新疆博斯腾湖表层沉积物的孢粉分析

通过对新疆博斯腾湖的表层沉积物孢粉的分析, 探讨了干旱区大型湖泊中表层沉积物孢粉与周围植被的关系以及孢粉在湖泊中的分布和传播规律. 研究发现, 博斯腾湖表层沉积物中孢粉主要是来源于周围荒漠植被中的藜科、蒿属、麻黄等植物以及湖泊四周浅水地带生长的香蒲等水生植物. 藜科、蒿属、麻黄等花粉的传播过程主要受大风作用的影响, 而且百分比含量在整个湖盆中的分布比较稳定, A/C值能够反映区域目前的干旱状况; 香蒲等水生植物花粉的百分比含量主要受其母体植物分布的影响. 云杉花粉在河口区域的相对高百分比含量表明其河流传播特征; 云杉花粉在湖泊中的低百分比含量与整个流域的植被及其分布状况以及区域的气候条件有关; 云杉花粉具有超代表性. 近河口湖泊表层样品孢粉组合受河流和近岸植被的影响较大, 但距离超过2 km以后湖泊沉积花粉百分比含量较为稳定. 孢粉的质量浓度分布显示, 孢粉在湖盆中的分布有比较强的趋中性. 博斯腾湖中部沉积物孢粉序列能够很好记录区域的植被变化历史和环境演化特征.     

Modern pollen assemblages from surface lake sediments and their environmental implications on the southwestern Tibetan Plateau

The relationships amongst modern pollen assemblages, vegetation, climate and human activity are the basis for reconstructing palaeoenvironmental changes using pollen records. It is important to determine these relationships at regional scales due to the development of vegetation under different climatic conditions and human activities. In this paper, we report on an analysis of modern pollen assemblages of 31 surface lake samples from 31 lakes (one sample per lake) on the southwestern Tibetan Plateau where the knowledge of modern pollen and their relationships with vegetation, climate and human activities is insufficient. The region includes five vegetation zones: sub‐alpine shrub steppe, alpine steppe, alpine meadow and steppe ecotone, mountain desert and alpine desert. The lakes span a wide range of mean annual precipitation (50–500 mm) and mean annual temperature (?8 to 6 °C). Modern pollen assemblages from our samples mainly consist of herb taxa (Artemisia, Cyperaceae, Poaceae, Chenopodiaceae, etc.) and some tree taxa (Pinus, Fagaceae, Alnus, etc.). The results indicate that modern pollen assemblages are able to reflect the main vegetation distribution. Redundancy analysis for the main pollen types and environmental variables shows that precipitation is the leading factor that influences the pollen distribution in the study area with the first axis capturing 13.7% of the variance in the pollen data set. The Artemisia/Chenopodiaceae ratio is valid for separating the desert component (<2) from the steppe and other vegetation zones (>2) but is unable to distinguish moisture variations. The Artemisia/Cyperaceae ratio is able to identify meadows (<1) and steppes (>1) and can be used as a moisture index on the southwestern Tibetan Plateau. Our results show that an appropriate range is needed for a modern pollen data set in order to perform pollen‐based quantitative climate reconstructions in one region. It is essential to perform modern studies before using pollen ratios to reconstruct palaeovegetation and palaeoclimate at a regional scale.     

晋冀低山丘陵区人工扰动植被花粉组合特征

54个晋冀低山丘陵区人工和人工扰动植被表土花粉样品分析表明:花粉组合以草本植物花粉(74.5％)为主,乔木花粉含量低于20％。人类活动较强的农田以谷物禾本科、蒿属和藜科花粉为主,人类活动较弱的荒地以蒿属、藜科和杂草禾本科花粉为主。人类活动强度不同,指示种不同,孢粉浓度亦不同;农田指示种为谷物禾本科、葡萄科和胡桃科,孢粉浓度约4380粒/g;而蒿属、蓼科和蔷薇科则为荒地指示种,孢粉浓度约10983粒/g。农作物种植方式不同,谷物禾本科花粉含量不同,单一禾本科作物种植区谷物禾本科花粉含量(40.2％)高于多种作物混作区(18.8％)和荒地(3.8％)的含量。谷物禾本科含量的变化可以为利用化石花粉提取和检测人类活动信息提供依据,也可为利用地层中谷物禾本科花粉判断古代农业活动提供依据。     

