宁夏及邻区石炭纪植物群

本文对宁夏及邻区石炭纪植物化石进行了系统研究，共鉴定植物化石２４属６２种，讨论了植物群的特征，划分了４个植物组合，即（１）Ｃａｒｄｉｏｐｔｅｒｉｄｉｕｍｓｐｅｔｓｂｅｒｇｅｎｓｅ－Ａｒｃｈａｅｏｃａｌａｍｉｔｅｓｓｃｒｏｂｉｃｕｌａｔｕｓ组合；（２）Ｍｅｓｏｃａｌｕｍｉｔｅｓｃｉｓｔｉｆｏｒｍｉｓ－Ｐｅｃｏｐｔｅｒｉｓａｓｐｅｒａ组合；（３）Ｃｏｎｃｈｏｐｈｙｌｌｕｍｒｉｃｈｔｈｏｆｅｎｉｉ－Ｐａｒｉｐｔｅｒｉｓｋａｉｐｉｎｇｉａｎａ组台；（４）Ｌｅｐｉｄｏｄｅｎｄｒｏｎｐｏｓｔｈｕｍｉｉ－Ｎｅｕｒｏｐｔｅｒｉｓｏｖａｔａ组合．这４个植物组合的地质时代大致与西欧的杜内期－维宪期、纳缪尔期、维斯发期和斯蒂芬期相当，其代表岩组分别为前黑山组－臭牛沟组、靖远组、羊虎沟组和太原组．     

巴彦浩特盆地东部上石炭统羊虎沟组孢粉组合

钻井资料证实，内蒙巴彦浩特盆地东部保存有较完整的石炭系，其中上石炭统羊虎沟组厚达４５９．６ｍ。经７２块岩样分析鉴定，共发现６２属１１９种孢粉，含６个新种。可概括为上下两个孢粉组合，上组合称Ｄｅｎｓｏｓｐｏｒｉｔｅｓｂｅｌｕｌｕｓ－Ｃｒａｓｓｉｓｐｏｒａｌａｔｉｇｒａｎｉｆｅｒ－Ｌｙｃｏｓｐｏｒａｇｒａｎｕｌａｔａ（简称ＢＬＧ）组合，下组合称Ｃｒａｓｉｓｐｏｒａｌａｔｉｇｒａｎｉｆｅｒ－Ｃａｍｐｔｏｒｉｌｅｔｅｓｒｅｔｉｃｕｌｏｆｏｒｍｉｓ－Ｖｅｓｔｉｓｐｏｒａｃｏｓｔａｔａ（简称ＬＲＣ）组合。地质时限为晚石炭世中期羊虎沟组（维斯法期Ａ—Ｃ）。     

新疆乌恰县莎里塔什组生物地层新知

对新疆乌恰县塔塔村下侏罗统莎里塔什组孢粉植物群的研究，共发现孢粉２２属２６种，植物化石１２属１５种。根据这些孢粉化石建立了一个孢粉组合，命名为Ｃｙａｔｈｉｄｉｔｅｓ－Ｃｌａｓｓｏｐｏｌｌｉｓ组合，再根据侏罗纪的标志植物锥叶蕨（Ｃｏｎｉｏｐｔｅｒｉｓ）的存在，将莎里塔什组的时代确定为早侏罗世，为乌恰地区早侏罗世生物地层提供了可靠依据     

青海北部早三叠世下环仓组孢粉组合

青海北部下环仓组孢粉组合,共计孢子花粉４３属７３种,其中包括１个新种,６个相似种、疑源类１１属。组合以Ｐｕｎｃｔａｔｉｓｐｏｒｔｔｅｓ－Ｌｕｎｄｂｌａｄｉｓｐｏｒａ－Ｃｙｃａｄｏｐｉｔｅｓ－Ｍｉｃｒｈｙｓｔｒｉｄｉｕｍ较多为特征,地质时代为旱三叠世期。     

