Structural and nutritional heterogeneity of riparian vegetation in Patagonia (Argentina) in relation to seasonal grazing by sheep

Riparian meadows of southern Patagonia are temporally and spatially heterogeneous habitats. They are thought to play a key role in regulating the dynamics of arid grazed steppes of the region. We conducted a 2-year study with sheep to determine the influence of two grazing conditions (deferment and spring grazing) on structural and nutritional parameters of three vegetation types apparently associated with a soil moisture gradient in a riparian meadow in southern Santa Cruz (Argentina). Spring deferment allowed forage accumulation in very moist and intermediate vegetation types by the beginning of the summer and had no detrimental impact on forage quality. Deferment had no effect on biomass accumulation of the drier vegetation type at our study site. Structural attributes of the vegetation appeared to affect spatial grazing patterns of sheep. Soil-related patchiness, rather than sheep grazing, appeared to control vegetation structure and nutritional value. The proportion of more mesic plant communities in riparian habitats of Patagonia could determine the outcome of plant–animal interactions in these systems. Practical recommendations of grazing capacity in paddocks, or more theoretical considerations of ecosystem dynamics of the Patagonian steppes need to explicitly consider the internal patchiness of riparian habitats.     

Structural and nutritional heterogeneity of riparian vegetation in Patagonia (Argentina) in relation to seasonal grazing by sheep

Riparian meadows of southern Patagonia are temporally and spatially heterogeneous habitats. They are thought to play a key role in regulating the dynamics of arid grazed steppes of the region. We conducted a 2-year study with sheep to determine the influence of two grazing conditions (deferment and spring grazing) on structural and nutritional parameters of three vegetation types apparently associated with a soil moisture gradient in a riparian meadow in southern Santa Cruz (Argentina). Spring deferment allowed forage accumulation in very moist and intermediate vegetation types by the beginning of the summer and had no detrimental impact on forage quality. Deferment had no effect on biomass accumulation of the drier vegetation type at our study site. Structural attributes of the vegetation appeared to affect spatial grazing patterns of sheep. Soil-related patchiness, rather than sheep grazing, appeared to control vegetation structure and nutritional value. The proportion of more mesic plant communities in riparian habitats of Patagonia could determine the outcome of plant–animal interactions in these systems. Practical recommendations of grazing capacity in paddocks, or more theoretical considerations of ecosystem dynamics of the Patagonian steppes need to explicitly consider the internal patchiness of riparian habitats.     

Grazing effects on patchy dryland vegetation in northern Patagonia

In this study the spatial patterns and dynamics of vegetation patches along a grazing gradient in the steppe ofLarrea divaricataandStipaspp. in NE Patagonia (Argentina) are described. A general effect of grazing is the reduction of total plant cover resulting from the decrease in cover of perennial grasses (Stipa speciosa, Poa ligularis, Stipa tenuis) and some tall shrubs (Chuquiraga hystrix, Bougainvillea spinosa, Lycium chilense). Dwarf shrubs (Nassauvia fueguianaandJunellia seriphioides) increase their cover under medium and/or high grazing pressures. Plant species are spatially grouped into patches which alternate with areas of bare soil. Eleven types of vegetation patch differing in the dominant plant functional type or species, floristic richness and size were identified with different relative frequency along the grazing gradient. Based on these results, it is postulated that grazing forces the replacement of large patches dominated by tall shrubs with high species richness, byLarrea divaricatapatches or small dwarf shrub patches with low species richness and the extinction of grass patches. This results from: (1) disruption of local balances of species deletions and additions; (2) fragmentation of large patches; and (3) formation of new vegetation patches. These changes lead to differing plant spatial organization and heterogeneity along the grazing gradient which may be described by characteristic arrays of vegetation patches.     

Impact of rainfall variability and grazing pressure on plant diversity in Mongolian grasslands

Climate and grazing are the main drivers of plant community composition and species richness in arid environments. This study aimed to examine the vegetation response to a spatial precipitation gradient, interannual rainfall variability, and grazing pressure in Mongolia. To examine the effect of a spatial precipitation gradient, we compared species richness among six sites. To investigate the effects of interannual rainfall variability and grazing pressure, we compared species richness for 2 years at two sites, in desert-steppe and steppe areas. The regional gradient in annual precipitation showed positive and negative relationships with grass richness and shrub richness, respectively, although total species richness did not vary significantly. The proportions of the different functional groups were affected by grazing pressure and rainfall variability in both zones. In the desert-steppe zone, species richness was lower in the drier year but did not vary with grazing pressure. In the steppe zone, species richness varied significantly with grazing pressure but did not vary between years. Precipitation would be more important than grazing pressure on vegetation changes in drier areas with high rainfall variability.     

