二叠纪—三叠纪之交凝块石（微生物岩）的沉积成岩过程——以华南广西百色地区作登剖面为例

二叠纪末大灭绝之后,在我国华南地区广泛发育了一套钙质微生物岩,这套微生物岩以凝块石为主,代表了大灭绝事件后特殊环境下的生物沉积建造。早期对凝块石的分布以及沉积特征有详细研究。通过镜下显微观察,阴极发光照相,背散射成像以及探针元素分析,对凝块石的显微特征进行了详细研究。通过对凝块石中化石的分布,矿物组成以及元素分布规律观察,对比现代凝块石沉积成岩过程总结了二叠纪末凝块石的沉积成岩过程,即早期生长阶段,沉积阶段,早期成岩阶段和后期成岩阶段。清晰的凝块石的沉积成岩过程为解析该时期微生物沉积建造打下基础。     

扬子地台与华南南盘江盆地大贵州滩三叠系沉积演化史

扬子地台是一个横跨华南地块的以浅海沉积为主的大型碳酸盐岩台地,南盘江盆地是发育在扬子地台碳酸盐岩台地背景之上的一个沉积盆地,从晚元古代到晚三叠世长期海相沉积演化历史中,扬子地台一南盘江盆地体系经历了多次重要的构造演化阶段。扬子地台从晚元古代到早三叠世末期一直保持为一个稳定的碳酸盐岩台地,在中三叠世末期扬子地块整体抬升,海平面下降,形成了遍及扬子主体的拉丁期大海退,从而使扬子地块大部分地区抬升为陆。南盘江盆地位于华南地块南缘,从晚元古代到晚三叠世沉积了一套厚度巨大的海相碳酸盐岩,晚三叠世发育了一套硅质碎屑的浊流沉积,区域沉积也由此转化为河流相沉积。二叠纪和三叠纪碳酸盐岩地层记录了碳酸盐岩台地长期演化历史及其特征多样的沉积建造和沉积环境,而硅质碎屑流和构造沉降速率的变化反映了盆地在三叠纪期间经历的聚合构造和前陆盆地发展过程。在三叠纪时期扬子地台沿西南一北东方向从云南围绕南盘江盆地向贵州延伸,在南盘江盆地中发育了几个孤立的碳酸盐岩台地,包括位于贵州南部和广西境内的大贵州滩和崇左平果台地。南盘江盆地在晚二叠世发生过一次区域性的海侵事件,早三叠世时期扬子地台和几个孤立台地为由鲕粒边滩组成的低角度斜坡,中三叠世（安尼期）变为由Tubiphytes边礁组成的陡倾斜坡。盆地范围内斜坡变陡激发了Tubiphytes礁和其它的生物体发育,而且它们组成了稳定碳酸盐岩台地的边缘。位于扬子地台西部地区的关林和贞丰一带与最北部的孤立台地（大贵州滩）在安尼期发育了陡倾的边礁。在拉丁期,扬子地台在关林一带进积并与盆地碎屑沉积互层穿插沉积,而位于贞丰的台地边缘出现了由断层控制的地貌特征。与此同时,扬子地     

早三叠世生物复苏期的特殊沉积——"错时相"沉积

经历了对二叠纪末大灭绝及相关地质灾变事件的多年热点研究后,近年来科学家们将注意力转移到灭绝后的事件效应上,即生态系和沉积体系状况。但紧随二叠纪末灭绝事件之后的早三叠世生态系以分异度极低的广适性分子和机会分子为主,这就突显沉积记录的重要,也使得下三叠统地层中的特殊沉积及相关构造——“错时相”沉积,如扁平砾石砾岩、蠕虫状灰岩、潮下皱纹构造、微生物岩、海底碳酸盐胶结岩扇、薄层灰岩和条带灰岩等,成为研究灭绝—残存—复苏领域的学科前沿。作为地质历史环境一次大跃变后的直接产物,“错时相”沉积紧接生物大灭绝后出现,并随中生代海洋生态系的重建而退出正常浅海环境,这种耦合关系表明沉积体系、生态系、生物灭绝与复苏、异常环境之间存在必然的联系。对于化石保存单调稀少的下三叠统地层,“错时相”沉积的研究,为探索二叠纪末生物灭绝与复苏提供了宝贵的材料和全新的视角。     