内蒙巴彦浩特盆地东部上石炭统靖远组孢子花粉

巴彦浩特盆地的上石炭统靖远组与羊虎沟组为连续沉积,厚度４７７．５ｍ。经２９块岩样分析鉴定,发现孢子花粉５１属９５种（含２新种）和３粒不能鉴定的分子。根据孢粉的纵向演化,可分为上下两个组合带,下组合：Ｔｒｉｐａｒｔｉｔｅｓｖｅｔｕｓｔｕｓ－Ｔｒｉｑｕｉｔｒｉｔｅｓｔｒｉｂｕｌａｔｕｓ组合（简称ＶＴ带）,分布于巴参２井３７７９～３５７０ｍ井段,该孢子带具环孢类含量较低,未发现单缝孢Ｌａｅｖｉｇａｔｏｓｐｏｒｉｔｅｓ和Ｐｕｎｃｔａｔｏｓｐｏｒｉｔｅｓ以及不能归类的Ｔｒｉｎｉｄｕｌｕｓｄｉａｍｐｈｉｄｉｏｓ;上组合：Ｓｉｎｕｓｐｏｒｅｓｓｉｎｕａｔｕｓ—Ｃｒａｓｉｓ－ｐｏｒａｋｏｓａｎｋｅｉ组合（简称ＳＫ带）,分布于同一井的３５７０～３３０１．５ｍ井段,孢粉丰度较下组合高得多,具环孢子含量一般在５０％以上,花粉含量增高。地质时代属晚石炭世早期（纳缪尔Ｂ—Ｃ期）。     

四川省若尔盖县巴西—求吉地区晚三叠世遗迹化石及遗迹相

四川若尔盖巴西－求吉地区晚三叠世首次发现了丰富的遗迹化石，经鉴定有２２个遗迹属、３４个遗迹种。根据不同遗迹属在不同层位中的相对丰度，建立了４个遗迹组合：（１）Ｈｅｌｍｉｎｔｈｏｐｓｉｓ－Ｐｈｙｃｏｓｐｈｏｎ组合；（２）Ｄｅｎｄｒｏｔｉｃｈｎｉｕｍ－Ｎｅｏｎｅｒｅｉｔｅｓ组合；（３）Ｐａｌｅｏｄｉｃｔｙｏｎ组合；（４）Ｔａｅｎｉｄｉｕｍ－Ｃｏｓｍｏｒｈａｐｈｅ组合。上述遗迹组合大致相当于Ｓｅｉｌａｃ     

新疆准噶尔盆地东缘下石炭统滴水泉组孢粉组合

新疆准噶尔盆地东缘克拉美丽地区双井子剖面滴水泉组孢粉组合中有孢粉化石２９属３０种，具环孢占优势，含量为５７．９％，其中Ｃｒａｓｓｉｓｐｏｒａ（２３．６％）、Ｓｔｅｎｏｚｏｎｏｔｒｉｌｅｔｅｓ（１４．３％）的含量最高；无环孢居次，含量为３４．５％，以Ｐｕｎｃｔａｔｉｓｐｏｒｉｔｅｓ（１２％）和Ｅｅｉｏｔｒｉｌｅｔｅｓ（７．１％）为主。地质时代为早石炭世晚期，可能属韦宪期早期，大致相当于甘肃靖远臭牛沟组下段Ⅰ孢子组合带和西欧的Ｐｕ孢子组合带。当时为亚热带温暖湿润气候。     

新疆三塘湖盆地老爷庙组古植物特征及其时代

老爷庙组的命名者认为这套地层属于泥盆—石炭系过渡层，在地质图上标为早石炭世。新材料表明，植物大化石Ｌｅｐｔｏｐｈｌｏｅｕｍｒｈｏｍｂｉｃｕｍ－Ａｒｃｈａｅｏｐｔｅｒｉｓｆｉｓｉｌｉｓ－Ｃｙｃｌｏｓｔｉｇｍａｓｐ．组合的时代应属晚泥盆世晚期，孢子化石Ｒｅｔｕｓｏｔｒｉｌｅｔｅｓｓｉｍｐｌｅｘ－Ａｐｉｃｕｌｉｔｕｓｉｓｐｏｒａｈｕｎａｎｅｎｓｉｓ（ＳＨ）组合显示的时代也为晚泥盆世晚期，从而纠正了这一地层属于泥盆—石炭系过渡层的观点。     