科尔沁沙地草地植被对围封和放牧的响应

研究了科尔沁沙地疏林草地、针茅草原和草甸植被盖度、地上和地下生物量、物种多样性对围封和放牧的响应。结果表明：（1）围封与放牧草地的优势植物不同,围封草地植物群落优势植物为多年生禾本科植物,放牧草地中一年生植物和小半灌木优势明显。（2）围封和放牧草地的植物盖度、凋落物量、地上生物量和物种丰富度存在明显差异（P<0.05）;围封显著提高了植被盖度和地上生物量,由疏林草地、针茅草原到草甸,植物盖度和地上生物量逐渐增加,而3种草地植被凋落物量大小顺序为针茅草原> 疏林草地> 草甸;放牧条件下植物盖度、凋落物量和物种丰富度差异不显著（P>0.05）。（3）3种草地之间的地下生物量无显著差异（P>0.05）,但围封与放牧之间、不同土壤层次之间地下生物量存在明显差异（P<0.05）;围封显著提高草地的地下生物量（P<0.05）;草地地下生物量随着土壤深度表现出下降趋势（P<0.05）。长期放牧增加了草地一年生植物和小半灌木植物的优势,消除了不同草地之间植被盖度和物种丰富度的差异;而围封能提高草地多年生禾本科植物的优势、增加其物种丰富度,对于草地质量和植物多样性的恢复和保育具有积极作用。     

北方草原区主要群落类型表土花粉分析

建立花粉与现代植被的关系是利用地层沉积物中的花粉记录恢复古植被、古气候研究的基础工作之一。中国北方草原区39个表土花粉研究表明,蒿属、藜科是草原区花粉组合的主要成分,以禾本科为建群种的草原群落花粉组合以蒿属花粉为主,禾本科花粉多低于10%。禾本科花粉的低代表性可能与其分蘖繁殖、花粉产量低有关。草原区表土花粉浓度与植被盖度相关不明显。典型草原与荒漠草原花粉组合存在着明显的区别,典型草原区藜科花粉百分比一般低于25%,荒漠草原区一般高于25%。DCA分析表明,草原区针叶树、莎草科、石竹科花粉代表较湿润的环境,蒿、藜科、锦鸡儿、柽柳、白刺花粉代表较干燥的环境,阔叶树、禾本科、菊科、毛茛科、蓼科、豆科、唇形科(百里香)、虎榛子+榛、蔷薇科花粉代表的环境介于前两组之间。DCA分析用于草原区花粉组合研究具有较好的环境指示意义。     

Long-term dynamics of a semiarid grass steppe under stochastic climate and different grazing regimes: A simulation analysis

We built a grid-based spatial explicit stochastic model that simulates grazing events and basic processes like seedling establishment, growth or mortality of the dominant species in the grass steppes of Patagonia. After evaluating the model with field data, we performed simulation experiments aimed to explore the interaction of precipitation and grazing regimes on vegetation dynamics. Grazing generated a reduction in tussock density which results in a decline in aboveground net primary production (ANPP). Both response variables presented a non-linear behavior including high temporal variability and delay effects, which may prolong for decades. There was a clear threshold in the response of the variables to stock density, though changes become evident only when a highly selective grazing scenario was used. Under high stock density conditions, precipitation use efficiency (PUE) was 82% lower than the values for non-grazed runs. The inter-annual variability of precipitation was more important than the grazing regime in explaining differences in tussock density. Simulation results highlight important issues regarding rangeland management: grazing regime might be as important as stocking density as a degradation agent, temporal lags might obscure degradation processes for decades, the definition of monitoring variables need to consider their response time constants.     