湖北崇阳二叠纪—三叠纪之交钙质微生物岩的时代及基本特征

湖北崇阳钙质微生物岩产在二叠纪末礁相绝灭界线之上, 绝灭界线之下即为晚二叠世长兴期的海绵礁灰岩、棘屑灰岩及藻有孔虫生物碎屑灰岩.微生物岩基本上由中粗晶-微晶方解石组成, 主要沉积构造有“花斑状构造”及藻叠层构造.目前微生物岩中发现的化石主要有球状菌藻类化石、介形虫、微型腹足、双壳、鱼牙及分类位置尚无法确定的棒状微型化石.微生物岩中已发现有牙形石化石: Hindeodusparvus, H.typicalis和H.latidentatus, 从牙形石的类型及产出层位分析, 崇阳钙质微生物岩的下部属于晚二叠世长兴期, 上部为早三叠世.崇阳微生物岩在沉积特征、生物组成及牙形石带上均基本可与华南其他地区已发现的微生物岩进行对比.      

东秦岭陕西聂家沟剖面二叠纪—三叠纪之交沉积特征及其古环境意义*

东秦岭陕西镇安聂家沟地区二叠纪—三叠纪之交发育一套海相碳酸盐岩沉积序列,较完整地记录了二叠纪末生物大灭绝事件前后浅海碳酸盐岩台地相生物群落演替及环境变化等信息,是研究二叠纪末生物大灭绝事件前后生态环境变化与沉积响应的理想区域。通过碳酸盐岩微相分析方法,对陕西镇安聂家沟剖面二叠系—三叠系界线附近的碳酸盐岩生物组合和微相类型进行了详细分析,共识别出11个微相类型: 斑点状凝块石、叠层石灰岩、鲕粒—纹层状叠层石灰岩、有孔虫颗粒灰岩、藻—海百合泥粒灰岩、含有被包壳和被磨蚀骨屑颗粒的粒泥灰岩、鲕粒颗粒灰岩、集合颗粒灰岩、含鲕粒的粒泥灰岩、泥晶灰岩和泥岩。根据碳酸盐岩微相特征及沉积相标志,在二叠系—三叠系界限附近划分出台地边缘、开阔台地和局限台地3种沉积相,其反映了二叠纪—三叠纪之交频繁的沉积相带变更的特点。该剖面碳酸盐岩微相反映的古生态和生物群落演替特征与中国华南同时期其他剖面具有较好的一致性,即二叠纪末生物大灭绝事件之后,早三叠世正常浅水碳酸盐岩台地生物类型和丰度极低,仅含有少量的双壳类、海百合等,灭绝事件界线附近以微生物碳酸盐岩等特殊微生物沉积构造占主导,之后微生物岩等消失,又出现了以双壳类为首的生物碎屑石灰岩。研究结果为了解二叠纪末生物大灭绝事件中生物和环境的协同演化提供了新的材料和认识。     