Pollen assemblages and their environmental implications in the Qaidam Basin,NW China

The Qaidam Basin is one of the most sensitive areas to climate change in China, owing to its unique geographical position and ecological condition. In this study, 32 surface‐soil pollen samples were collected to reveal the relationship between modern pollen assemblages, vegetation and precipitation in the eastern region of the Qaidam Basin. The results show that Chenopodiaceae (3.8–87%, average 48%), Artemisia (1.7–64.2%, average 17.5%) and Ephedra (0–90%, average 16.3%) are the dominant pollen types in all samples, and that different pollen assemblages correspond to different vegetation types. DCA and CCA of major pollen types demonstrate that precipitation is an important factor in the control of the distribution of vegetation in the study area. The content and concentration of the three major pollen types (Artemisia, Chenopodiaceae and Ephedra) change with the mean annual precipitation, and the optimum mean annual precipitation for Ephedra, Chenopodiaceae and Artemisia is <80, 80–200 and >160 mm, respectively. Correlation analysis between the variation in grain size of the three major pollen types and the main environmental variables shows that the grain size of the three pollen types is positively correlated with precipitation in the Qaidam Basin. The results confirm that precipitation is the most important environmental factor in the Qaidam Basin, and that it has an important effect on pollen grain size in the study area.     

呼伦湖表层沉积物花粉空间分布特征及影响因素探讨

对呼伦湖29个表层沉积物花粉组合和环境影响因子进行主成分分析(PCA)和冗余分析(RDA),探讨呼伦湖表层沉积物花粉组合的空间分布特征及造成空间分异的现代过程机制.研究结果表明,呼伦湖表层沉积物花粉组合空间分布差异显著,藜科和松属等花粉在湖泊东侧近岸地带百分含量较高,蒿属与桦属花粉在湖泊中部和西部百分含量较高.呼伦湖东岸盐生非地带性植被、主导风向形成的漂流、与水深相关的二次悬移再沉积过程以及河流搬运作用是影响呼伦湖表层沉积物花粉空间分异的主要因素,其中湖泊东岸盐生非地带性植被造成了藜科花粉百分含量在近东岸地带偏高,主导风向形成的由西向东的漂流作用可能导致了松属花粉百分含量在近东岸地带偏高.     

Post-Glacial climatic change on Boothia Peninsula, Nunavut, Canada

A high temporal resolution pollen diagram from a lake in the mid-Arctic region of the Boothia Peninsula, Nunavut, Canada, documents the history of the regional vegetation and climate for the past 7200 yr. Major tundra pollen taxa in the core include Cyperaceae and Salix, with Cyperaceae comprising over 50% of the pollen in the early and late Holocene. Tree pollen, transported from far to the south, comprised a large percentage of the pollen sum, with Pinus accounting for 30% of the pollen in some levels of the core. Pollen percentages and concentrations of taxa typical of the mid-Arctic were highest in the mid-Holocene, corresponding to warm conditions. Decreasing pollen concentrations indicate cooling temperatures, with more rapid decreases occurring around 4200, 3800-3400, and 2500 cal yr BP. Pollen percentages of Salix, Cyperaceae, and Artemisia increased in the past 35 yr in response to global warming. Reconstructions of July temperature using the modern analog technique showed that the mid-Holocene (5800-2800 cal yr BP) was approximately 1 °C higher than during the past 1000 yr.     

