林木多样性模型及生长模拟

林木生存环境复杂多变,导致林木形态千差万别,为了准确描述林木的形态多样性特征,提出了一种基于模型分解的林木生长模拟方法。首先构建多样性林木三维模型,使用林木形态的实际测量数据,基于B样条函数约束树干、树冠形态,依据削度方程模拟树干直径的变化趋势,利用Direct3D实现林木形态的三维建模;其次,按林木形态特征进行林木模型分解,将林木模型分为9个子模型;最后,使用场景节点按照拓扑结构关系对子模型进行动态组织,基于胸径、树高、枝下高、冠高和冠幅生长模型,使用多核CPU多线程并行的计算方式,获得各年龄阶段林木东、南、西、北方向形态参数,从而对子模型生长状态进行控制,实现林木不同方向生长变化的可视化模拟。结果表明：该方法可将林木三维模型与生长模型紧密结合起来,林木各方向的枝下高、冠高和冠幅严格按照生长模型进行生长,可有效模拟出林木各方向生长状态的多样性;三维场景渲染帧率（FPS）保持在25以上,平均FPS可以达到50,实现林木生长的方向异质性模拟。     

Forest structure of a maple old-growth stand: a case study on the Apennines mountains (Southern Italy)

In Europe, very small forest areas can be considered to be old-growth, and they are mainly located in Eastern Europe. The typical structures of old growth forests infrequently occur in Mediterranean mountainous environments, since they have been affected by human activities for centuries. This study focused on a remote and almost pure Italian maple stand located in southern Italy, which has not been managed for long time due to its inaccessibility. The effects of natural evolution on the forest stand were evaluated through the analysis of the spatial and chronological structure and the regeneration patterns, then estimating the amounts and quality of deadwood occurrence. Across the whole stand, all the trees with DBH (diameter at breast height) larger than 50 cm (LLT, large living trees) were measured (DBH and height) and age was also determined through a dendrochronological approach. The diameters observed ranged between 50 and 145 cm with ages of 120 to 250 years. The Latham index calculated for trees within the sample plot highlighted a multilayered canopy with a dominant layer of large living trees (age > 120 years). The size-class distribution of stems had a reverse-J shape, and basal area was 52 m² ha^-1. Deadwood was exclusively constituted by standing dead trees and CWD and its volume was on average 31 m³ ha^-1.Pure Italian maple forests are generally rare in Europe, and it was unexpected to find a forest stand characterized by a so complex structure with old growth attributes. The study of complex forest stand, even if small, could give precious information on the forest evolution, clarifying also diverse auto-ecological traits of tree species that usually are not common in our forests.     

Integrated Analysis of the Trees and Associated UnderCanopy Species in a Subalpine Forest of Western Himalaya,Uttarakhand, India

Subalpine forests are known as outstanding habitats due to co-existence of both temperate and alpine vegetation and are classic example of ecotonal zones. Limited but diverse physiognomy of trees inhabiting in subalpine forest results in variability within under-canopy habitat conditions. Studies were undertaken to assess population status, habitat preference and interferences to the trees and associated undercanopy herbs in a subalpine forest of western Himalaya. A total of 10 woody and 23 under-canopy herbs were recorded in the selected subalpine forest area. At each stand, the number of tree species and under-canopy herbs ranged from 2 to 4 and 8 to 10 respectively. Abies spectabilis, Acer caesium, Quercus floribunda, Q. semecarpifolia and Rhododendron arboreum were key tree species in this area. The density of main woody species was 280 to 1190 individuals ha-1 at different stands. Herbaceous plants with rosette and clump growth habits were observed to have higher values for total basal cover and importance value index. Presence of some undercanopy herbs like; Dactylorhiza hatagirea, Malaxis muscifera, Picrorhiza kurrooa, Polygonatumcirrhifolium and Skimmia laureola in the specific habitats also showed that they are habitat specific.However, the presence of Frageria nubicola and Viola sp. was common in the selected stands. Felling of trees for timber, construction of temporary huts,fuel wood and lopping for fodder were main interferences for trees. On the other hand, trampling driven damage due to grazing, habitats degradation and overexploitation were observed key threats for under-canopy herbs. Integrated analysis including population studies, habitats preference and interferences to the trees and under-canopy herbs in this sensitive and important ecosystem will be useful for determining the conservation plans and ecosystem management.     