东秦岭陕西聂家沟剖面二叠纪—三叠纪之交沉积特征及其古环境意义*

东秦岭陕西镇安聂家沟地区二叠纪—三叠纪之交发育一套海相碳酸盐岩沉积序列,较完整地记录了二叠纪末生物大灭绝事件前后浅海碳酸盐岩台地相生物群落演替及环境变化等信息,是研究二叠纪末生物大灭绝事件前后生态环境变化与沉积响应的理想区域。通过碳酸盐岩微相分析方法,对陕西镇安聂家沟剖面二叠系—三叠系界线附近的碳酸盐岩生物组合和微相类型进行了详细分析,共识别出11个微相类型: 斑点状凝块石、叠层石灰岩、鲕粒—纹层状叠层石灰岩、有孔虫颗粒灰岩、藻—海百合泥粒灰岩、含有被包壳和被磨蚀骨屑颗粒的粒泥灰岩、鲕粒颗粒灰岩、集合颗粒灰岩、含鲕粒的粒泥灰岩、泥晶灰岩和泥岩。根据碳酸盐岩微相特征及沉积相标志,在二叠系—三叠系界限附近划分出台地边缘、开阔台地和局限台地3种沉积相,其反映了二叠纪—三叠纪之交频繁的沉积相带变更的特点。该剖面碳酸盐岩微相反映的古生态和生物群落演替特征与中国华南同时期其他剖面具有较好的一致性,即二叠纪末生物大灭绝事件之后,早三叠世正常浅水碳酸盐岩台地生物类型和丰度极低,仅含有少量的双壳类、海百合等,灭绝事件界线附近以微生物碳酸盐岩等特殊微生物沉积构造占主导,之后微生物岩等消失,又出现了以双壳类为首的生物碎屑石灰岩。研究结果为了解二叠纪末生物大灭绝事件中生物和环境的协同演化提供了新的材料和认识。     

楚雄盆地会基关反转背斜的确立及其意义

楚雄盆地处在扬子地台西缘,发育在早—中三叠世古隆起上。晚三叠世早—中期伸展裂陷,沉积形成了近海泥质、灰质生油岩、含中基性火山碎屑复理石建造;晚三叠世晚期—早白垩世整体坳陷,四川盆地与楚雄盆地一度连为统一的巨型坳陷,沉积形成了内陆含煤磨拉石建     

上扬子地区下三叠统飞仙关组一段:大灭绝后从停滞海洋到动荡海洋的沉积记录

在P—T界线生物大灭绝事件以后的早三叠世早期,上扬子地区广泛沉积了低能环境纹层状微晶灰岩、灰质页岩或条带状灰岩沉积,代表了P—T事件之后早三叠世最早期上扬子地区所处的古特提斯海海洋循环的近乎停滞;该套低能环境静水沉积广泛见于江油、广元、旺苍、重庆等地剖面中。上覆于停滞海洋沉积之上的是角砾状灰岩、扁平砾石灰岩、丘状交错层理颗粒灰岩、沙纹层理粉砂岩等和风暴作用有关的动荡海水沉积;低能环境沉积与动荡海水沉积之间常见冲刷、侵蚀界面,这都反映了上扬子地区从停滞海洋到动荡海洋的地质记录。川西北地区角砾状灰岩分布面积大、成因特殊,而扁平砾石灰岩是早三叠世典型的与风暴作用有关的错时相灰岩。沉积岩石显示的从静水条件到与风暴有关的动荡水条件的环境变化,预示着早三叠世早期生物大灭绝以后不同寻常的古海洋和古气候变化。P—T生物大灭绝可能对早三叠世早期古气候和大气循环起到了显著作用,P—T事件后生物对整个地球系统的调节作用减弱,地球系统向极端情况发展的趋势将得不到有效制约,全球古环境与古气候可能因此变得极端恶劣。     

湖北松滋地区下奥陶统微生物岩

首次系统性地对湖北松滋地区下奥陶统微生物岩开展研究。根据微生物岩的生长方式,文中将微生物岩划分为原地生长型和非原地生长型两大类别,前者包括叠层石、凝块石,后者主要为核形石。根据几何形态特征,将研究区的叠层石分为层状、波状、柱状和丘状4种类型,新发现并命名了双锥柱状叠层石。将凝块石划分为斑状、网状和条带状3种类型,在豹斑状和网状凝块石中,发现了许多钻孔捕食软体动物Ecculiomphalus化石以及生物扰动的痕迹,文中认为这两种凝块石均属于生物扰动型凝块石。生物扰动型凝块石的发现不仅填补了国内、外奥陶纪该领域的部分空白,而且为凝块石成因的研究提供新的材料。在对各类微生物岩的沉积特征分析基础上,阐述了各类微生物岩的沉积环境,并总结其沉积环境分布模式。对微生物岩发育与中奥陶世后生动物大辐射之间的关系进行了探讨,根据早奥陶世晚期至中奥陶世初期后生动物的阶梯式和快速辐射与微生物岩突然减少的对应关系,结合研究区叠层石、凝块石中发现许多后生底栖钻孔捕食腹足类Ecculiomphalus化石等现象,认为微生物岩随时间推移而逐渐减少、衰退与后生动物丰度的快速增加有一定关系,指出食草动物不仅啃食了形成叠层石的菌藻类微生物,造成叠层石数量的不断减少,而且破坏了叠层石的生长状态,结果形成了研究区独具特征的生物扰动型凝块石。此外,作者认为,中奥陶世开始,海平面的快速上升也是研究区微生物岩减少直至消失的原因之一。     