Interpretation of floodplain pollen in alluvial sediments from an Arid Region

Pollen was collected from modern alluvium and from the atmosphere to document the nature and amount of paleoenvironmental information reflected by alluvial pollen chronologies. Results indicate that pollen in alluvium is a homogeneous mixture derived almost entirely from the floodplain itself. The few pollen grains derived from nonfloodplain plant communities and preserved in alluvial sediments are so well mixed that their frequencies no longer reflect the geographic distribution of the specific plant communities in which they originated. In contrast, the abundance of alluvial pollen grains, derived from the major floodplain taxa (Chenopodiineae, Ambrosia type), varies with summer and winter climate. This annual variation is preserved in alluvial pollen assemblages through a combination of processes within sedimentation basins involving discontinuous deposition events and mechanical pollen degradation. The high-frequency, wide-amplitude pollen variance in alluvial pollen assemblages contrasts with the low-frequency, narrow-amplitude pollen variance in sediments of lakes and ponds. The slight geographic variance in alluvial pollen assemblages, in contrast to the large variance in soil pollen, allows use of alluvial pollen to infer climate throughout the watershed in which pollen is sampled.     

天山南坡小尤尔都斯盆地表土孢粉初步研究

通过天山中部小尤尔都斯盆地南北方向采集的27个表土样品的孢粉分析, 探讨了孢粉和植被带的关系及盆地中美天鹅湖的孢粉来源问题. 结果表明: 孢粉组合主要成分是云杉属、 菊科、 禾本科、 蒿属、 藜科、 唐松草属、 莎草科、 水龙骨和其他蕨类孢子, 表土孢粉组合与各植被带之间有良好的对应关系.主成分分析表明, 研究区植被分布主要受温度因素控制. 上升气流和水流是影响针叶林带云杉属花粉传播的主要因素, 由山谷风从巩乃斯云杉林带来的云杉花粉受到美天鹅湖北侧小山阻挡, 沉降在湖泊以南的山坡上, 之后随降雨和雪山融水进入美天鹅湖. 大部分表土孢粉属种的空间分布与相应的植物分布一致, 而蒿属和藜科花粉百分含量在一定程度上代表了区域花粉雨的信号, 利用A/C比值重建古环境时须要谨慎.蒿、 藜花粉计数较少时, A/C比值波动较大, 计数值大于约280粒时, A/C比值趋于稳定.     

Ecological significance of common pollen ratios: A review

Pollen ratios have been commonly used to indicate landscape change and climate variation. However, the reliability of these indicators needs to be verified by studies on modern pollen process. Here, we synthesized the major pollen ratios used in previous studies and found that pollen ratios are valuable indicators for the change of vegetation types and climate, e.g., precipitation and moisture. Artemisia/Chenopodiaceae (A/C) ratio could increase from desert to steppe and positively correlate with mean annual precipitation (MAP). Artemisia/Cyperaceae (A/Cy) ratio could be used to identify cool meadow and warm steppe, and it is positively correlated with temperature of July (T_July) and negatively correlated with MAP. Arboreal pollen/nonarboreal pollen (AP/NAP) ratio can be used as a semi-quantitative indicator for landscape and regional precipitation changes. In spite of the significant climatic and environmental implications of the pollen ratios, they were also questioned in some studies under various circumstances and thus caution is needed when using them to indicate climate in different vegetation zones.     

Late Holocene vegetation and climate oscillations in the Qaidam Basin of the northeastern Tibetan Plateau

Pollen evidence from sediment cores at Hurleg and Toson lakes in the Qaidam Basin was obtained to examine vegetation and climatic change in the northeastern Qinghai-Tibetan Plateau. The chronologies were controlled by ²¹⁰Pb and ¹³⁷Cs analysis and AMS ¹⁴C dating. Pollen assemblages from both lakes are dominated by Chenopodiaceae (∼ 40%), Artemisia (∼ 30-35%) and Poaceae (∼ 20-25%), with continued occurrence but low abundance of Nitraria, Ephedra, and Cyperaceae. Artemisia/Chenopodiaceae (A/C) pollen ratios from two lakes show coherent large oscillations at centennial timescale during the last 1000 yr. A/C ratios were high around AD 1170, 1270, 1450, 1700 and 1920, suggesting that the vegetation was more “steppe-like” under a relatively moist climate than that during the intervening periods. Wet-dry climate shifts at the two lakes (2800 m asl) are in opposite phases to precipitation changes derived from tree-ring records in the surrounding mountains (> 3700 m asl) and to pollen and snow accumulation records from Dunde ice core (5300 m asl), showing that a dry climate in the basin corresponds with a wet interval in the mountains, especially around AD 1600. This contrasting pattern implies that topography might have played an important role in mediating moisture changes at regional scale in this topographically complex region.     

