The role of temperature on treeline migration for an eastern African mountain during the Last Glacial Maximum

Paleo-data suggest that East African mountain treelines underwent an altitudinal shift during the Last Glacial Maximum (LGM). Understanding the ecological and physiological processes underlying treeline response to such past climate change will help to improve forecasts of treeline change under future global warming. In spite of significant improvements in paleoclimatic reconstruction, the climatic conditions explaining this migration are still debated and important factors such as atmospheric CO₂ concentration, the impact of lapse rate decreasing temperature along altitudinal gradients and rainfall modifications due to elevation have often been neglected or simplified. Here, we assess the effects of these different factors and estimate the influence of the most dominant factors controlling changes in past treeline position using a multi-proxy approach based on simulations from BIOME4, a coupled biogeography and biogeochemistry model, modified to account for the effect of elevation on vegetation, compared with pollen, and isotopic data. The results indicate a shift in mountain vegetation at the LGM was controlled by low pCO₂ and low temperatures promoting species morphologically and physiologically better adapted to LGM conditions than many trees composing the forest belt limit. Our estimate that the LGM climate was cooler than today’s by ?4.5 °C (range: ?4.3 to ?4.6 °C) at the upper limit of the treeline, whereas at 831 m it was cooler by ?1.4 °C (range: ?2.6 to ?0.6 °C), suggests that a possible lapse rate modification strongly constrained the upper limit of treeline, which may limit its potential extension under future global warming.     

The simulated atmospheric response to expansion of the Arctic boreal forest biome

Over the last century, the Arctic has warmed at twice the rate of the planet as a whole. Observational evidence indicates that this rapid warming is affecting the tundra and boreal forest biomes by changing their structure and geographic distribution. A global climate model (GCM) was used to explore the atmospheric response to boreal forest expansion by applying a one-grid cell shift of the forest into tundra. This subtle shift is meant to represent the expansion that would occur this century rather than more extreme scenarios predicted by dynamic vegetation models. Results show that this shift causes an average annual warming of 0.3 °C over the region because of a reduction in the surface albedo and an increase in net radiation. A warming of ~1.0 °C occurs in spring when the forest masks the higher albedo snow-covered surface and results in snowmelt and a reduction in cloud cover. Results fail to show a larger-scale dynamical response although some warming of the lower and mid troposphere occurs in July. No changes were found in the position or strength of the Arctic frontal zone as some studies have indicated will occur with a shift in the boreal forest-tundra boundary. These findings suggest that coupled model simulations that predict larger changes in vegetation distribution are likely overemphasizing the amount of Arctic warming that will occur this century. These findings also indicate that a realistic dynamical response to subtle land cover change might not be correctly simulated by GCMs run at coarse spatial resolutions.     

Global Warming and Potential Changes in Fish Habitat in U.S. Streams

To project potential habitat changes of 57 fish species under global warming, their suitable thermal habitat at 764 stream gaging stations in the contiguous United States was studied. Global warming was specified by air temperature increases projected by the Canadian Centre of Climate Modelling General Circulation Model for a doubling of atmospheric CO₂. The aquatic thermal regime at each gaging station was related to air temperature using a nonlinear stream temperature/air temperature relationship.Suitable fish thermal habitat was assumed to be constrained by both maximum temperature and minimum temperature tolerances. For cold water fishes with a 0 °C lower temperature constraint, the number of stations with suitable thermal habitat under a 2×CO₂ climate scenario is projected to decrease by 36%, and for cool water fishes by 15%. These changes are associated with a northward shift of the range. For warm water fishes with a 2 °C lower temperature constraint, the potential number of stations with suitable thermal habitat is projected to increase by 31%.     

Ecological responses of plant species and communities to climate warming: upward shift or range filling processes?