Cattle grazing effects on annual plants assemblages in the Central Monte Desert,Argentina

Plant assemblages' dynamics have been studied to evaluate the influence of different environmental factors. The aim of our work was to assess the effect of a disturbance in the form of livestock grazing on annual plants, in a South American desert. We tested the exclusion of cattle grazing by comparing the composition of annual plants in three major vegetation types within a MaB Reserve in the Monte Desert of Argentina, with those of an adjacent grazed field. Sampling was conducted in two consecutive years that differed in precipitation. We established three sampling sites within each vegetation type at the reserve and the grazed field. Transects were set to assess plant cover, abundance, and richness. Theoretically, changes in diversity are explained by changes in one of its components: species richness. Species richness of annual plants was not different between the grazed and ungrazed sites. However, plant cover and diversity were lower in grazed sites, whereas abundance increased. Owing to a strong species-specific effect, we propose that it is evenness the main parameter involved in diversity dynamics in the heterogeneous vegetation mosaic of the Monte desert. Finally, vegetation types (spatial heterogeneity) and rainfall regime (temporal heterogeneity) greatly interacted with grazing effects. We endorse the idea that rapid responses of annual plant assemblages to changes in rainfall conditions coupled with herbivore control, might result in a restoration pathway for degraded arid landscapes.     

Patterns of alien plant invasions in northwestern Patagonia, Argentina

Invasive plant species are a growing concern in many regions of the world, yet little information is available on the effects of habitat type and distance to urban settlements on the ability of alien plants to become established. We studied plant communities from the arid steppes to the Andean forest within two national parks and surroundings, in northwestern Patagonia, Argentina. Alien and native vascular plant species were sampled using 90 × 100 m² plots and analyzed in relation to both climatic and environmental variables and distance to urban centres. We also compared life-forms of alien and native species. Precipitation and distance to an urban centre were important determinants of alien species community richness and composition, and shrubland sites had significantly more alien species than forest or steppe sites. Alien flora (15% of the species) was composed of a higher proportion of annuals and biennials than native flora. Our results suggest that precipitation, distance to source population, and anthropogenic disturbance influence the invasion process in this region, together with the availability of open niches. This work stresses the importance of monitoring invasive species in natural reserves, and of considering habitat types as well as idiosyncratic characteristics of the non-natives for developing management strategies.     

Effects of Grazing on the Grassland Vegetation Community Characteristics in Inner Mongolia

The continuous increase of livestock production in Inner Mongolia has caused severe degradation of the grassland ecosystems in recent years. Previous grazing experiments have shown a wide range of vegetation responses between the biome types on a global scale, but there is still a lack of sufficient studies to discern the relative responses of a given biome type. We conducted a meta-analysis of vegetation coverage (VC), plant density (PD), total biomass (TB), above-ground biomass (AGB), under-ground biomass (UGB) and Shannon-Weaver Index (SI) in different grassland types in Inner Mongolia obtained under conditions of different grazing intensities and durations. The results showed that grazing decreased VC, TB, AGB, UGB, and PD significantly. Compared to the global and national average values, the negative effects of grazing to steppe biomass in Inner Mongolia were higher than that on the global scale, while less pronounced than that in China. TB of the meadow steppe in Inner Mongolia increased by 40% under moderate grazing intensity and duration because of compensatory growth. SI of the desert and meadow steppe showed negative linear relationships with the grazing intensity in Inner Mongolia. The percentage changes in AGB, PD, and SI to grazing showed quadratic relationships with the mean annual temperature of the experimental year. With increasing mean annual precipitation, the negative effects of grazing on UGB and SI first decreased and then increased, with that of VC and grazing showing a cubic relationship.     

Is the analysis of plant community structure better than common species-diversity indices for assessing the effects of livestock grazing on a Mediterranean arid ecosystem?

Grazing in arid Mediterranean ecosystems brings about changes in species diversity, but the best way to measure such changes is unclear. In this study we compared various methods in order to identify indicators that might be useful for the management and conservation of grazed arid Mediterranean ecosystems.Changes in community structure and composition were compared along a previously studied grazing gradient in Cabo de Gata-Nijar Natural Park. Plant diversity was measured by calculating diversity (Shannon information index), evenness (Pielou index) and richness (species number). Rarefaction curves were used to measure plant species richness and the results were compared with traditional richness values. Community structure was measured as the percentage of bare ground and the coverage of overall perennial grasses, Stipa tenacissima L., perennial shrubs and annual plants. Our results showed that the proportion of bare ground, the cover of perennial shrubs and the relative abundance of S. tenacissima are good indicators of grazing effects on vegetation. Overall plant community structure was more sensitive to the grazing gradient than were the diversity indices. Finally, the adequacy of standardized methods for comparing species richness (i.e. rarefaction curves) along a grazing gradient are discussed.     