Extending the known distribution of the Younger Dryas Vedde Ash into northwestern Russia

The known distribution of wind‐blown Vedde Ash (ca. 10.3 ka BP) has been extended to the Karelian Isthmus in northwestern Russia. This has been possible as the result of a density separation technique that separates the rhyolitic Vedde Ash shards from the minerogenic host sediment. The Vedde Ash occurs in the middle of a pollen zone with high percentages of, for example, Artemisia and Chenopodiaceae, suggesting that the Younger Dryas (or GS‐I in the GRIP ice‐core event stratigraphy) was cold and dry throughout its duration. This is in agreement with sites in south Sweden where the Vedde Ash also occurs in the middle of a pollen zone dominated by Artemisia, Chenopodiaceae and Cyperaceae. Copyright © 2000 John Wiley & Sons, Ltd.     

Pollen taphonomy in a canyon stream

Surface soil samples from the forested Chuska Mountains to the arid steppe of the Chinle Valley, Northeastern Arizona, show close correlation between modern pollen rain and vegetation. In contrast, modern alluvium is dominated by Pinus pollen throughout the canyon; it reflects neither the surrounding floodplain nor plateau vegetation. Pollen in surface soils is deposited by wind; pollen grains in alluvium are deposited by a stream as sedimentary particles. Clay-size particles correlate significantly with Pinus, Quercus, and Populus pollen. These pollen types settle, as clay does, in slack water. Chenopodiaceae-Amaranthus, Artemisia, other Tubuliflorae, and indeterminate pollen types correlate with sand-size particles, and are deposited by more turbulent water. Fluctuating pollen frequencies in alluvial deposits are related to sedimentology and do not reflect the local or regional vegetation where the sediments were deposited. Alluvial pollen is unreliable for reconstruction of paleoenvironments.     

An 11 000-year record of vegetation and environment from Lago di Martignano,Latium, Italy

A pollen diagram from Lago di Martignano, a maar lake in central Italy, provides an 11000-year record of vegetation and environment change. The earliest pollen spectra are dominated by Artemisia and Gramineae, representing late glacial steppe vegetation typical of the Mediterranean region. Broad-leaved forests were established by ca. 11 000 yr BP. Although Quercus initially dominated their canopy, a wide range of other mesophyllous trees were also present. Pollen values for sclerophyllous tree and shrub taxa characteristic of Mediterranean woodlands and scrub are initially low (<10%). After ca. 7000 yr BP, however, they begin to increase and rise to a peak of >40% of total land pollen at ca. 6700 yr BP, with Olea europaea the single most abundant taxon. Human influence upon the vegetation only becomes significant somewhat after this peak, with progressive clearance of woodland and expansion of herbaceous communities. Castanea sativa and luglans regia pollen is recorded consistently from the beginning of the rise in pollen values for taxa characteristic of Mediterranean scrub communities. Pollen values for arable crops increase progressively after ca. 5500 yr BP, following the peak pollen values for taxa characteristic of Mediterranean scrub vegetation. Late glacial and Holocene climate changes have been complex in this region, with the present character of the climate developing only during the last millennium. Rates of change of pollen spectra peak during this period.     

Palynological evidence for early Holocene aridity in the southern Sierra Nevada, California

Sediments of Balsam Meadow have produced a 11,000-yr pollen record from the southern Sierra Nevada of California. The Balsam Meadow diagram is divided into three zones. (1) The Artemisia zone (11,000–7000 yr B.P.) is characterized by percentages of sagebrush (Artemisia) and other nonarboreal pollen higher than can be found in the modern local vegetation. Vegetation during this interval was probably similar to the modern vegetation on the east slope of the Sierra Nevada and the climate was drier than that of today. (2) Pinus pollen exceeded 80% from 7000 to 3000 yr B.P. in the Pinus zone. The climate was moister than during the Artemisia zone. (3) Fir (Abies, Cupressaceae, and oak (Quercus) percentages increased after 3000 yr B.P. in the Abies zone as the modern vegetation at the site developed and the present cool-moist climatic regime was established. Decreased fire frequency after 1200 yr B.P. is reflected in decreased abundance of macroscopic charcoal and increased concentration of Abies magnifica and Pinus murrayana needles.     

