中国古尔班通古特沙漠东部中心地区植物分布与植物—土壤关系(英文)

The distribution of plant species and relationships between species and soil factors in the east central part of Gurbantunggut Desert was studied to provide more insight into the flora and determine differences in vegetation across various parts of the desert. Two-way Cluster Analysis showed that the vegetation in the area could be divided into three groups, the first group was dominated by the shrub species, Ephedra przewalskii and the grass species, Carex physodes mainly in areas of flat grounds and gentle slopes; the second group was dominated by C. physodes, Artemisia songorica and A. xerophytica mainly on the slope of sand dunes and the third group was dominated by the shrub species, Haloxylon persicum mainly on the top of sand dunes. There was no difference in plant density between Groups 1 and 2 but there was a significant decrease in Group 3. Soil water under vegetation Group 3 was much lower than that in the other two groups at all soil depths. The EC, organic matter, total P and soluble Na, Ca and Mg varied very similarly with soil water. Canonical correspondence analysis(CCA) satisfactorily assessed the species-soil relations in the area. The distribution of plant species was strongly correlated with the soil factors of water content, organic matter, EC and nutrients. The variations in species occurrence explained by the three CCA axes were about 70%, indicating that some explanatory site variables may exist outside our studied parameters. Soil texture is suggested to be included in future studies to improve the explanation of CCA.     

河西走廊荒漠绿洲过渡带封育对土壤和植被的影响

在河西走廊荒漠绿洲过渡带,封育天然植被是植被群落恢复、防止绿洲沙漠化的有效措施。以流动沙丘作为对照(0年),对封育5年和15年的半固沙和固定沙丘植被群落以及土壤进行调查取样和分析。结果表明:随着封育年限增加,天然固沙植被群落生物多样性增加,灌木层和草本层植物密度、盖度和生物量都显著增加,灌木层盖度从10%增加到40%,草本层以一年生草本植物为主,物种从5种增加到8种,生物量从1 g·m^-2增加到13 g·m^-2。随着天然植被盖度增加,土壤表层沙土细粒化明显,沙土中黏粉粒含量显著增加,土壤质地由粗质沙粒向细质沙粒转变;随着沙土中黏粉粒成分的增加,沙土有机质、全氮、全磷含量也增加,灌丛下土壤养分含量高于灌丛间,“沃岛效应”明显。同时,在灌丛下表层土壤出现明显的盐分集聚现象,其中SO₄^2-、K⁺、Na⁺含量分别增加了6、3、17倍。在降水100 mm左右的荒漠绿洲过渡带,封育可以显著恢复固沙植被群落和提高沙土质地和养分。     

灌丛对流动沙地土壤特性和草本植物的影响

通过对流动沙地灌丛内外土壤特性、土壤养分含量、土壤种子库和草本植物群落特征的差异性调查,分析了灌丛对沙地土壤特性和林下草本植被的影响。结果表明,在流动沙地0—20 cm土壤中细沙、极细沙、粘粉粒、有机质、总氮和总磷、有效磷和土壤水分含量,小叶锦鸡儿灌丛下分别较灌丛外高17.3%、4.4%、 49.5%、43.8%、40.0%、23.1%、16.3%和10.8%,黄柳灌丛下较灌丛外分别高3.5%、21.3%、0.0%、20.0%、16.7%、8.3%、10.6%和28.1%。小叶锦鸡儿、差不嘎蒿和黄柳灌丛下凋落物蓄积量要比灌丛外分别高18.3倍、365.2倍和15.5倍。差不嘎蒿灌丛下土壤种子库密度较灌丛外高10.9倍。原为半固定、半流动沙地优势种的多年生草本植物白草,不仅能在流动沙地灌丛下存活,而且具有较高的密度、高度、盖度和地上生物量。结果还表明,从灌丛中心到灌丛边缘,凋落物产量、土壤种子库密度、草本植物密度、盖度、生物量均存在明显的递减梯度,在灌丛外不远处消失。这些结果说明,在流动沙地,灌丛具有明显的“肥岛”效应和“保种”作用。     