The fate of alpine species in response to climate warming is still unclear. We analyze effects of climate warming on the composition of alpine plants communities and unravel the range filling of communities within a belt from long-term true upward shift processes. In the European Alps we re-sampled in 2003 the vegetation at sites studied in 1953 and analyzed the changes at intra- and inter-community level. Since 1953 all communities experienced a high species turnover, leading to an overall increase in species richness as new species exceeded species losses. The dominant species mainly declined allowing the potential expansion of competitors and/or of new species. The main recruitment sources are neighbor communities within the same elevation belt performing biotic exchanges with other plant communities in the same altitudinal belts. The changes of species distribution curves with elevation emphasized that more than half of the most widespread persisting species exhibited downward shifts instead of upward shifts. Upward shifts from lower elevation belts and of nonnative species were very limited. One third of the persisting species declined and could be used as a proxy to measure the extinction debt. Therefore the fate of plant communities will depend on the ability of the original species to persist and fill the available ecological gaps. Species persistence may be crucial in developing adaptation and environmental protection strategies.     

Ungulate herbivory modifies the effects of climate change on mountain forests

Recent temperature observations suggest a general warming trend that may be causing the range of tree species to shift to higher latitudes and altitudes. Since biotic interactions such as herbivory can change tree species composition, it is important to understand their contribution to vegetation changes triggered by climate change. To investigate the response of forests to climate change and herbivory by wild ungulates, we used the forest gap model ForClim v2.9.6 and simulated forest development in three climatically different valleys in the Swiss Alps. We used altitudinal transects on contrasting slopes covering a wide range of forest types from the cold (upper) to the dry (lower) treeline. This allowed us to investigate (1) altitudinal range shifts in response to climate change, (2) the consequences for tree species composition, and (3) the combined effect of climate change and ungulate herbivory. We found that ungulate herbivory changed species composition and that both basal area and stem numbers decreased with increasing herbivory intensity. Tree species responded differently to the change in climate, and their ranges did not change concurrently, thus causing a succession to new stand types. While climate change partially compensated for the reductions in basal area caused by ungulate herbivory, the combined effect of these two agents on the mix of the dominant species and forest type was non-compensatory, as browsing selectively excluded species from establishing or reaching dominance and altered competition patterns, particularly for light. We conclude that there is an urgent need for adaptive forest management strategies that address the joint effects of climate change and ungulate herbivory.     

Modeling the Influence of Topographic Barriers on Treeline Advance at the Forest-Tundra Ecotone in Northwestern Alaska

The response of terrestrial ecosystems to climate warming has important implications to potential feedbacks to climate. The interactions between topography, climate, and disturbance could alter recruitment patterns to reduce or offset current predicted positive feedbacks to warming at high latitudes. In northern Alaska the Brooks Range poses a complex environmental and ecological barrier to species migration. We use a spatially explicit model (ALFRESCO) to simulate the transient response of subarctic vegetation to climatic warming in the Kobuk/Noatak River Valley (200 × 400 km) in northwest Alaska. The model simulations showed that a significantly warmer (+6 °C) summer climate would cause expansion of forest through the Brooks Range onto the currently treeless North Slope only after a period of 3000–4000 yr. Substantial forest establishment on the North Slope didnot occur until temperatures warmed 9 °C, and only following a 2000 yr time lag. The long time lags between change in climate and change in vegetation indicate current global change predictions greatly over-estimate the response of vegetation to a warming climate in Alaska. In all the simulations warming caused a steady increase in the proportion of early successional deciduous forest. This would reduce the magnitude of the predicted decrease in regional albedo and the positive feedback to climate warming. Simulation of spruce forest refugia on the North Slope showed forest could survive with only a 4 °C warming and would greatly reduce the time lag of forest expansion under warmer climates. Planting of spruce on the North Slope by humans could increase the likelihood of large-scale colonization of currently treeless tundra. Together, the long time lag and deciduous forest dominance would delay the predicted positive regional feedback of vegetation change to climatic warming. These simulated changes indicate the Brooks Range would significantly constrain regional forest expansion under a warming climate, with similar implications for other regions possessing major east-west oriented mountain ranges.     

Capelin migrations and climate change – a modelling analysis

The capelin is a small pelagic fish that performs long distance migrations. It is a key species in the Barents Sea ecosystem and its distribution is highly climate dependent. Here we use an individual based model to investigate consequences of global warming on capelin distribution and population dynamics. The model relies on input on physics and plankton from a biophysical ocean model, and the entire life cycle of capelin including spawning of eggs, larval drift and adult movement is simulated. Spawning day and adult movement strategies are adapted by a genetic algorithm. Spawning has to take place in designated near-shore spawning areas. The output generated by the model is capelin migration/distribution and population dynamics. We present simulations with present day climate and a future climate scenario. For the present climate the model evolves a spatial distribution resembling typical spatial dynamics of capelin with the coasts of Northern Norway and Murman as the main spawning areas. For the climate change simulation, the capelin is predicted to shift spawning eastwards and also utilize new spawning areas along Novaya Zemlya. There is also a shift in the adult distribution towards the north eastern part of the Barents Sea and earlier spawning associated with the warming.     

