from the Lower Devonian Posongchong Formation of Southeastern Yunnan, China, and Its Evolutionary Significance

A new plant, Zhenglia radiata gen. et sp. nov., from the Posongchong Formation （Early Devonian, Pragian age） of the Wenshan District of southeastern Yunnan Province, China is a small herbaceous lycopsid. The aerial branches divide into foliar and fertile stems. The arrangement of the leaf bases on the stem shows lepidodendroid-like phyllotaxy. Possibly the leaf base bears a ligulate pit. The microphyll consists of a swollen, decurrent base, and an entire, linear lamina. The fertile axis is composed of sporophylls arranged helically, forming a compact area similar to cone-like structure. Each sporophyll consists of a widened lateral extension and a distal lamina. The ovoid-elongate sporangia are attached adaxially on the surface of sporophylls. Based on morphological similarities, especially the undivided microphylls, the style of phyllotaxy, the form of the cone-like structures and sporangia, the new lycopsid Zhenglia radiata is placed in the order Protolepidodendrales and a possible evolutionary relationship with the arborescent lycopsids of the Late Devonian and Carboniferous is considered.     

A New Zosterophyll Plant, Ramoferis gen. nov., from the Posongchong Formation of Lower Devonian (Pragian) of Southeastern Yunnan, China

A new genus and species,Ramoferis amalia gen.et sp.nov.,is described from the Early Devonian(Pragian)Posongchong Formation,Wenshan district of southeastern Yunnan,China.This plant has isodichotomous major axes,which also divide anisotomously distally to produce closely spaced lateral branches,often within fertile regions.The stalked round to reniform sporangia,borne laterally and spirally over several levels of branches,are distantly to moderately spaced or aggregated into distinct spikes.The stalks incr...     

Researches on Plants from the Lower Devonian Xujiachong Formation in the Qujing District, Eastern Yunnan

Fossil plants Hedeia sinica Hao et Gensel 1998, Huia gracilis Wang et Hao 2001 and Guangnania cuneata Wang et Hao are described from the Lower Devonian Xujiachong Formation, the Qujing district, eastern Yunnan, China. They contribute to our knowledge of the flora in this district. Based on the occurrence of common plants (Hedeia, Huia, Guangnania and Zosterophyllum australianum) and their horizons, it is proved for the first time that the mid-lower assemblage of the Xujiachong flora is comparable with the Early Devonian ((late) Pragian) Posongchong flora of southeastern Yunnan and the upper Baragwanathia flora of Australia. In view of this fact and the stratigraphic sequence, the mid-lower part of the Xujiachong Formation is considered to be of (late) Pragian age. Through comprehensive analyses of plant, bivalve and fish assemblages and the lithology, the upper part of the Xujiachong Formation is dated to be of early Emsian age. The Xujiachong flora belonged to the northeastern Gondwana palaeophytogeogr     

Two Zosterophyll Plants from the Lower Devonian (Lochkovian) Xitun Formation of Northeastern Yunnan, China

Abstract: Two zosterophyll plants are described from the Lower Devonian (Lochkovian) Xitun Formation of Qujing, Yunnan, China. Xitunia spinitheca gen. et sp. nov. has stalked sporangia laterally attached on the axis in a helical arrangement. Sporangia are dorsoventrally flattened and composed of two unequal valves; the adaxial valve is round in face view, while the abaxial valve is larger than the former, triangular or wedge-shaped, and radially bears long spiny appendages along the distal margin. Xitunia shows new variation of sporangial morphology within the zosterophylls. Zosterophyllum minorstachyum sp. nov. has K-shaped branchings at the basal parts and small-sized terminal spikes, which consist of round to elliptical sporangia arranged helically. This paper provides new data on the diversity of plant types during Lochkovian when rare vascular plants were reported. As for various species of Zosterophyllum in South China, their apparent evolutionary trend of features from the Late Silurian to Early Devonian (Emsian) is discussed.     

The oldest flora of the South China Block, and the stratigraphic bearings of the plant remains from the Ngoc Vung Series, northern Vietnam

Several outcrops of the Late Silurian and Devonian of the Ngoc Vung Series, northern Vietnam, yielded plant remains. The Late Silurian localities delivered the earliest known flora of the South China block. Although the fossils are fragmentary, they complement our knowledge about the global composition of the flora. The major components of the flora are plants with dichotomous habit and terminal bivalvate sporangia, which are close relatives to zosterophylls, and zosterophylls. Plants with possible euphyllophyte affinities and bryophytes are occasionally present. This floral composition is similar to that of the rich, younger South China block assemblages from the Posongchong and Xujiachong Formations of China, considered Pragian in age. The South China block flora is therefore likely to have been dominated by zosterophylls and pre-zosterophylls at least from the Late Silurian to the Pragian (i.e. a 20 million years long period). It also strengthens the hypothesis that more derived plants were present on eastern Gondwana earlier that elsewhere, in the first steps of tracheophyte evolution. The Devonian localities of the Ngoc Vung Series delivered a thick fibrous stem fragment and a basal euphyllophyte. These latter plant remains provide some stratigraphic data. The large stem fragment is consistent with an Eifelian age for the Duong Dong Formation (part of the Ngoc Vung Series), as suggested by the brachiopod fauna. The accompanying basal euphyllophyte displays a combination of characters (axes 3-4 mm wide and lateral branchings) that is also consistent with an Eifelian age, but possibly more characteristic of the Emsian flora. It is therefore suggested that the stratigraphic range of the Duong Dong Formation might be extended down to the Emsian.     