乌兰布和沙漠典型植物群落土壤风蚀可蚀性研究

为探明植被恢复对乌兰布和沙漠土壤风蚀可蚀性的影响,以乌兰布和沙漠内不同沙地固 定阶段的 8 种典型植物群落及群落内表土作为研究对象,对土壤物理因子（可蚀性颗粒含量、土壤 含水量、有机质含量）、土壤结皮因子、植被因子 3 类土壤风蚀可蚀性因子指标进行监测,分析土壤 风蚀可蚀性因子在不同植物群落类型间、沙地固定阶段间的差异。结果表明：（1）在乌兰布和沙漠 典型植物群落中,沙蓬、沙生针茅、盐爪爪等草本植物群落的土壤风蚀可蚀性最强,白刺、梭梭、沙 冬青等灌木植物群落土壤风蚀可蚀性弱于草本植物群落,说明灌木林能显著降低土壤风蚀作用。 （2）随着沙地的不断固定,土壤结构不断发育,土壤可蚀性不断降低,土壤风蚀可蚀性强弱表现为 固定沙地<半固定沙地<流动沙地。（3）土壤可蚀性颗粒含量、土壤有机质含量、土壤含水量、土壤结 皮、植被因子与植被类型及沙地固定阶段具有显著相关关系。因此,在沙区生态建设工程中,为了 减少土壤风蚀量,不仅要考虑物种的选择,还要促进人工生态系统的演替和恢复,从而有效降低土 壤风蚀可蚀性。研究结果可为乌兰布和沙区植被生态系统服务功能的科学评价、防沙治沙工程的 建设与管理提供一定参考。     

Vegetation patterns and species–environment relationships in the Gurbantunggut Desert of China

The Gurbantunggut Desert is the largest stable and semi-stable sand desert in China, yet few data exist on vegetation pattern and species-environment relationships for these diverse desert landscapes. The sand dunes of the survey area are mainly of the longitudinal form from north to south, but checkerboard-shaped and honeycomb-shaped forms are also present, with the height of 10–50 m. We measured vegetation and soil data on north–south transects and compared them with vegetation and soil data on east–west transects. Analysis revealed that the varying trend and strength of the species diversity, dominance and cover of the plant communities in the longitudinal and transverse directions across the landscape are significantly different. The results of CCA ordination show that the microhabitats of soil organic matter (OM), soil salts (TS), sorting index (σ), topsoil water-content (SM) and pH relate to the differences in vegetation observed as differences in species assemblage from salt-intolerant ephemerals, micro-subshrubs and subshrubs to salt-tolerant shrubs and micro-arbors. The terrain (alt.) and soil texture (the contents of Mz and φ1) affect the spatial differentiation of many species. However, this spatial differentiation is not so marked on transects running longitudinally with the landscape, in the same direction as the dunes. The species of the desert vegetation have formed three assemblages under the action of habitat gradients, relating to three sections running transversely across the landscape, at right angles to the direction of the dune crests. In the mid-east section of the study area the topography is higher, with sand-lands or dune-slopes with coarse particles. Here the dominant vegetation comprises shrubs and subshrubs of Seriphidium santolium and Ephedra distachya, with large numbers of ephemeral and ephemeroid plants of Senecio subdentatus, and Carex physodes in spring and summer. On the soil of the dune-slopes in the mid-west of the study area, with coarse particles and abundant TK, the plant assemblage of Haloxylon persicum, Soranthus meyeri and Agriophyllum squarrosum is developed. The species composition in the east marginal belt of the study area has similar characteristics to the mid-west section. There is no corresponding section in the north–south transects (except for the north and south margins). This is because the habitats of most plants are located in the middle of the microhabitat gradients in the north–south direction in the desert.     

沙坡头固定沙丘结皮层的微生物区系动态

对中科院沙坡头沙漠研究试验站的人工植被固定沙丘、红卫自然植被固定沙丘的结皮层和流动沙丘表层(0~ 05cm)中的微生物类群数量的研究结果表明:①好气性细菌数量影响着微生物总数量的变化趋势;②微生物总数量分布依次排列为:自然植被固定沙丘>1956年栽植区>1964年栽植区>自然半固定沙丘>1982年栽植区>流动沙丘;③土壤微生物类群数量与结皮层的形成、植物覆盖度和土壤含水率等因子有密切的关系;④结皮层中的微生物类群数量与流动沙丘的固定程度呈正向关系。     