鄂东南地区早三叠世中晚期沉积相和沉积古地理

鄂东南地区早三叠世大冶群为一套以碳酸岩为主的地层。在发育较全的阳新龙港新屋铺剖面可以划分出10个岩性段。本文着重探讨早三叠世中、晚期的沉积相特征。早三叠世鄂东南地区为下扬子海域的一部分,推测早三叠世早期奉区与赣西北为统一的开放海,可能与分隔华北和扬子大陆的秦岭-大别山海域相连。沉积物以深水盆地相和陆棚相泥质页岩和灰岩为主,夹风暴成因的砾屑灰岩和重力流成因的碎屑灰岩。早三叠世中期,大别山、九岭山古陆成为局限盆地,发育潮坪、渴湖、障壁丘和鲕粒滩;晚期因周期性的海平面变化导致形成局限浅水盆地、蒸发岩沉积和深水盆地泥岩和重力流沉积。这些变化与Haq等人(1988)提出的全球海平面变化曲线相吻合。     

The Permian–Triassic mass extinction: Ostracods (Crustacea) and microbialites

The end-Permian mass extinction (EPE), about 252 Myr ago, eradicated more than 90% of marine species. Following this event, microbial formations colonised the space left vacant after extinction of skeletonised metazoans. These post-extinction microbialites dominated shallow marine environments and were usually considered as devoid of associated fauna. Recently, several fossil groups were discovered together with these deposits and allow discussing the palaeoenvironmental conditions following the EPE. At the very base of the Triassic, abundant Ostracods (Crustacea) are systematically present, only in association with microbialites. Bacterial communities building the microbial mats should have served as an unlimited food supply. Photosynthetic cyanobacteria may also have locally provided oxygen to the supposedly anoxic environment: microbialites would have been refuges in the immediate aftermath of the EPE. Ostracods temporarily disappear together with microbialites during the Griesbachian.     

Current perspectives on the Permian–Triassic boundary and end-Permian mass extinction: Preface

The end-Permian mass extinction is now robustly dated at 252.6 ± 0.2 Ma (U–Pb) and the Permian–Triassic (P–T) GSSP level is dated by interpolation at 252.5 Ma. An isotopic geochronological timescale for the Late Permian–Early Triassic, based on recent accurate high-precision U–Pb single zircon dating of volcanic ashes, together with calibrated conodont zonation schemes, is presented. The duration of the Early Triassic (Induan + Olenekian stages) is estimated at only 5.5 million years. The duration of the Induan Stage (Griesbachian + Dienerian sub-stages) is estimated at ca. one million years and the early Olenekian (Smithian sub-stage) at 0.7 million years duration. Considering this timescale, the “delayed” recovery following the end-Permian mass extinction may not in fact have been particularly protracted, in the light of the severity of the extinction. Conodonts evolved rapidly in the first 1 million years following the mass extinction leading to recognition of high-resolution conodont zones. Continued episodic global environmental and climatic stress following the extinction is recognized by multiple carbon isotope excursions, further faunal turnover and peculiar sedimentary and biotic facies (e.g. microbialites). The end-Permian mass extinction is interpreted to be synchronous globally and between marine and non-marine environments. The nature of the double-phased Late Permian extinction (at the Guadalupian–Lopingian boundary and the P–T boundary), linked to large igneous provinces, suggests a primary role for superplume activity that involved geomagnetic polarity change and massive volcanism.     