Crown fuel characteristics and carbon emission from Japanese red pine stands burned by crown fire in Mt. Palgong, South Korea

Carbon emissions from forest fires are considered an important factor of ecosystem carbon balance and global climate change. Carbon emissions from Japanese red pine stands (Pinus densiflora S. et Z.) burned by crown fire were estimated at Mt. Palgong in Daegu Metropolitan City, and crown fuel characteristics, including crown bulk density, crown base height, and fuel moisture content of Japanese red pine, were analyzed. Total biomass combusted was calculated by subtracting the biomass of burned stands from that of unburned stands exhibiting similar stand structures and site environments. Ten trees in the unburned area and five trees in the burned area were cut by using direct harvesting techniques to estimate crown layer biomass. All biomass sampled was oven-dried and weighed. The dry weight ratios of stems, branches, and needles were 70%, 21%, and 9%, respectively. The available fuel load susceptible to combustion during the crown fire spread was equivalent to 55% of the crown layer biomass. The crown bulk density was 0.24 kg/ m 3 on average. The estimated amount of CO 2 was 23,454 kg CO 2 /ha for the crown layer. These results will be useful for calculating the amount of CO 2 emitted from forest fires and for developing a forest carbon model in P. densiflora forests.     

Thinning effects on walnut stem and crown diameter growth and fruiting in the walnut-fruit forests of Kyrgyzstan

The walnut-fruit forests (Juglans regia L.) in southern Kyrgyzstan are considered to be biodiversity hotspots and of considerable economic and ecological importance to local populations. The fruiting ability and stability of these forests are, however, decreasing because of infrequent silvicultural interventions and non-existent natural regeneration. Continuous inventories and walnut harvest measurements were carried out on 18 trial and control plots over the course of 13 year. Results showed that i) thinning had a positive influence on diameter at breast height (dbh) increase, ii) dbh growth and crown area increases are strongly and positively correlated, iii) large crowns enhanced the production of fruits, and iv) temperature conditions had a major influence on the production of fruits. The above-mentioned points are particularly relevant for young plantations but are also valid for relatively old plantations (> 50 years). We conclude that regular thinning substantially improves diameter growth, fruit production and ability of trees to withstand wind and snow damage in both young stands and in older walnut-fruit forests.     

Effects of environmental factors on tree seedling regeneration in a pine-oak mixed forest in the Qinling Mountains, China

Natural seedling regeneration and tree establishment are affected by various environmental factors.In this study,we established eight,eight,six,and four independent forest stands(each stand was further divided into five subplots) respectively based on the altitudinal gradient,stand density,slope location,and slope aspect to investigate the effects of environmental factors on tree seedling regeneration in a pine-oak mixed forest.The results indicated that the seedling density was significantly higher at altitudes of 1,283 m to 1,665 m,whereas the sapling density did not differ with altitudes.The seedling and sapling density decreased significantly at 1,835 m.The seedling densities on the upper slopes were much higher than those on the middle and lower slopes,whereas the sapling density had no difference.The sapling density decreased gradually from the southwest(20°-75°) whereas it increased on the shady slopes to the northeast(40°).The seedling density increased from southwest(20°) to northeast(40°).The seedling and sapling densities increased with the stand density(850 trees ha-1to 1,525 trees ha-1) whereas the sapling density was significantly lower in stands(1,900 trees ha-1).Principal components analysis showed that the slope aspect and stand density had more important roles in tree regeneration in this study region compared with the other two factors.Therefore,our findings suggest that it will be beneficial to keep stands at a moderate density on shady slopes.Appropriate thinning of higher density stands is also expected to promote the natural regeneration of pine-oak mixed forest.     

基于标记控制分水岭分割方法的高分辨率遥感影像单木树冠提取

树冠是树木的重要组成,可以直接反映树木健康状况。高空间分辨率遥感影像和遥感技术为快速获取详细的树冠信息和实时监测林冠变化提供了有效的途径。因此,基于高空间分辨率遥感影像的单木树冠提取方法研究对现代森林管理具有重要意义。本文以黄河三角洲地区孤岛林场人工刺槐林和旱柳为研究对象,以QuickBird影像为数据源,首先利用面向对象方法实现研究区林地和非林地分类;然后以林地为掩膜,提取出树冠分布范围;在此基础上,分别选取疏林区和密林区为试验区域,通过形态学开闭重建滤波,平滑图像,去除噪声;最后,利用标记控制分水岭分割方法分别对疏林区和密林区进行树冠提取。本文以人工勾绘结果为参考进行精度验证,结果显示疏林区F测度达到87.8%,密林区F测度达到65.5%,表明该提取方法简单易行,精度可靠。     