Changes in soil and vegetation on moving sand dunes after exclosure in Horqin Sandy Land, Northern China

In the semiarid Horqin Sandy Land of northern China, land desertification is the main causation in vegetation degradation and formation of moving dunes. A study was conducted from 1996 to 2005 to monitor the changes of vegetation characteristics and soil properties after moving dunes were fenced. The changes were compared between moving sand dunes with exclosure and without exclosure to evaluate the effectiveness of vegetation and soil restoration after exclosure establishment. The results show that exlosure establishment facilitated the colonization and development of plant species by ameliorating stressful environmental conditions. Species diversity, average coverage, and plant density significantly increased after exclosure of moving sand dunes along sequence compared with sand dunes without exclosure. Vegetation recovery on moving sand dunes accelerated by exclosure resulted in significant changes in soil properties including increased silt and clay contents, organic C and total N and decreased sand content, especially at the 0-5 cm depth. The results implied that moving sand dunes can be rapidly fixed by construction of exclosure.     

古尔班通古特沙漠纵向沙垄植被空间异质性

在古尔班通古特沙漠南部纵向沙垄上按5 m×10 m的网格测度植被样方100个,采集浅层风沙土样品184件。调查发现,乔灌木层片和草本层片的分布及相互关系明显受沙垄地形影响,并且在干旱环境下种群间的竞争和空间上的互补使两层片生态优势度具有显著的相关性。地统计分析结果显示,纵向沙垄上植被空间异质性表现为木本层片的物种多样性呈斑块状分布,草本层片物种多样性呈现平行沙垄的带状结构和斑块结构叠加的特点,而植被盖度则在垂直沙垄走向的方向上具有强烈的梯度性变化。和沙垄土壤理化性质空间格局特征相对比,植被盖度及物种多样性与风沙土浅层含水量在自相关空间（变程）及空间异质性程度上比较接近。这是纵向沙垄小生态系统中草本植物和浅层土壤水分耦合关系在空间上的反映,也是木本植物对水分和土壤资源在小尺度（m）上竞争及中小尺度（10～100 m）上互补的结果。     

沙坡头地区铁路两侧流沙上人工植被的生态效果

包兰铁路通过的沙坡头地区, 位于腾格里沙漠东南缘, 地貌为起伏连绵的高大格状流动沙丘。因地处西北内陆, 加之地势较高, 具有降雨少而集中, 气候干旱, 温度变化剧烈和风大沙多、植被稀少的特点。是典型的半荒漠地带。     

腾格里沙漠不同类型沙丘土壤水分含量与地形-植被因子关系研究

土壤水分是沙区主要的生态限制因子,其分布受气候、地形和植被等众多因素的影响。以腾格里沙漠沙坡头地区3种类型的沙丘（固定沙丘、半固定沙丘和流动沙丘）为研究对象,利用方差分析和冗余分析（RDA）等方法对沙丘不同部位和不同深度土壤水分的分布特征及其与地形-植被因子之间的关系进行了综合分析。结果表明：（1） 不同类型沙丘上0~300 cm的土壤水分随着深度的增加而增加,表层土壤水分的波动程度大于中层和深层。（2） 固定沙丘不同部位及不同深度的土壤水分之间没有明显的差异,半固定沙丘和流动沙丘迎风坡与丘底的土壤水分高于背风坡和丘顶。（3） 固定沙丘上的土壤水分受地形-植被因子的影响较半固定沙丘和流动沙丘小,影响固定沙丘土壤水分的主要因子有坡向、高差和灌木多度。（4） 地形-植被因子与研究区绝大多数半固定沙丘和流动沙丘的土壤水分均有负相关关系。研究揭示了腾格里沙漠土壤水分的分布规律及其与地形-植被因子的关系,对制定相应的防风固沙措施以及建立科学合理的植物固沙模式有积极的指导作用。     