内蒙古岱海表层沉积物中孢粉的分布及来源

岱海位于半干旱区东部边缘及东亚夏季风的尾闾区，生态环境体系脆弱，气候和环境变化敏感，是研究环境变化的理想地区。65个湖泊表层及入湖河流冲积物样品的孢粉分析揭示了岱海表层孢粉沉积特征及孢粉来源。岱海表层孢粉主要来源于区域性植被的孢粉，其总体特征反映了区域植被的组成，但不同部位孢粉组合的差异反映了湖滨当地植被的变化。受沉积环境的影响，岱海湖区孢粉浓度总体上要低于入湖河流孢粉浓度，湖区中部孢粉浓度最高。岱海孢粉以河流携带的为主，同时受风力传播的影响。蒿、藜科、桦、虎榛子等花粉和蕨类孢子以河流搬运的为主，松、禾本科及莎草科花粉可能还受风力传播的影响。一些仅在个别样品中出现且百分比很低的花粉类型如豆科、十字花科等还有可能是昆虫、鸟类或人类活动携带而来。     

Paleoecological studies at Lake Patzcuaro on the west-central Mexican Plateau and at Chalco in the basin of Mexico

A 1520-cm sediment core from Lake Patzcuaro, Michoacan, Mexico, is 44,000 yr old at the base. All parts of the core have abundant pollen of Pinus (pine), Alnus (alder), and Quercus (oak) with frequent Abies (fir). The interval dated from 44,000 to 11,000 yr ago has a homogeneous flora characterized by abundant Juniperus (juniper) pollen and frequent Artemisia (sagebrush). It is believed to represent an appreciably drier and colder climate than at present. The Holocene at Lake Patzcuaro is characterized by a moderate increase in Pinus pollen and the loss of Juniperus pollen, as the modern type of climate succeeded. Alnus was abundant until about 5000 yr ago; its abrupt decrease with the first appearance of herbaceous weed pollen may reflect the cutting of lake-shore and stream-course alder communities for agricultural purposes, or it may simply reflect a drying tendency in the climate. Pollen of Zea (corn) appears at Lake Patzcuaro along with low peaks of chenopod and grass pollen at 3500 yr B.P. apparently recording a human population large enough to modify the natural environment, as well as the beginning of agriculture. A rich aquatic flora in this phase suggests eutrophication of the lake by slope erosion. In the most recent period corn is absent from the sediments, perhaps reflecting a change in agricultural practices. The environment changes at Lake Patzcuaro are similar to and correlate with those in the Cuenca de Mexico, where diatom stratigraphy from the Chalco basin indicates fluctuations in lake levels and lake chemistry in response to variations in available moisture. Before 10,000 yr ago climates there were cool and dry, and the Chalco basin was occupied by a shallow freshwater marsh that drained north to Lake Texcoco, where saline water accumulated by evaporation. Increases in effective moisture and possible melting of glaciers during the Holocene caused lake levels to rise throughout the Cuenca de Mexico, and Lake Texcoco flooded the Chalco basin with brackish water. After 5000 yr ago such flooding decreased, and shallow freshwater ponds and marshes were restored in the Chalco basin. This environmental change coincides with the appearance of Zea pollen and suggests cultural control of lake levels and salinity.     