顾及单木三维形态的无人机立体影像单木识别算法

单木参数对当前的森林资源管理、生态研究以及生物多样性保护等具有重要意义。无人机立体影像数据与单木识别算法为单木参数的低成本、自动化获取提供了基础。现有研究表明,常用的基于局部最大值搜索的单木识别算法面对密集林分时存在严重的漏识别问题,影响了参数提取的精度,因此本文提出了顾及单木三维形态的无人机立体影像单木识别新算法。算法首先综合利用无人机立体影像的高程与RGB光谱信息,通过随机森林分类进行林冠区的提取;然后利用形态学的多层腐蚀、膨胀与连通区标记进行树冠相连单木的分离与树冠中心点的提取,从而实现单木自动化识别。本文选取内蒙古大兴安岭林区和四川王朗林区的4块样地进行验证,以目视解译数据为参考,分别与基于高程值的局部最大值搜索算法（算法A）、基于RGB光谱亮度值的局部最大值搜索算法（算法B）进行比较。结果显示：本文提出的算法在4个样地的平均F1-score为94.17%,与算法A和算法B相比分别提高了15.85%和9.37%;而对于密集样地,本文提出的算法在查全率上相比算法A和算法B分别提高51.79%和35.64%。结果表明本文提出的算法在不同林区均能够实现较好的单木识别效果,特别是能够有效避免密集林分下的漏识别问题,为基于无人机立体影像的单木识别研究提供了一种新的思路。     

Tree wave migration across an elevation gradient in the Altai Mountains,Siberia

The phenomenon of tree waves(hedges and ribbons) formation within the alpine ecotone in Altai Mountains and its response to observed air temperature increase was considered. At the upper limit of tree growth Siberian pine(Pinus sibirica) forms hedges on windward slopes and ribbons on the leeward ones. Hedges were formed by prevailing winds and oriented along winds direction. Ribbons were formed by snow blowing and accumulating on the leeward slope and perpendicular to the prevailing winds, as well as to the elevation gradient. Hedges were always linked with microtopography features, whereas ribbons were not. Trees are migrating upward by waves and new ribbons and hedges are forming at or near tree line, whereas at lower elevations ribbons and hedges are being transformed into closed forests. Time series of high-resolution satellite scenes(from 1968 to 2010) indicated an upslope shift in the position ribbons averaged 155±26 m(or 3.7 m yr~(-1)) and crown closure increased(about 35%–90%). The hedges advance was limited by poor regeneration establishment and was negligible. Regeneration within the ribbon zone was approximately 2.5 times(5060 vs 2120 ha~(-1)) higherthen within the hedges zone. During the last four decades, Siberian pine in both hedges and ribbons strongly increased its growth increment, and recent tree growth rate for 50 year-old trees was about twice higher than those recorded for similarly-aged trees at the beginning of the 20~(th) century. Hedges and ribbons are phenomena that are widespread within the southern and northern Siberian Mountains.     

Structure and regeneration dynamics of three forest types at different succession stages of spruce – fir mixed forest in Changbai Mountain,northeastern China

Analyzing and understanding the structure and growth dynamics of forests at different stages is helpful to promote forest succession, restoration and management. Three spots representing three succession stages of spruce-fir mixed forest(SF: polar-birch secondary forest, MF: spruce-fir mixed forest and PF: spruce-fir near primary forest) were established. Structure, growth dynamics during two growth seasons for dominant tree species, regeneration were examined, and a univariate O-ring function statistic was used to analyze the spatial patterns of main regeneration tree species. Results showed that,(1) composition of tree species, periodic annual increment(PAI) of the diameter at breast height(DBH), basal area for overstory trees and of ground diameter(DGH) for saplings, were significantly different with the succession;(2) the current species composition and regeneration dynamics of SF suggested a development towards spruce-fir mixed forests. Pioneer species like Betula platyphyllaa will gradually disappear while climax species, such as Abies nephrolepis, Pinus koraiensis, Picea koraiensis and Tilia amurensis will dominate forest stands;(3) Despite the highest volume occurring in PF, and saplings in it grew better than in the others, this forest type is unstable because of its unsustainable structure of DBH class and insufficient regeneration; and(4) MF had the most reasonable distribution of DBH class for adult trees(DBH 5.0 cm) and DGH class for saplings(H ≥30 cm and DBH ≤5 cm), as well as an optimal volume increment. Limiting canopy opening size can lessen the physiological stress and promote the growth and competitive status of regeneration. Management implications for increasing the gaps and thus creating better growth conditions for understory saplings and facilitating forest succession were discussed.     