古尔班通古特沙漠生物土壤结皮影响下的土壤酶分布特征

对古尔班通古特沙漠典型沙垄生物土壤结皮分布区的土壤酶分布特征进行了定量研究。结果表明：①土壤酶的分布在典型沙垄的不同地貌部位具有空间异质性,表现为垄间低地>沙垄坡部>垄顶,垄顶与垄间低地的过氧化氢酶、脲酶、中性磷酸酶、碱性磷酸酶、蛋白酶活性存在极显著差异（P<0.01）;②垂直分布上,土壤酶的积累以表层0～2 cm为主,随土壤深度增加显著递减,以垄间低地表现的尤为明显;③地貌部位与土层深度的交互效应对中性磷酸酶、碱性磷酸酶、转化酶均有极显著影响（P<0.01）;④相关性分析表明,有机质、全N是影响土壤酶活性的重要因素;各种酶促反应既是专性的,又是相互联系的;⑤主成分分析表明,过氧化氢酶和脲酶能够较好地反映土壤综合状况,与大多数土壤因子均呈极显著相关（P<0.01）。尤其是脲酶,能够作为反映典型沙垄生物土壤结皮区土壤肥力的主要指标。     

Species diversity and its relation with soil factors under different site conditions in a desert-oasis ecotone

Vegetation and soil surveys were conducted under different site conditions in 2007–2011 to study species diversity using richness, evenness and diversity indices, in the middle portion of the Heihe River Basin. The relationship between species distribution and soil environmental factors was also studied by Canonical Correspondence Analysis (CCA). Results show that vegetation coverage and species diversity were the highest in the interdune lowland, and the lowest in the mobile dune. Results of the Hill’s index (diversity ordering) shows that species diversity is reduced along decreasing soil water content, and the order of species diversity was interdune lowland, flat slope, fixed dune, semifixed dune and mobile dune. The influence degree of soil factors on vegetation distribution was soil water content > pH > total K > organic matter > available N > total N > available K > total P > saline content > available P. Soil water content and pH were important factors significantly affecting spatial distribution difference of vegetation, the environmental explanation was 98%.     

科尔沁沙地两种固沙灌木林地土壤理化性质和酶活性比较

小叶锦鸡儿(Caragana microphylla)和差不嘎蒿(Artemisia halodendron)是科尔沁沙地典型的固沙灌木,它们在退化沙地的恢复过程中起着重要的作用。本研究对这两种灌木林地表层土壤理化性质和酶活性进行了比较。结果表明:小叶锦鸡儿比差不嘎蒿对沙丘土壤理化性质和酶活性的改善作用较大。其中,小叶锦鸡儿林地土壤黏粉粒含量比差不嘎蒿高3倍左右,土壤含水量是差不嘎蒿的1.31倍,土壤有机碳和全氮分别是差不嘎蒿的1.35和1.36倍,土壤过氧化物酶和脲酶活性分别是差不嘎蒿林地土壤的2和3倍。科尔沁沙地流动沙丘中种植小叶锦鸡儿和差不嘎蒿后,两种灌木均能改善沙丘土壤理化性质和酶活性,但是其改善能力存在一定差异,总体来说,在退化沙地土壤肥力和生物活性的恢复方面,小叶锦鸡儿的优势高于差不嘎蒿。     

腾格里沙漠沙坡头地区固沙植被对生物多样性恢复的长期影响

地处腾格里沙漠东南缘的沙坡头人工固沙植被始建于1956年,46 a来不仅确保了包兰铁路沙漠地段的畅通无阻,而且对区域生态环境的恢复产生了巨大的影响,成为我国干旱沙漠地区交通干线荒漠化防治与生态恢复的成功模式。长期定位监测结果表明:人工固沙植被建立4~5 a后,沙丘表面物理结构初步得到稳定,并由大气降尘形成的无机土壤结皮逐渐演变形成土壤微生物结皮。荒漠藻类、苔藓和地衣等隐花植物在结皮层中得到了大量的繁衍:固沙植被建立46 a后出现藻类24种;苔藓仅有5 种,少于天然固定沙丘结皮上的种类,此外,地衣也在植被区发现,这说明固定沙丘景观逐渐趋于稳定的状态;相对于流沙区,固沙植被区近地面风速降低了 40%,土壤有机质含量增加了60%,其中氮、磷、钾等荒漠生态系统主要限制养分因子及土壤理化性质得到了改善,沙丘表层成土过程明显;土壤水分循环的时空变异驱动了植被的演变,为大量的草本的侵入和定居创造了条件;此外,对鸟类、昆虫和土壤动物及荒漠动物的生存产生了积极的影响。46 a后,固沙植被区共有鸟类 28 种,昆虫 50 种,动物 23种。生物多样性的恢复使原有的相对单一的固沙植被演变成一个结构、组成和功能相对复杂的荒漠生态系统。沙坡头地区生态环境在人为促进下的恢复为我国西部生态建设提供了科学依据。     