新疆准噶尔盆地东缘下石灰统滴水泉组孢粉组合

新疆准噶尔盆地东缘克拉美丽地区双井子剖面滴水泉组孢粉组合中有孢粉化石２９属３０种，具环孢占优势，含量为５７．９％，其中Ｃｒａｓｓｉｓｐｏｒａ（２３．６％）、Ｓｔｅｎｏｚｏｎｏｔｒｉｌｅｔｅｓ（１４．３％）的含量最高；无环孢居次，含量为３４．５％，以Ｐｕｎｃｔａｔｉｓｐｏｒｉｔｅｓ（１２％）和Ｌｅｉｏｔｒｉｌｅｔｅｓ（７．１％）为主，地质时代为早石炭世晚期，可能属韦宪期早期，大致相当于甘肃靖远臭牛沟     

青海北祁连区西大沟组孢粉组合

青海北祁连区西大沟组孢粉组合含有８３属１１０种（类型）,包括２新种,其中蕨类植物孢子为３３．６％,裸子植物花粉为５０．５％,疑源类为１５．８％。本组合以Ａｐｉｃｕｌａｔｉｓｐｏｒｉｓ－Ｃｏｒｄａｉｔｉｎａ－Ｓｃｈｉｚｏｓｐｏｒｉｓ较多为特征地质时代为早三叠世。     

松辽盆地深层侏罗系-下白垩统生物组合和沉积环境

本文描述了松辽盆地探井揭露的深层地层序列及其所产的5个牛物门类化石组合特征.9个孢粉组合是Monosulcites-Cyathidites组合、Pinaceae-Cyathidites-Osmundaeidites组合、Classopollis-Piceites组合、Lophotriletes-Cicatricosisporites组合、Paleoconiferus-Lygodiumsporites组合、Cyathidites-Leiotriletes-Clavatipollenites组合、Cicatricosisporites-Leiotriletes-Polyporopollenites组合、Leiotriletes-Schizaeoisporites-Classopollis组合和Trilobosporites-Cyathidites-Tricolpollenites组合.3个藻类组合是Vesperopsis-Australisphaera组合、Balmula granorugosa组合和Vesperopsis zhaodongensis组合.2个介形类组合是Cypridea unicostata-Limnocypridesadscondida组合和Mongolocypris limpida-Paracandona planiuscula组合.4个植物组合是Elatocladus submanchurica-Ginkgoites orientalis组合、Acanthopteris gothani-Pterophyllum组合、Sphenopteris johnstrupii-Coniopteris nympharum组合和Platanus septentrionalis-Protophyllum undulaturn组合.3个大孢子组合是Minerisporites sp.组合、Ricinospora leavigata-Galamospora sp.组合和Arcellites组合.还介绍了叶肢介、轮藻、双壳类和鱼类化石的产出情况.结合同位素资料认为：大庆群时代为中侏罗世,火石岭组可能为贝里阿斯期,沙河子组为凡兰吟期-欧特里夫期,营城组为欧特里夫期-巴列姆期,登娄库组为巴列姆期-阿普第期早期,泉头组一-二段为阿普第-阿尔卑期,并对各组沉积环境进行了初步分析.     

谈女星介(Cypridea)在我国出现的最低层位

Cypridea(女星介 )分布在世界许多国家和地区。它的最早出现时代 ,除欧洲为中侏罗世外 ,都见于晚侏罗世。在我国Cypridea分布广泛 ,已在辽、吉、黑、京、冀、晋、蒙、沪、鲁、苏、皖、浙、赣、闽、豫、鄂、湘、粤、桂、琼、滇、川、陕、甘、宁、青、新等 2 7个省、市、自治区以及渤海、黄海海域发现。关于Cypridea在我国出现的最低层位有着不同的意见 ,或上侏罗统或下白垩统。究其原因 ,或是对生物群认识上的分歧 ,或是对含化石地层划分上的异议。通过对四川、湖北、冀北、辽西、甘肃河西走廊、青海、陕甘宁盆地、新疆及东南沿海Cypridea出现最低层位的分析 ,将它们中女星介出现的相同、相似的属种与英国Purbeck层CinderBedsMember上下层位的带化石对比 ,确定了我国晚侏罗世及晚侏罗—早白垩世临界时的生物群 ,从而确认Cypridea在我国出现的最低层位为上侏罗统。     