Assessing the influence of the minimum measured diameter on forest spatial patterns and nearest neighborhood relationships

Forest structure analysis is important for understanding the properties and development of a forest community,and its outcomes can be influenced by how trees are measured in sampled plots.Although there is a general consensus on the height at which tree diameter should be measured[1.3 m:diameter at breast height(DBH)],the minimum measureddiameter(MMD)often varies in different studies.In this study,we assumed that the outcomes of forest structure analysis can be influenced by MMD and,to this end,we applied g(r)function and stand spatial structural parameters(SSSPs)to investigate how different MMDs affect forest spatial structure analysis in two pine-oak mixed forests(30 and 57 years old)in southwest China and one old-growth oak forest(120years old)from northwest China.Our results showed that 1)MMD was closely related to the distribution patterns of forest trees.Tree distribution patterns at each observational scale(r=0-20 m)tended tobecome random as the MMD increased.The older the community,the earlier this random distribution pattern appeared.2)As the MMD increased,neighboring trees became more regularly distributed around a reference tree.In most cases,however,nearest neighbors of a reference tree were randomly distributed.3)Tree species mingling decreased with increasing diameter,but it decreased slowly in older forests.4)No correlations can be found between individual tree size differentiation and MMD.We recommend that comparisons of spatial structures between communities would be more effective if using a unified MMD criterion.     

结合DSM的机载LiDAR单木树高提取研究

机载LiDAR在提取地形坡度较大区域的冠层高度模型（CHM）时易产生畸变,降低单木树高的提取精度,为此提出一种CHM与数字表面模型（DSM）相结合的树高估算方法。首先基于预处理后的点云生成的CHM,利用局部最大值算法和标记控制分水岭分割算法进行分割,得到单木树冠轮廓多边形;然后结合DSM,采用固定窗口的局部最大值算法探测树顶点并提取其高程,继而与使用狄洛尼三角网和高程内插得到的地面点相减获取树高;最后,以广西兴安县富江村附近地形起伏较大的针叶林为试验区,测试3种不同坡度下,在CHM、CHM结合DSM获得的树高与实测树高分别进行精度分析。结果表明,当树木分别位于平均坡度为32°、27°和15°的试验区时,CHM中提取的树高与实测数据拟合的R²分别为0.84、0.85和0.87,RMSE为1.48、1.41和1.58 m,结合DSM后R²为0.92、0.91和0.93,RMSE为0.93、1.02和1.16 m;在地形坡度较大的区域,本文方法可以有效提高单木树高的估算精度。     

Alternative stable states in mountain forest ecosystems: the case of European larch (<Emphasis Type="Italic">Larix decidua</Emphasis>) forests in the western Alps

European larch (Larix decidua) forests of the western Alps form extensive cultural landscapes whose resilience to global changes is currently unknown. Resilience describes the capacity of ecological systems to maintain the same state, i.e., the same function, processes, structure, and composition despite disturbances, environmental changes and internal fluctuations. Our aim is to explore the resilience of larch forests to changes in climate and land use in the western Italian Alps. To do so, we examined whether larch forests can be described as an alternative stable state in mountain forest ecosystems. We used tree basal area data obtained from field forest inventories in combination with topography, forest structure, land use, and climate information. We applied three different probabilistic methods: frequency distributions, logistic regressions, and potential analyses to infer the resilience of larch forests relative to that of other forest types. We found patters indicative of alternative stable states: bimodality in the frequency distribution of the percent of larch basal area, and the presence of an unstable state, i.e., mixed larch forests, in the potential analyses. We also found: (1) high frequency of pure larch forests at high elevation, (2) the probability of pure larch forests increased mostly with elevation, and (3) pure larch forests were a stable state in the upper montane and subalpine belts. Our study shows that the resilience of larch forests may increase with elevation, most likely due to the altitudinal effect on climate. Under the same climate conditions, land use seems to be the main factor governing the dominance of larch forests. In fact, subalpine larch forests may be more resilient, and natural succession after land abandonment, e.g., towards Pinus cembra forests, seems slower than in montane larch forests. In contrast, in the upper montane belt only intense land use regimes characterized by open canopies and forest grazing may maintain larch forests. We conclude that similar approaches could be applied in other forest ecosystems to infer the resilience of tree species.     