Effects of the abandonment of the burrowing mounds of fat sand rat (Psammomys obesus cretzschamar 1828) on vegetation and soil surface attributes along the coastal dunes of North Sinai,Egypt

Fat sand rats (Psammomys obesus) are strictly herbivorous and live in densely complex burrows in the desert of North Africa and the Middle East. Little is known however about the effect of their foraging and burrowing activities on surface morphology and plant community attributes. This study evaluated such effects by comparing burrow and mound surface morphology, canopy of the main host chenopod shrub Anabasis articulata, vegetation cover, and plant abundance and species richness on and off both active and abandoned colonies in the semi-stabilized sand dunes of the Northern Sinai. In general, active burrow systems were characterized by reduced A. articulata canopy area, and more soil disturbance, with higher and larger burrow mounds dominated by bare ground, dung, and dead and fresh litter. The abandonment of mounds for five years has resulted in significant increases of plant cover, canopy height, abundance and species richness. Vegetation structure and plant species assemblages differed between mound and non-mound patches of both active and abandoned sites. The results suggest that fat sand rats can have significant direct and indirect, short-term and long-term effects on vegetation dynamics and structure through their mound building and foraging.     

呼伦贝尔沙地北部沙带植物群落分布格局与土壤特性的关系

研究呼伦贝尔沙地植物群落与土壤特征相互关系有助于促进中国北方的生态恢复与重建工作。分析了呼伦贝尔沙地北部沙带58个样地174个样方的植物和土壤的调查数据,调查到94种植物,分属于26科64属,其中菊科（Compositae）、禾本科（Gramineae）、豆科（Leguminosae）和蔷薇科（Rosaceae）为前四大科,占总物种数55.33%。采用Ward系统分类法,将所调查植被分为4类。寸草薹-二裂委陵菜群落（Carex duriuscula-Potentilla bifurca） Tsallis多样性最高,冰草-星毛委陵菜-糙隐子草群落（Agropyron cristatum-Potentilla acaulis-Cleistogenes squarrosa） Tsallis多样性最低。不同植物群落间土壤物理特性有显著差异,化学特征差异不显著。随着土层深度的增加,各群落土壤全氮、全磷、有机质含量逐渐降低,表聚现象明显,脚薹草-贝加尔针茅群落（Carex pediformis-Stipa baicalensis）和冰草-星毛委陵菜-糙隐子草群落养分层次特征显著,羊草群落（Leymus chinensis）和寸草薹-二裂委陵菜群落养分层次特征不明显。CCA排序结果表明影响植被群落物种分布的土壤因子有全氮、全磷、有机质、pH和土壤水分,其中深层土壤全氮、全磷、有机质（20~40 cm土层）的作用最显著。呼伦贝尔沙地北部沙带植物群落分布存在较显著差异,呈现出斑块状分布规律,而土壤养分对植被群落格局具有重要影响。     

Comparison of soil physico-chemical properties under different land-use and cover types in northeastern China's Horqin Sandy Land

The Horqin Sandy Land of northeastern China was originally a grassland with plenty of water and lush vegetation dominated by palatable grass species along with sparsely scattered woody species. However, it has experienced severe desertification in recent decades due to its fragile ecology together with inappropriate human activities. Currently, the landscape of the Horqin Sandy Land is dominated by irrigated croplands and sand dunes with different degrees of vegetation cover, as the region has become the most important part of the semiarid agro-pastoral ecotone of northern China. In this study, we compared soil physical and chemical properties under different land-use and cover types (irrigated cropland, rainfed cropland, sandy grassland, fixed dunes, and mobile dunes). We found that soil particle size distribution; organic C, total N, and total mineral element, microelement, and available microelement and nutrient contents; pH; CEC; and bulk density differed significantly among the land-use and cover types. In general, soil quality was highest in the cropland, intermediate in the sandy grassland, and lowest in the dunes. The most important soil quality attribute, soil organic carbon (SOC) storage, decreased in the following order: irrigated cropland (5,699 g/m²) > sandy grassland (3,390 g/m²) > rainfed cropland (2,411 g/m²) > fixed dunes (821 g/m²) > mobile dunes (463 g/m²). SOC was significantly positively correlated with a large proportion of the other soil physico-chemical parameters. Our results suggest that the key issue in restoration of the degraded soils will be to increase SOC storage, which would also create a high potential for sequestering soil C in desertified areas of the Horqin Sandy Land.     