鲁西南莱芜、蒙阴、平邑盆地早第三纪地层的划分和对比

鲁西南地区分布着近东西向大致平行排列的早第三纪盆地群。在蒙阴、莱芜盆地早第三纪沉积称为官庄组，在平邑盆地称为卞桥组。依据沉积物特征和沉积旋回都能分为下段、中段和上段。通过对哺乳动物、介形类、轮藻、腹足类和孢粉等化石的研究可确定中段的沉积时代为早始新世，通过对轮藻、介形类和哺乳类动物的研究能确定下段和上段分别为始新世早期和中期。在卞桥组中段和官庄组中段发现介形类Ｃｙｐｒｉｄｅａ属，该属起源于晚侏罗世或早白垩世，在早白垩世繁盛，晚白垩世逐渐衰落，灭绝于晚古新世或早始新世，在早始新世沉积中的发现，是迄今在我国发现的最高层位。     

Paleoproterozoic arc and ophiolitic rocks on the northwest-margin of the Trans-Hudson Orogen,Saskatchewan, Canada: their contribution to a revised tectonic framework for the orogen

A new lithotectonic framework for the northwestern Reindeer Zone of the Trans-Hudson Orogen divides rocks into five northwest- to north-dipping volcano-sedimentary assemblages: (1) at the structural base, the 1.92–1.87 Ga largely sedimentary Levesque Bay Assemblage (partly equivalent to former ‘MacLean Lake gneisses’), which lies within the confines of the Kisseynew Domain and is tectonically imbricated with metasedimentary rocks of the <1.85 Ga McLennan and Burntwood groups; (2) the turbiditic Duck Lake Assemblage, also located along the northern edge of the Kisseynew Domain; it contains detrital zircons ranging in age between 1.92 and 1.87 Ga; (3) the ?1.92 Ga mafic–ultramafic volcano-plutonic Lawrence Point Assemblage of the La Ronge Domain; (4) the ≥1.88 Ga felsic to intermediate volcano-plutonic Reed Lake Assemblage of the La Ronge Domain; and (5) the turbiditic Milton Island Assemblage of the Rottenstone Domain, which contains detrital zircons ranging in age between 2.83 and 1.86 Ga. The assemblages are intruded by a variety of 1.91–1.78 Ga mafic to felsic plutons.The Lawrence Point Assemblage is interpreted as a dismembered supra-subduction zone ophiolite. High-MgO refractory harzburgite (‘Group 1’ ultramafic rocks), at the structural base of the assemblage, is geochemically identical to the upper mantle section of selected supra-subduction zone ophiolites and mantle tectonites. Chromite and olivine compositions of the ‘Group 1’ ultramafic rocks are also comparable to those of ophiolitic harzburgite and mantle tectonite. Mafic metavolcanic rocks of the assemblage are classified as subalkaline tholeiitic basalts. Their trace element patterns and Hf, Ta, Th, Y, Nb, and La element ratios resemble those of modern back-arc basin basalts. The Reed Lake Assemblage represents a subduction-generated arc complex that was built on top of the Lawrence Point Assemblage; its mafic metavolcanic rocks are subalkaline basalts, with calc-alkaline trends, and elevated Th and Ce concentrations and negative Nb anomalies. Feldspar porphyry dykes intruding the Lawrence Point and Duck Lake assemblages constrain timing of Lawrence Point ophiolite emplacement onto the Duck Lake Assemblage to 1.86–1.84 Ga. The trace element geochemistry of the dykes suggests continued arc volcanism after ophiolite emplacement. Mafic metavolcanic rocks of the Levesque Bay Assemblage are geochemically similar to those of the Lawrence Point Assemblage. Other ultramafic rocks (peridotite to pyroxenite) are abundant in the Lawrence Point Assemblage, but have similar geochemistry to small ultramafic bodies intruding the Reed Lake, Duck Lake and Levesque Bay Assemblages. They represent a separate, later phase (?1.86 Ga) of ultramafic plutonism, which post-dates ophiolite emplacement.Timing of Lawrence Point ophiolite emplacement (between 1.86 and 1.84 Ga) and geochemistry of later felsic and mafic/ultramafic volcanism suggest that the Lawrence Point ophiolite and overlying Reed Lake arc assemblage were not accreted to the Hearne Craton prior to 1.86 Ga, but were first accreted to the Flin Flon–Glennie Complex after 1.86 Ga.     