湖北崇阳二叠纪—三叠纪之交生物灭绝和沉积微相演化

二叠纪—三叠纪之交,湖北崇阳地区处于浅水碳酸盐岩台地环境.二叠纪末的全球事件在该剖面的沉积微相和生物演化上均留下了清楚的记录.二叠纪末生物大灭绝之前,崇阳地区为典型的正常浅海台地环境,生物种类多样,数量丰富,主要生物化石有钙藻、有孔虫、腕足、棘皮类和海绵等.生物大灭绝之后,钙藻、(筳)类、棘皮类、海绵、绝大部分有孔虫开始消失,取而代之的是个体微小的腹足、介形虫和大量的蓝细菌化石.大灭绝界线之上,首先出现的是25 cm厚的纹层状的微生物岩,含较丰富的种类单调的有孔虫化石.之后逐渐相变为花斑状微生物岩和穹隆状微生物岩,厚度分别为6.4,2.3m.不同类型微生物岩在结构构造和生物组成上存在差别.微生物岩沉积结束之后,则相变为浅滩相鲕粒灰岩.共划分出3种沉积相,即开阔台地相、潮坪相和浅滩相.崇阳剖面的生物灭绝和沉积微相变化是二叠纪—三叠纪之交浅水台地环境生物与环境过程的典型代表,为认识二叠纪末浅海沉积相演化和全球事件提供了新材料.     

上扬子石炭-二叠纪海相碳酸盐的锶同位素演化与全球海平面变化

地质历史中海水的锶同位素组成是时间的函数,全球海平面变化是其最主要的控制因素,上扬子地区石炭-二叠纪海相碳酸盐的锶同位素演化曲线与海平面变化曲线有着很好的一致性。锶同位素演化曲线说明:1)早石炭世是一个海水逐渐加深的全球海平面上升时期,锶同位素最小值所显示的最大海泛面的年龄为 34 2Ma,位于杜内阶和韦宪阶的界线上 ;2 )晚石炭世是一个全球海平面下降时期 ;3)整个二叠纪都是全球海平面上升时期,晚二叠世的海平面上升不仅幅度大,而且海水在短时间内迅速加深 ;4)晚二叠世具有古生代海相碳酸盐的锶同位素最小值,显示晚二叠世末的全球淹没事件,最大海泛面的年龄为 2 5 0Ma,正好在二叠 /三叠纪界线附近 ;5 )二叠 /三叠纪之交的全球生物绝灭事件可能与二叠世末的全球淹没事件有关。     

Microbial deposits in the aftermath of the end‐Permian mass extinction: A diverging case from the Mineral Mountains (Utah,USA)

The Lower Triassic Mineral Mountains area (Utah, USA) preserves diversified Smithian and Spathian reefs and bioaccumulations that contain fenestral‐microbialites and various benthic and pelagic organisms. Ecological and environmental changes during the Early Triassic are commonly assumed to be associated with numerous perturbations (productivity changes, acidifica‐tion, redox changes, hypercapnia, eustatism and temperature changes) post‐dating the Permian–Triassic mass extinction. New data acquired in the Mineral Mountains sediments provide evidence to decipher the relationships between depositional environments and the growth and distribution of microbial structures. These data also help to understand better the controlling factors acting upon sedimentation and community turnovers through the Smithian–early Spathian. The studied section records a large‐scale depositional sequence during the Dienerian(?)–Spathian interval. During the transgression, depositional environments evolved from a coastal bay with continental deposits to intertidal fenestral–microbial limestones, shallow subtidal marine sponge–microbial reefs to deep subtidal mud‐dominated limestones. Storm‐induced deposits, microbialite–sponge reefs and shallow subtidal deposits indicate the regression. Three microbialite associations occur in ascending order: (i) a red beds microbialite association deposited in low‐energy hypersaline supratidal conditions where microbialites consist of microbial mats and poorly preserved microbially induced sedimentary structure; (ii) a Smithian microbialite association formed in moderate to high‐energy, tidal conditions where microbialites include stromatolites and associated carbonate grains (oncoids, ooids and peloids); and (iii) a Spathian microbialite association developed in low‐energy offshore conditions that is preserved as multiple decimetre thick isolated domes and coalescent domes. Data indicate that the morphologies of the three microbialite associations are controlled primarily by accommodation, hydrodynamics, bathymetry and grain supply. This study suggests that microbial constructions are controlled by changes between trapping and binding versus precipitation processes in variable hydrodynamic conditions. Due to the presence of numerous metazoans associated with microbialites throughout the Smithian increase in accommodation and Spathian decrease in accommodation, the commonly assumed anachronistic character of the Early Triassic microbialites and the traditional view of prolonged deleterious conditions during the Early Triassic time interval is questioned.     