Birth and early evolution of metazoans

The reconstruction of the phylogenetic tree of animals (TOA) has long been one of the central interests in biological and paleobiological sciences. We review the latest results of paleontological and stratigraphical studies on the Ediacaran–Cambrian sequences mainly from South China for revising the TOA in accordance with modern genome biology. A particular focus is given to the pattern of animal diversification based on the fossil first appearances of high-rank clades chiefly in phylum-level. The results show an abrupt divergence of lineages during the Ediacaran–Cambrian transition; however, the appearances of metazoan phyla were obviously diachronous, with three major phases recognized herein. The first phase is marked by the appearances of basal metazoan phyla in the latest Ediacaran. Very few unequivocal bilaterian clades were present at this phase. The second phase occurred in the Terreneuvian (Cambrian Stages 1–2), represented by the occurrences of many lophotrochozoan lineages. This phase also involves the appearances of calcified basal metazoan lineages, and possibly, those of contentious ecdysozoans in the latest Terreneuvian, but no deuterostome has been known from this age. The third and also the largest phase occurred in the Cambrian Stage 3, which involve all the three supraphylogenetic clades of the Eubilateria. A number of lophotrochozoan lineages, the bulk of ecdysozoans, and all deuterostome phyla, appeared for the first time in this phase. Since there is no unambiguous evidence for bilaterians in the Ediacaran, the Cambrian explosion sensu stricto was an abrupt diversification of bilateral lineages in a short time of ca. 25 million years across the Ediacaran–Cambrian boundary. Next critical issues in research include high-resolution chrono- and chemostratigraphic analyses, correlations between biotic events and environmental perturbations, physiological approach to the biological connotation of biomineralization, and exploration for the lost mid-oceanic biota and environments, which are crucial in understanding the entire picture of the Cambrian explosion.     

中国瓣甲鱼类化石的生物地层学意义

瓣甲鱼类在我国泥盆系中分布较广,具有相当重要的生物地层学意义。根据化石在地层中的分布,我国的辦甲鱼类自上而下可分为5个组合(带):拟辦鱼一广西辦甲鱼(Quasipetalichthys-Guangsipetalichthys)组合,宽头鱼(Eurycaraspis)带,中华瓣甲鱼-西南瓣甲鱼(Sinipetalichthys-Xinanpetalichthys)组合,新瓣鱼(Neopetalichthys)带,滇东瓣甲鱼(Diandongpetalichthys)带。它们的时代分别相当于吉维期,吉维期早期,布拉格期,布拉格期早期和洛赫科夫期早期。     

广西南宁下泥盆统华南鱼类化石新发现及其地层和古地理意义*

笔者描述了广西南宁下泥盆统布拉格阶那高岭组下部盔甲鱼类的新发现,建立了华南鱼科2个新种——丘比特华南鱼(Huananaspis cupido sp. nov.)和梁氏亚洲鱼(Asiaspis liangi sp. nov.)。前者以头甲个体较小、不发育中背棘、两角末端超出头甲后缘等特征区别于属型种武定华南鱼(H. wudingensis),后者以头甲长大于宽、角向头甲侧后方延伸、角后缘发育锯齿、中背孔呈纵长椭圆形等特征区别于属型种宽展亚洲鱼(A. expansa)。广西南宁地区那高岭组具有可靠的地质时代证据,其中所产的徐家冲鱼类化石组合能够为华南地区早泥盆世地层的对比研究提供鱼类化石标准,而华南鱼属(Huananaspis)和亚洲鱼属(Asiaspis)在那高岭组的发现,丰富了华南鱼类的多样性,扩大了徐家冲鱼类组合的化石内容,并为云南武定坡松冲组与广西南宁那高岭组之间的对比提供了直接化石证据。广西南宁莲花山组与那高岭组过渡层中的徐家冲鱼类化石组合物种多样性最高,几乎涵盖了其他地区该组合的所有属种,说明广西南宁地区可能是盔甲鱼类辐射和扩散的中心之一。该地区早泥盆世布拉格期的古鱼类、早期维管植物和腕足动物均表现出显著的地方性色彩,可能跟华南海是一个半封闭的陆表海湾有关,即由于封闭海效应,鱼类在这个封闭的环境中独立演化,表现出很高的多样性和很强的土著性,最终在华南板块上形成一个独立的动物区系——“盔甲鱼—云南鱼”动物区系。