生长模型驱动的单株杉木三维动态模拟

虚拟植物是一种潜在、有力的植物分析辅助工具,而单株木生长模拟是森林生态系统模拟的基础。为了动态模拟杉木生长发育过程,提出了参数化单株木三维形态结构建模和与距离无关的单木生长模型的集成方法,其主要过程为：首先,根据杉木的形态结构特征,采用参数化建模方法建立特定生长阶段的三维静态模型;其次,根据生长模型预测不同年龄杉木的树高、胸径、枝下高和分枝轮数,并与树木三维静态模型几何描述参数建立联动;最后,采用参数曲线调整干径和枝径变化、枝条长度、分枝角等形态结构参数,使模型形态随树龄增长而变化。在自主研发的ParaTree系统上,扩展杉木动态生长模拟模块,并以福建省漳平五一林场的二类调查数据为例,动态模拟了杉木的生长过程。模拟结果表明,本方法可直观表达林分中林木个体平均生长状况和大体形态结构特征。树木三维模型描述参数与传统树木统计生长模型结合,有利于重用林业领域淀积大量的生长模型。     

Succession Features and Dynamic Simulation of Subalpine Forest in the Gongga Mountain, China

The Gongga Mountain of eastern Tibet Plateau is a representative of the alpine regions with high peaks and deep valleys. Climate change over the last thousand years has controlled the dynamics of glacier and debris flow occurrence, which resulted in substantial changes in the mountainous environment. The authors surveyed the community structure of primary forests in Gongga Mountain and forest successign processes in woodland plots. The changing features in the subalpine environment are discussed in this paper. Tree species and sizes between the glacier shrinking areas and debris flow fans in Hailuogou Valley are compared. The pioneer species that settle in debris flow fans and the glacier shrinking areas are Salix spp. and Populus purdomii. Abies fabri and Picea brachytyla are the climax tree species. The succession process of primary vegetation in Hailuogou (2700 ～ 3200 m) can be divided into four stages:     

Alternative stable states in mountain forest ecosystems: the case of European larch(Larix decidua) forests in the western Alps

European larch(Larix decidua) forests of the western Alps form extensivecultural landscapes whose resilience to global changes is currently unknown. Resilience describes the capacity of ecological systems to maintain the same state, i.e., the same function, processes, structure, and composition despite disturbances, environmental changes and internal fluctuations. Our aim is to explore the resilience of larch forests to changes in climate and land use in the western Italian Alps.To do so, we examined whether larch forests can be described as an alternative stable state in mountain forest ecosystems. We used tree basal area data obtained from field forest inventories in combination with topography, forest structure, land use, and climate information. We applied three different probabilistic methods: frequency distributions, logistic regressions, and potential analyses to infer the resilience of larch forests relative to that of other forest types.We found patters indicative of alternative stable states: bimodality in the frequency distribution of the percent of larch basal area, and the presence of an unstable state, i.e., mixed larch forests, in the potential analyses. We also found:(1) high frequency ofpurelarchforestsathighelevation,(2)the probability of pure larch forests increased mostly with elevation, and(3) pure larch forests were a stable state in the upper montane and subalpine belts.Our study shows that the resilience of larch forests may increase with elevation, most likely due to the altitudinal effect on climate. Under the same climate conditions, land use seems to be the main factor governing the dominance of larch forests. In fact, subalpine larch forests may be more resilient, and natural succession after land abandonment, e.g., towards Pinuscembra forests, seems slower than in montane larch forests. In contrast, in the upper montane belt only intense land use regimes characterized by open canopies and forest grazing may maintain larch forests.We conclude that similar approaches could be applied in other forest ecosystems to infer the resilience of tree species.     

Reconstruction of debris-flow activity in a temperate mountain forest catchment of central Mexico

The use of dendrochronology to study and date geomorphic processes in volcanic environments is still incipient, even more so on the volcanic slopes covered by temperate forests in central Mexico. Mass movements, such as debris flows, often impact forest stands where they cause damage to individual trees, thereby generating growth disturbances(GD) in the tree-ring records. The identification and dating of GD enables reconstruction of the age of trees colonizing bare surfaces after major events, but also allows the assessment of the frequency or spatial distribution of past geomorphic process activity. Here we used increment cores from 65 Pinus leiophylla, Abies religiosa, and Alnus jorullensis trees growing in the Axal gorge, on the southern slopes of La Malinche volcano, to unravel past debris-flow activity both temporally and spatially. Based on the combination of GD records, a weighted tree response index(Wit), field evidence and hydrometeorological records, we reconstructed 23 debris flows since 1933.Interestingly, almost two-thirds of the reconstructed years with debris-flow activity in Axal gorge match with events recorded in Axaltzintle gorge located on the NE slopes of La Malinche. These findings suggest a regional triggering mechanism, most likely related to the occurrence of hurricanes. This research could be useful for disaster risk management of the La Malinche National Park.     