A 12.5-kyr history of vegetation dynamics and mire development with evidence of Younger Dryas larch presence in the Verkhoyansk Mountains, East Siberia, Russia

Werner, K., Tarasov, P. E., Andreev, A. A., Müller, S., Kienast, F., Zech, M., Zech, W. & Diekmann, B. 2009: A 12.5‐kyr history of vegetation dynamics and mire development with evidence of Younger Dryas larch presence in the Verkhoyansk Mountains, East Siberia, Russia. Boreas, 10.1111/j.1502‐3885.2009.00116.x. ISSN 0300‐9483. A 415 cm thick permafrost peat section from the Verkhoyansk Mountains was radiocarbon‐dated and studied using palaeobotanical and sedimentological approaches. Accumulation of organic‐rich sediment commenced in a former oxbow lake, detached from a Dyanushka River meander during the Younger Dryas stadial, at ～12.5 kyr BP. Pollen data indicate that larch trees, shrub alder and dwarf birch were abundant in the vegetation at that time. Local presence of larch during the Younger Dryas is documented by well‐preserved and radiocarbon‐dated needles and cones. The early Holocene pollen assemblages reveal high percentages of Artemisia pollen, suggesting the presence of steppe‐like communities around the site, possibly in response to a relatively warm and dry climate ～11.4–11.2 kyr BP. Both pollen and plant macrofossil data demonstrate that larch woods were common in the river valley. Remains of charcoal and pollen of Epilobium indicate fire events and mark a hiatus ～11.0–8.7 kyr BP. Changes in peat properties, C31/C27 alkane ratios and radiocarbon dates suggest that two other hiatuses occurred ～8.2–6.9 and ～6.7–0.6 kyr BP. Prior to 0.6 kyr BP, a major fire destroyed the mire surface. The upper 60 cm of the studied section is composed of aeolian sands modified in the uppermost part by the modern soil formation. For the first time, local growth of larch during the Younger Dryas has been verified in the western foreland of the Verkhoyansk Mountains (～170 km south of the Arctic Circle), thus increasing our understanding of the quick reforestation of northern Eurasia by the early Holocene.     

Seasonality of modern pollen and sediment deposition in an estuarine context: the Severn Estuary Levels,southwest England

Recent sedimentological and palynological research on subfossil Holocene banded sediments from the Severn Estuary Levels suggested seasonality of deposition, registered by variations in mineral grain‐size and pollen assemblages between different parts of the bands. Here we provide data that strengthen this interpretation from sampling of modern sediments and pollen deposition on an active mudflat and saltmarsh on the margin of the Severn Estuary, and comparison with a vegetation survey and contemporary records of climate, river and tidal regimes. The results of grain‐size analysis indicate deposition of comparatively coarse‐grained silts during the relatively cool and windy conditions of winter and comparatively fine‐grained sediments during relatively warm and calm summer months. Pollen analysis demonstrates the significance of long‐term storage of pollen grains and fern spores in the estuarine waterbody, superimposed on which seasonal variations in pollen inputs from local and regional vegetation remain detectable. Copyright © 2007 John Wiley & Sons, Ltd.     

Late Wisconsinan Vegetation and Environment of the Tunica Hills Region, Louisiana/Mississippi

Pollen, plant macrofossil, and radiocarbon-dating studies of seven exposures of fluvial sediments in the Tunica Hills region of southeastern Louisiana and southwestern Mississippi provide new information on late Wisconsinan vegetation, flora, and environment of the region. The assemblages date between 25,250 and 17,530 yr B.P. Pollen and macrofossil assemblages are dominated by Picea, which comprises 40-70% of the pollen assemblages. Abies and Larix pollen and macrofossils are absent, in contrast to sites to the north in the central Mississippi Valley. Deciduous hardwoods (Quercus, Fagus, Fraxinus, Carya, Juglans nigra, Acer, Ulmus) are minor components of both pollen and macrofossil assemblages. Radiocarbon dates of Picea and Quercus wood indicate that these two genera grew contemporaneously in the region. Regional upland forests were dominated by Picea. Picea cones and cone fragments are not typical of any extant North American species, and probably represent either an extinct species or an extinct variety or subspecies of Picea glauca. Late Wisconsinan climate of the region was cooler than present, but not necessarily as cool as implied by P. glauca or other "boreal" taxa.