Biostratigraphy and facies analysis of the Upper Cretaceous–Danian? platform carbonate succession in the Kuyucak area,western Central Taurides,S Turkey

The Upper Cretaceous succession outcropping in the Anamas–Akseki Autochton, consists of approximately 500 m thick purely platform carbonate sediments. It begins with Cenomanian limestones intercalated with limestone breccias (Unit-1) containing mainly Pseudorhapydionina dubia, Pseudonummoloculina heimi, Spiroloculina cretacea (Assemblage I) and unconformably overlies the Lower Cretaceous (Barremian–Aptian) limestones with Vercorsella laurentii, Praechrysalidina infracretacea and Salpingoporella hasi. The Cenomanian limestones include foraminiferal packstone–wackestone, peloidal packstone–wackestone and mudstone microfacies deposited in restricted platform conditions. The Cenomanian succession is truncated by an unconformity characterised by locale bauxite deposits. Immediately above the unconformable surface, dolomitic limestones and rudistid limestones (Unit-2) are assigned to the upper Campanian based on the benthic foraminiferal assemblage (Assemblage II) comprising mainly Murciella gr. cuvillieri, Pseudocyclammina sphaeroidea, Accordiella conica, Scandonea samnitica and Fleuryana adriatica (smaller-sized populations). The upper Campanian limestones composed of dominantly foraminiferal-microbial packstone–wackestone microfacies deposited in shallow water environment with low energy, restricted circulation. The following limestones of the Unit-2 is characterised by sporadic intercalation of “open shelf” Orbitoides, Omphalocyclus, Siderolites assemblage (Assemblage III), assigned to the Maastrichtian, in addition to pre-existing “restricted platform” species. In the upper part of this biozone, the Rhapydionina liburnica/Fleuryana adriatica concurrent range subzone (Assemblage IIIb) is distinguished by the presence of Valvulina aff. triangularis, Loftusia minor as well as the nominal species. The Maastrichtian limestones with sporadically open marine influence consist of bioclastic (rudist-bearing) packstone–floatstone, foraminiferal packstone–wackestone with rudist fragments and peloidal/intraclastic packstone–wackestone microfacies deposited in shallow subtidal–subtidal (lagoonal) environments. The Upper Cretaceous succession passes upwardly into 70 m thick limestones and clayey limestones (Unit-3) which do not contain rudists and pre-existing foraminiferal assemblage with one exception Valvulina aff. triangularis. Variable amounts of ostracoda, discorbids, miliolids, dasycladacean algae and Stomatorbina sp. (Assemblage IV) occur into mud-rich microfacies suggesting restricted conditions with low water energy. A probable Danian age is proposed for the Unit-3 based on the occurrence of Valvulina aff. triangularis and Stomatorbina sp. which were previously recorded from Danian of peri-Tethyan platforms.     

Jurassic-Cretaceous Boundary Strata of the Somanakamura Group in NE Japan and their Correlation with Coeval Terrestrial Deposits in China

Defining the Jurassic-Cretaceous boundary is a controversy in stratigraphic study of the world. It has been widely accepted that this boundary can be defined at the bottom of Berriasian in Tethys, with the appearance of the ammonite Berriasella jacobi dating to ca. 145 Ma. However, it is difficult for the widespread terrestrial deposits in China to correlate with the international standard of marine facies. The Somanakamura Group in Japan is represented by a succession of marine-continental transitional strata. It provides a bridge of marine and nonmarine stratigraphic correlation. The ammonite and radiolarian fossils preserved in this group suggest an age from Bajocian to early Valanginian. The J-K boundary was defined in or atop the Tomizawa Formation of the group according to the ammonite data. The present authors study the fossil spores and pollen newly found from the Tomizawa and Koyamada formations. Three assemblages have been recognized. They are Assemblage 1 (Cyathidites-Classopollis) from the upper part of the Tomizawa Formation, Assemblage 2 (Cyathidites-Jiaohepollis) from the lower part of the Koyamada Formation, and Assemblage 3 (Cyathidites-Spheripollenites-Ephedripites) from the middle to upper part of the Koyamada Formation. With the reference of ammonite evidence, the J-K boundary can be defined between Assemblage 1 and Assemblage 2. This palynological J-K boundary can be correlated with that of terrestrial sequence in China. However, local biostratigraphy imply that the continental J-K boundary in China is of 135 or 137 Ma age. It has a considerable discrepancy from the marine standard. Biogeographically, the distribution pattern of spores and pollen in southern China is in accordance with that in the Somanakamura Group, which parallels the Tuchengzi Formation in northeastern China. By the palynological correlation between the Somanakamura Group and the strata in southern China, and then with the sequence in northeastern China, it is suggested that the continental J-K boundary is located in the Tuchengzi Formation.     