Numerical Simulation of Global Temperature Change during the 20th Century with the IAP／LASG GOALS Model

The IAP/LASG GOALS coupled model is used to simulate the climate change during the 20th century using historical greenhouse gases concentrations, the mass mixing ratio of sulfate aerosols simulated by a CTM model, and reconstruction of solar variability spanning the period 1900 to 1997. Four simulations, including a control simulation and three forcing simulations, are conducted. Comparison with the observational record for the period indicates that the three forcing experiments simulate reasonable temporal and spatial distributions of the temperature change. The global warming during the 20th century is caused mainly by increasing greenhouse gas concentration especially since the late 1980s; sulfate aerosols offset a portion of the global warming and the reduction of global temperature is up to about 0.11℃ over the century; additionally, the effect of solar variability is not negligible in the simulation of climate change over the 20th century.     

An assessment of the current and future thermal regimes of three streams located in the Wenatchee River basin, Washington State: some implications for regional river basin systems

We examine summer temperature patterns in the Wenatchee River and two of its major tributaries Icicle and Nason Creeks, located in the Pacific Northwest region of the United States. Through model simulations we evaluate the cooling effects of mature riparian vegetation corridors along the streams and potential increases due to global warming for the 2020s–2080s time horizons. Site potential shade influences are smaller in the mainstream due to its relatively large size and reduced canopy density in the lower reaches, proving a modest reduction of about 0.3°C of the stream length average daily maximum temperature, compared with 1.5°C and 2.8°C in Icicle and Nason Creeks. Assuming no changes in riparian vegetation shade, stream length-average daily maximum temperature could increase in the Wenatchee River from 1–1.2°C by the 2020s to 2°C in the 2040s and 2.5–3.6°C in the 2080s, reaching 27–30°C in the warmest reaches. The cooling effects from the site potential riparian vegetation are likely to be offset by the climate change effects in the Wenatchee River by the 2020s. Buffers of mature riparian vegetation along the banks of the tributaries could prevent additional water temperature increases associated with climate change. By the end of the century, assuming site potential shade, the tributaries could have a thermal condition similar to today’s condition which has less shade. In the absence of riparian vegetation restoration, at typical summer low flows, stream length average daily mean temperatures could reach about 16.4–17°C by the 2040s with stream length average daily maxima around 19.5–20.6°C, values that can impair or eliminate salmonid rearing and spawning. Modeled increases in stream temperature due to global warming are determined primarily by the projected reductions in summer streamflows, and to a lesser extent by the increases in air temperature. The findings emphasize the importance of riparian vegetation restoration along the smaller tributaries, to prevent future temperature increases and preserve aquatic habitat.     

Shrubland encroachment in southern New Mexico,U.S.A.: An analysis of desertification processes in the American southwest

The area dominated by the shrubs creosotebush (Larrea tridentata) and mesquite (Prosopis glandulosa) in the American southwest has increased several-fold over the last century, with a corresponding decrease in areal coverage of productive grasslands and increased surface soil erosion throughout the region. The factors thought to be responsible for this regional shift in vegetation are: (1) overgrazing by domestic livestock; (2) fire suppression; and (3) historical changes in climate. We examine the evidence concerning each of these factors and develop a synthetic model outlining the principles affecting shrubland encroachment, which focuses on life history characteristics of the dominant shrubs and a number of positive biotic and edaphic feedback mechanisms contributing to their establishment and persistence. We conclude that the expansion of shrub dominance that has occurred over the last century may have been triggered by extreme live-stock overgrazing at the end of the nineteenth century, which coincided with rainfall regimes that were unfavorable for perennial grass growth. Hence, the landscape we observe today may be a product of positive feedback mechanisms triggered over a century ago by management practices that were uninformed with regard to the importance of historical climate patterns and the life history characteristics of important rangeland species. Our consideration of these issues also addresses potential land surface - climate interactions that could occur as a result of regional alterations in vegetation dominance and physiognomy.     