不同固沙区结皮中微生物生物量和数量的比较研究

沙地生态系统是温带干旱区、半干旱区的重要草地类型, 沙地的稳定性是生态系统健康的重要条件。在中国科学院沙坡头沙漠试验研究站, 笔者对不同程度固沙区结皮和流动沙丘表层中的微生物生物量和数量进行了比较研究。结果表明: 自然和人工植被固沙区结皮及流沙表层的微生物数量分布不同, 细菌数量比真菌和放线菌数量多数百倍, 在所有微生物类群中, 细菌数量最大, 微生物总数的变化取决于好气性细菌数量的多少, 但生物量却与之不同。在不同程度固沙区结皮中, 微生物生物量大小依次排列为: 自然植被区>1956年人工植被区>1964年人工植被区>1982年人工植被区>流沙区, 微生物生物量在自然植被固沙区中最多, 分别是1956年、1964年、1982年人工植被固沙区和流沙区的2.63、4.17、9.25和44.29倍, 表明微生物生物量随人工植被的栽植年代增加而增大, 在流动沙丘中最小, 而菌丝生物量方面是1956年人工植被固沙区已与自然植被固沙区十分接近。微生物生物量和数量与沙丘固定程度、人工植被栽植年代、结皮厚度、苔藓种类等均呈正相关, 人工植被栽植年代越长, 结皮越厚、微生物生物量和数量也越大。在不同年代人工植被固沙区结皮中, 微生物生物量和数量与相对稳定的自然植被固沙区相比, 仍未达到稳定状态。     

毛乌素沙地沙障环境下的沙丘迎风坡植被及土壤养分变化

沙障在固定流沙、促进植被恢复和改良土壤等生态建设方面起着重要作用。对毛乌素沙地设置沙柳（Salix psammophila）沙障6、8、10、11、12年的沙丘迎风坡（裸露沙丘地作为对照）开展了植被调查和土壤养分测试分析。结果显示：设障8年后的沙丘迎风坡0—30 cm深度内的土壤有机质、全氮含量显著（P<0.05）高于流动沙丘,且设障10—11年沙丘迎风坡植被盖度、0—80 cm深度内土壤有机质和全氮含量达到最大,但全磷和全钾含量无明显的变化,所有样地间和3个深度层间的全磷、全钾含量均无显著性差异;植被总盖度、0—80 cm深度内土壤有机质含量及全氮含量相互之间存在极显著线性正相关关系（P<0.01）,而且土壤pH值随有机质含量增加而线性降低。沙柳沙障固沙地的植被盖度、土壤有机质和全氮相互之间存在趋同性,且布设10年的沙障土壤改良和植被恢复成效达到最大,之后表现出下降趋势。据此建议活体沙柳沙障在维持其固沙功效的前提下以10年为限需进行平茬处理。     

荒漠人工固沙植被区土壤结皮斥水性发展特征

土壤结皮深刻影响绿洲边缘固沙植被区的土壤水文过程。土壤结皮斥水性是量化上述影响的潜在指标。采用滴水穿透时间（WDPT）法,研究了河西走廊荒漠边缘不同建植年限梭梭植被区固定沙丘土壤结皮斥水性的时间演变规律及影响因素。结果表明：建植20 a后,丘顶与丘间地土壤结皮出现显著斥水性（WDPT>5 s）,而丘坡土壤结皮无斥水性（WDPT<5 s）。丘顶土壤结皮斥水性与总有机碳、微生物量碳、有机碳C-H组分以及电导率显著相关（P<0.01）;而丘间地土壤结皮斥水性与黏、粉粒含量显著相关（P<0.01）。植被形成的“碳岛”及“盐岛”效应是丘顶土壤结皮斥水性形成的主要原因,而黏、粉粒在土壤表层的积聚是丘间地土壤结皮斥水性形成的主要原因。