冀北滦平陆相侏罗—白垩系生物地层界线及候选层型研究进展

滦平盆地张家沟侏罗—白垩系界线上下为连续沉积的滨浅湖—半深湖序列,介形类和叶肢介类化石丰富,是中国陆相J\K界线理想的层型候选剖面。通过晚侏罗世和早白垩世介形类、叶肢介生物组合及沉积学与成因地层学的综合研究,认为陆相侏罗—白垩系生物地层界线以介形类大个体滦平介Luanpingella的消失、女星介Cypridea的出现,叶肢介Nestoria、Pseudograpta及Keratestheria等的消失以及Eosestheria的始现,扇三角洲平原或前缘亚相或等同亚、微相带的结束为标志。陆相界线层型研究和定义异于海相盆地,陆相环境生物化石的自然缺欠可以通过地层格架建立及沉积亚微相序列高精度地划分、对比予以弥补,并应纳入陆相界线层型的定义中。     

东海沉积地层矿物研究

本文据东海五口探井的559个碎屑矿物样资料,用主要重矿物的含量及其组合特征编绘了含量变化图,试图进行分层比较。位于西湖凹陷的平湖一井和玉泉1井重矿组合特征可比性较好;龙井2井三潭组以下与上两井缺少可比特点;灵峰1井除东海群外,其余地层与上列3井相比,差异是大的。     

Duration of the first biozone in the Siberian hypostratotype of the Vendian

The first biozone (Anabarites trisulcatus Assemblage Zone) in the Siberian hypostratotype of the Vendian (northwestern slope of the Olenek Uplift) is represented by the Turkut Formation of the Khorbusuonka Group and most of the Syhargalakh Formation of the Kessyusa Group. The lower part of the Kessyusa Group in some of the sections includes stratiform breccia coeval with the middle part of the Syhargalakh Formation. The breccia is shown to be the alteration product of tuff breccia, which is widely distributed in the region and occurs as diatremes. A U-Pb zircon date of 543.9 ± 0.24 Ma for tuff breccia provides the best constraint on the age of the boundary between the Anabarites trisulcatus and Purella antiqua Assemblage Zones. The first appearance of small skeletal fossils Cambrotubulus decurvatus (which define the base of the Anabarites trisulcatus Assemblage Zone) is 1.4 m above the lower boundary of the Turkut Formation. Ichnofabric in the underlying Khatyspyt Formation is globally distributed in the strata 553-551 million years old, always predating the first appearance of small skeletal fossils of the Anabarites trisulcatus Assemblage Zone. The base of the Anabarites trisulcatus Assemblage Zone is therefore younger than 553-551 Ma, whereas the duration of the assemblage zone does not exceed six million years.     

日本早白垩世非海相软体动物群组合

日本具有4个标志早白垩世的半咸水和淡水软体动物群组合:欧特里沃期—早巴列姆期的立川和菖蒲动物群组合,晚巴列姆期的濑林动物群组合和早阿普特期动物群组合。纯非海相下白垩统大量分布于日本东北部(本州岛北部)的部分地区,以及日本中部、西南部的内带。在日本东北部的部分地区以及日本西南部的外带(四国岛与本州岛中部),从巴列姆阶下部往上有一些海相地层夹在非海相层中,这为研究非海相动物群提供了准确的年代依据。生物组合之间在种一级水平上的区别,主要由环境的差异而引起。1)立川动物群组合(欧特里沃期—早巴列姆期),含有半咸水Hayamina naumanni组合和淡水Megasphaerioides okurodaniensis组合;手取群的Okurodani、Kuwajima和Izuki三个组内的淡水软体动物可与本动物组合相对比;2)菖蒲动物群组合(欧特里沃期—早巴列姆期),含有Eomiodon nipponicus组合;3)濑林动物群组合(晚巴列姆期),含有Costocyrena radiatostriata组合,可与手取群的Kitadani组和日本东北部关门群的Sengoku组以及Monomiyama组的淡水软体动物组合相对比;4)日比原动物群组合(早阿普特期),含有Costocyrena minor组合,关门群Wakamiya组的淡水软体动物可与本动物组合相对比。