Behavior of lophophorates during the end-Permian mass extinction and recovery

The end-Permian mass extinction devastated most marine communities and the recovery was a protracted event lasting several million years into the Early Triassic. Environmental and biological processes undoubtedly controlled patterns of recovery for marine invertebrates in the aftermath of the extinction, but are often difficult to single-out. The global diversity and distribution of marine lophophorates during the aftermath of the end-Permian mass extinction indicates that stenolaemate bryozoans, rhynchonelliform brachiopods, and lingulid brachiopods displayed distinct recovery patterns.Bryozoans were the most susceptible of the lophophorates, experiencing relatively high rates of extinction at the end of the Permian, and becoming restricted to the Boreal region during the Early Triassic. The recovery of bryozoans was also delayed until the Late Triassic and characterized by very low diversity and abundance. Following the final disappearance of Permian rhynchonelliform brachiopod survivors, Early Triassic rhynchonelliform brachiopod abundance remained suppressed despite a successful re-diversification and a global distribution, suggesting a decoupling between global taxonomic and ecological processes likely driven by lingering environmental stress.In contrast with bryozoans and rhynchonelliforms, lingulid brachiopods rebounded rapidly, colonizing shallow marine settings left vacant by the extinction. Lingulid dominance, characterized by low diversity but high numerical abundance, was short-lived and they were once again displaced back into marginal settings as environmental stress changed through the marine recovery. The presence in lingulid brachiopods of the respiratory pigment hemerythrin, known to increase the efficacy of oxygen storage and transport, when coupled with other morphological and physiological adaptations, may have given lingulids a survival advantage in environmentally stressed Early Triassic settings.     

Behavior of lophophorates during the end-Permian mass extinction and recovery

The end-Permian mass extinction devastated most marine communities and the recovery was a protracted event lasting several million years into the Early Triassic. Environmental and biological processes undoubtedly controlled patterns of recovery for marine invertebrates in the aftermath of the extinction, but are often difficult to single-out. The global diversity and distribution of marine lophophorates during the aftermath of the end-Permian mass extinction indicates that stenolaemate bryozoans, rhynchonelliform brachiopods, and lingulid brachiopods displayed distinct recovery patterns.Bryozoans were the most susceptible of the lophophorates, experiencing relatively high rates of extinction at the end of the Permian, and becoming restricted to the Boreal region during the Early Triassic. The recovery of bryozoans was also delayed until the Late Triassic and characterized by very low diversity and abundance. Following the final disappearance of Permian rhynchonelliform brachiopod survivors, Early Triassic rhynchonelliform brachiopod abundance remained suppressed despite a successful re-diversification and a global distribution, suggesting a decoupling between global taxonomic and ecological processes likely driven by lingering environmental stress.In contrast with bryozoans and rhynchonelliforms, lingulid brachiopods rebounded rapidly, colonizing shallow marine settings left vacant by the extinction. Lingulid dominance, characterized by low diversity but high numerical abundance, was short-lived and they were once again displaced back into marginal settings as environmental stress changed through the marine recovery. The presence in lingulid brachiopods of the respiratory pigment hemerythrin, known to increase the efficacy of oxygen storage and transport, when coupled with other morphological and physiological adaptations, may have given lingulids a survival advantage in environmentally stressed Early Triassic settings.