结合Gabor小波和形态学的高分辨率图像树冠提取方法

树冠信息的遥感提取能有效辅助森林参数反演、林分长势监测、树种识别等森林调查活动。随着遥感信息自动化提取的需求不断加强,本文基于高空间分辨率遥感数据,以滁州市皇甫山林场为研究区域,设计了一种结合Gabor小波和形态学的树冠提取方法。该方法首先采用Gabor小波提取出纹理特征,其次结合K-means聚类分析方法,对PCA降维后的纹理特征向量提取出阔叶林区,最后基于形态学理论降低影像噪声,并利用前景后景标记的分水岭方法进行单木树冠提取。经过与人工解译的树冠信息结果对比发现,在郁闭度较高的阔叶林区,该自动化方法提取树冠精度较高,分割准确率A_d为79.59%,F测度达到了79.00%能有效提供精确的单木树冠信息,为林业经济调查技术的发展具有一定的实践意义。     

Tree population dynamics of three altitudinal vegetation communities on Mount Cameroon (1989�C2004)

Changes in permanent sample plots in the lowland,submontane and montane forests on Mount Cameroon(4,095 m above sea level),an active volcano,are described for 15 years from 1989 to 2004.Throughout the study period,the stocking level of trees with a diameter at breast height(DBH) ≥ 10 cm in the three forests were lower than in pan-tropical stands suggesting a significant impact of volcanic and human-related activities on the vegetation communities on the mountain.Annual mortality rates in the submontane and montane forests were consistent with those reported for comparable altitudinal ranges in the Blue Mountains of Jamaica.The annual mortality rate was higher in the lowland forest than other lowland sites included.Divergence between recruitment and mortality rates was large suggesting that the three vegetation communities have not reached their climax.The seven-year difference in half-life of large trees(with a DBH ≥ 50 cm) in the submontane and montane forests suggests an altitudinal effect on turnover of larger trees that in turn contributes to the frequent small stature of high altitude forests.There was little evidence of an altitudinal effect on species turnover and growth rate.This finding supports generalizations about the zero effect of growth on the stature of high altitude trees.Understanding forest dynamics is crucially important in the management of tropical montane environmentsand in this instance particularly so given the recent creation of the Mount Cameroon National Park.     

Age-related changes of carbon accumulation and allocation in plants and soil of black locust forest on Loess Plateau in Ansai County,Shaanxi Province of China

The effects of reforestation on carbon(C) sequestration in China′s Loess Plateau ecosystem have attracted much research attention in recent years. Black locust trees(Robinia pseudoacacia L.) are valued for their important use in reforestation and water and soil conservation efforts. This forest type is widespread across the Loess Plateau, and must be an essential component of any planning for C sequestration efforts in this fragile ecological region. The long-term effects of stand age on C accumulation and allocation after reforestation remains uncertain. We examined an age-sequence of black locust forest(5, 9, 20, 30, 38, and 56 yr since planting) on the Loess Plateau to evaluate C accumulation and allocation in plants(trees, shrubs, herbages, and leaf litter) and soil(0–100 cm). Allometric equations were developed for estimating the biomass of tree components(leaf, branch, stem without bark, bark and root) with a destructive sampling method. Our results demonstrated that black locust forest ecosystem accumulated C constantly, from 31.42 Mg C/ ha(1 Mg = 10~6 g) at 5 yr to 79.44 Mg C/ha at 38 yr. At the ′old forest′ stage(38 to 56 yr), the amount of C in plant biomass significantly decreased(from 45.32 to 34.52 Mg C/ha) due to the high mortality of trees. However, old forest was able to accumulate C continuously in soil(from 33.66 to 41.00 Mg C/ha). The C in shrub biomass increased with stand age, while the C stock in the herbage layer and leaf litter was age-independent. Reforestation resulted in C re-allocation in the forest soil. The topsoil(0–20 cm) C stock increased constantly with stand age. However, C storage in sub-top soil, in the 20–30, 30–50, 50–100, and 20–100 cm layers, was age-independent. These results suggest that succession, as a temporal factor, plays a key role in C accumulation and re-allocation in black locust forests and also in regional C dynamics in vegetation.