Temporal and Spatial Variations in the Climate Controls of Vegetation Dynamics on the Tibetan Plateau during 1982–2011

The ecosystem of the Tibetan Plateau is highly susceptible to climate change. Currently, there is little discussion on the temporal changes in the link between climatic factors and vegetation dynamics in this region under the changing climate.By employing Normalized Difference Vegetation Index data, the Climatic Research Unit temperature and precipitation data,and the in-situ meteorological observations, we report the temporal and spatial variations in the relationships between the vegetation dynamics and climatic factors on the Plateau over the past three decades. The results show that from the early 1980s to the mid-1990s, vegetation dynamics in the central and southeastern part of the Plateau appears to show a closer relationship with precipitation prior to the growing season than that of temperature. From the mid-1990s, the temperature rise seems to be the key climatic factor correlating vegetation growth in this region. The effects of increasing temperature on vegetation are spatially variable across the Plateau: it has negative impacts on vegetation activity in the southwestern and northeastern part of the Plateau, and positive impacts in the central and southeastern Plateau. In the context of global warming, the changing climate condition(increasing precipitation and significant rising temperature) might be the potential contributor to the shift in the climatic controls on vegetation dynamics in the central and southeastern Plateau.     

Global vegetation change predicted by the modified Budyko model

A modified Budyko global vegetation model is used to predict changes in global vegetation patterns resulting from climate change (CO₂ doubling). Vegetation patterns are predicted using a model based on a dryness index and potential evaporation determined by solving radiation balance equations. Climate change scenarios are derived from predictions from four General Circulation Models (GCM's) of the atmosphere (GFDL, GISS, OSU, and UKMO). Global vegetation maps after climate change are compared to the current climate vegetation map using the kappa statistic for judging agreement, as well as by calculating area statistics. All four GCM scenarios show similar trends in vegetation shifts and in areas that remain stable, although the UKMO scenario predicts greater warming than the others. Climate change maps produced by all four GCM scenarios show good agreement with the current climate vegetation map for the globe as a whole, although over half of the vegetation classes show only poor to fair agreement. The most stable areas are Desert and Ice/Polar Desert. Because most of the predicted warming is concentrated in the Boreal and Temperate zones, vegetation there is predicted to undergo the greatest change. Specifically, all Boreal vegetation classes are predicted to shrink. The interrelated classes of Tundra, Taiga, and Temperate Forest are predicted to replace much of their poleward (mostly northern) neighbors. Most vegetation classes in the Subtropics and Tropics are predicted to expand. Any shift in the Tropics favoring either Forest over Savanna, or vice versa, will be determined by the magnitude of the increased precipitation accompanying global warming. Although the model predicts equilibrium conditions to which many plant species cannot adjust (through migration or microevolution) in the 50–100 y needed for CO₂ doubling, it is nevertheless not clear if projected global warming will result in drastic or benign vegetation change.     

Elevation gradients of European climate change in the regional climate model COSMO-CLM

A transient climate scenario experiment of the regional climate model COSMO-CLM is analyzed to assess the elevation dependency of 21st century European climate change. A focus is put on near-surface conditions. Model evaluation reveals that COSMO-CLM is able to approximately reproduce the observed altitudinal variation of 2 m temperature and precipitation in most regions and most seasons. The analysis of climate change signals suggests that 21st century climate change might considerably depend on elevation. Over most parts of Europe and in most seasons, near-surface warming significantly increases with elevation. This is consistent with the simulated changes of the free-tropospheric air temperature, but can only be fully explained by taking into account regional-scale processes involving the land surface. In winter and spring, the anomalous high-elevation warming is typically connected to a decrease in the number of snow days and the snow-albedo feedback. Further factors are changes in cloud cover and soil moisture and the proximity of low-elevation regions to the sea. The amplified warming at high elevations becomes apparent during the first half of the 21st century and results in a general decrease of near-surface lapse rates. It does not imply an early detection potential of large-scale temperature changes. For precipitation, only few consistent signals arise. In many regions precipitation changes show a pronounced elevation dependency but the details strongly depend on the season and the region under consideration. There is a tendency towards a larger relative decrease of summer precipitation at low elevations, but there are exceptions to this as well.     

Influence of dynamic vegetation on climate change arising from increasing CO<Subscript>2</Subscript>

A dynamic global vegetation model (DGVM) is coupled to an atmospheric general circulation model (AGCM) to investigate the influence of vegetation dynamics on climate change under conditions of global warming. The model results are largely in agreement with observations and the results of previous studies in terms of the present climate, present potential vegetation, present net primary productivity (NPP), and pre-industrial carbon budgets. The equilibrium state of climate properties are compared among pre-industrial, doubled, and quadrupled atmospheric CO₂ values using DGVM–AGCM and current AGCM with fixed vegetation to evaluate the influence of dynamic vegetation change. We also separated the contributions of temperature, precipitation and CO₂ fertilization on vegetation change. The results reveal an amplification of global warming climate sensitivity by 10% due to the inclusion of dynamic vegetation. The total effects of elevated CO₂ and climate change also lead to an increase in NPP and vegetation coverage globally. The reduction of albedo associated with this greening results in enhanced global warming. Our separation analysis indicates that temperature alters vegetation at high latitudes such as Siberia or Alaska, where there is a switch from tundra to forest. On the other hand, CO₂ fertilization provides the largest contribution to greening in arid/semi-arid region. Precipitation change did not cause any drastic vegetation shift.     

Arctic climate change with a 2 ∘C global warming: Timing, climate patterns and vegetation change

The signatories to United Nations Framework Convention on Climate Change are charged with stabilizing the concentrations of greenhouse gases in the atmosphere at a level that prevents dangerous interference with the climate system. A number of nations, organizations and scientists have suggested that global mean temperature should not rise over 2 ^°C above preindustrial levels. However, even a relatively moderate target of 2 ^°C has serious implications for the Arctic, where temperatures are predicted to increase at least 1.5 to 2 times as fast as global temperatures. High latitude vegetation plays a significant role in the lives of humans and animals, and in the global energy balance and carbon budget. These ecosystems are expected to be among the most strongly impacted by climate change over the next century. To investigate the potential impact of stabilization of global temperature at 2 ^°C, we performed a study using data from six Global Climate Models (GCMs) forced by four greenhouse gas emissions scenarios, the BIOME4 biogeochemistry-biogeography model, and remote sensing data. GCM data were used to predict the timing and patterns of Arctic climate change under a global mean warming of 2 ^°C. A unified circumpolar classification recognizing five types of tundra and six forest biomes was used to develop a map of observed Arctic vegetation. BIOME4 was used to simulate the vegetation distributions over the Arctic at the present and for a range of 2 ^°C global warming scenarios. The GCMs simulations indicate that the earth will have warmed by 2 ^°C relative to preindustrial temperatures by between 2026 and 2060, by which stage the area-mean annual temperature over the Arctic (60–90^°N) will have increased by between 3.2 and 6.6 ^°C. Forest extent is predicted by BIOME4 to increase in the Arctic on the order of 3 × 10⁶ km² or 55% with a corresponding 42% reduction in tundra area. Tundra types generally also shift north with the largest reductions in the prostrate dwarf-shrub tundra, where nearly 60% of habitat is lost. Modeled shifts in the potential northern limit of trees reach up to 400 km from the present tree line, which may be limited by dispersion rates. Simulated physiological effects of the CO₂ increase (to ca. 475 ppm) at high latitudes were small compared with the effects of the change in climate. The increase in forest area of the Arctic could sequester 600 Pg of additional carbon, though this effect is unlikely to be realized over next century.     

全球变暖中的科学问题

2013年各国政府间气候变化专门委员会（IPCC）第一工作组发布了第五次气候变化科学评估报告,以大量的观测分析和气候模式模拟证据,继续强调由于人类排放增加,全球正在变暖,未来将继续变暖的观点。本文综述研究全球变暖的几个深层次的科学问题,即多套全球气温观测资料的差异、不同标准气候态时段的作用、20世纪全球变暖的检测和归因及未来全球气温变化的走向,以此提出需进一步研究的科学问题。结果表明;需要进一步提高观测资料的质量;注意不同标准气候态时段对应的数值的不同;应进一步改善气候模式模拟年代际变率的能力及研究近15 a全球变暖减缓和停滞的原因,从而改善气候模式的模拟效果;造成预估未来全球气候变化的不确定性主要来自气候模式的差异、未来排放情景的差异及气候系统内部变率影响和自然外强迫的作用。     

Modelling Regional Climate Change Effects On Potential Natural Ecosystems in Sweden

This study aims to demonstrate the potential of a process-based regional ecosystem model, LPJ-GUESS, driven by climate scenarios generated by a regional climate model system (RCM) to generate predictions useful for assessing effects of climatic and CO₂ change on the key ecosystem services of carbon uptake and storage. Scenarios compatible with the A2 and B2 greenhouse gas emission scenarios of the Special Report on Emission Scenarios (SRES) and with boundary conditions from two general circulation models (GCMs) – HadAM3H and ECHAM4/OPYC3 – were used in simulations to explore changes in tree species distributions, vegetation structure, productivity and ecosystem carbon stocks for the late 21st Century, thus accommodating a proportion of the GCM-based and emissions-based uncertainty in future climate development. The simulations represented in this study were of the potential natural vegetation ignoring direct anthropogenic effects. Results suggest that shifts in climatic zones may lead to changes in species distribution and community composition among seven major tree species of natural Swedish forests. All four climate scenarios were associated with an extension of the boreal forest treeline with respect to altitude and latitude. In the boreal and boreo-nemoral zones, the dominance of Norway spruce and to a lesser extent Scots pine was reduced in favour of deciduous broadleaved tree species. The model also predicted substantial increases in vegetation net primary productivity (NPP), especially in central Sweden. Expansion of forest cover and increased local biomass enhanced the net carbon sink over central and northern Sweden, despite increased carbon release through decomposition processes in the soil. In southern Sweden, reduced growing season soil moisture levels counterbalanced the positive effects of a longer growing season and increased carbon supply on NPP, with the result that many areas were converted from a sink to a source of carbon by the late 21st century. The economy-oriented A2 emission scenario would lead to higher NPP and stronger carbon sinks according to the simulations than the environment-oriented B2 scenario.     

21世纪气候变化情景下环北极地区植被生长季与活动积温变化

基于部门间影响模式比较计划（ISI-MIP, Inter-Sectoral Impact Model Intercomparison Project)对CMIP5中5个气候（地球）系统模式模拟结果的降尺度数据,利用多模式集合预估了气候变化情景下21世纪环北极地区植被生长季与活动积温变化。研究发现:1）多模式集合模拟能够基本再现观测的初、终霜日及无霜期长度与>10°C积温的空间分布特征以及1979～2004年各指标变化趋势的空间分布特征,但其对气候变化年际变率的模拟能力较弱;2）至21世纪末,终霜日最多将提前60 d,初霜日将推迟20～40 d,无霜期延长幅度最高可达100 d,积温将增加1000～1200°C。其中RCP8.5情景下,各指标变幅最大,RCP2.6情景下变幅最小;3）各指标变幅呈现出较大的空间差异,亚欧大陆中西部的变幅普遍较大,随着气候变暖,>10°C积温增加幅度表现出明显的纬度地带性,南部增幅较大,北部增幅较小。     

Using a global climate model to evaluate the influences of water vapor, snow cover and atmospheric aerosol on warming in the Tibetan Plateau during the twenty-first century

We examine trends in climate variables and their interrelationships over the Tibetan Plateau using global climate model simulations to elucidate the mechanisms for the pattern of warming observed over the plateau during the latter half of the twentieth century and to investigate the warming trend during the twenty-first century under the SRES A1B scenario. Our analysis suggests a 4°C warming over the plateau between 1950 and 2100. The largest warming rates occur during winter and spring. For the 1961–2000 period, the simulated warming is similar to the observed trend over the plateau. Moreover, the largest warming occurs at the highest elevation sites between 1950 and 2100. We find that increases in (1) downward longwave radiation (DLR) influenced by increases in surface specific humidity (q), and (2) absorbed solar radiation (ASR) influenced by decreases in snow cover extent are, in part, the reason for a large warming trend over the plateau, particularly during winter and spring. Furthermore, elevation-based increases in DLR (influenced by q) and ASR (influenced by snow cover and atmospheric aerosols) appear to affect the elevation dependent warming trend simulated in the model.