退化沙质草地土壤种子库和地上植被对封育的响应机理

从植物种和植物功能类群两个水平系统探讨退化沙质草地土壤种子库和地上植被对短期封育管理响应机理的研究尚不多。以内蒙古东部的科尔沁沙地退化沙质草地为研究对象,采取随机样方法对禁牧6 a的封育区及与其相邻的自由放牧草地的土壤种子库和地上植物群落的物种密度进行了比较研究,草本植物群落被区分为一年生禾本科植物、非禾本科一年生植物和多年生植物3个主要功能类群。利用测定数据,分析研究了短期禁牧封育对土壤种子库和地上群落不同植物种和各植物功能类群的生态恢复作用机理。主要研究结果是：①短期封育显著促进了退化沙质草地植被的生态恢复,但发现土壤种子库和地上植被对禁牧的响应模式和机理存在明显差异,短期禁牧使土壤种子库中所有3个植物功能类群的密度均得到显著提高,而仅提高了地上植被一年生禾本科植物和多年生植物两个类群的密度;相反,禁牧显著提高了地上植被所有3个功能类群的物种丰富度和多样性,仅提高了土壤种子库中多年生植物类群的物种多样性。②短期禁牧显著增加了地上植物群落组成中可食和优良牧草的比例,提高了草地的经济利用价值。主要结论是：短期禁牧封育是一种经济有效的提高退化沙质草地植被多样性、生产力及对植物群落物种组成和草地品质进行优化调控和改良的重要草地管理手段。研究建议,今后应重点开展不同禁牧时间下对退化沙质草地土壤种子库和地上植被多样性、生产力及其物种组成变化影响的长期生态学控制实验研究。     

科尔沁沙质草地植物群落的放牧退化及其自然恢复过程

 为了了解不同放牧强度对我国北方沙质草地的影响及其封禁停牧后草地植被的自然恢复过程,我们于1992—2006年在科尔沁沙地开展了草地放牧和封育试验,分析研究了人类放牧活动对草地植被的影响,比较了不同程度退化草地恢复速度及其所受气候变化影响。研究结果表明:①在科尔沁沙地,随着放牧强度的增加,草地植被受放牧影响的程度会加重,但只有持续重牧才会导致植被的严重退化,而且持续重牧对植被盖度、高度和草地生产力的危害程度要大于对物种丰富度和多样性的危害程度;中度放牧可以导致植被盖度、高度、生物量的轻度下降,但对物种丰富度、多样性没有不良影响;对于退化草地而言,持续轻牧可促进植被的有效恢复;②封禁停牧以后,草地原受干扰程度不同,其恢复的相对速度也不同,表现为原草地退化越严重,其恢复速度相对越快,而且是植被高度、盖度、生物量的恢复速度要大于物种丰富度、多样性和频度的恢复速度;③草地植被的恢复演替明显受到气候变化的影响,暖湿气候有利于植被的恢复,暖干气候对于植被的恢复不利。     

Small-scale vegetation dynamics in semi-arid steppe in Inner Mongolia

Small-scale vegetation dynamics were followed for ecotones and in uniform stands inArtemisia-dominated steppe vegetation under grazing and when recovering from heavy grazing. Species composition was followed annually for 5 years in 1 m²and 0·25 m²plots for (1) presence–absence; (2) density; and (3) biomass.More rapid vegetation dynamics, in terms of change of type of vegetation and distance moved in DCA species space between sampling occasions, were observed in smaller plots and in early seral stages, where species were few and alpha diversity low. Only the plots recently protected from grazing showed a directional vegetation change; those protected for more than 3 years and those under continued grazing showed, at this scale, non-directional dynamics.     

Assessment of grazing effect on sheep fescue (Festuca valesiaca) dominated steppe rangelands, in the semi-arid Central Anatolian region of Turkey

In the semi-arid steppe rangelands of Central Turkey, Festuca valesiaca and Thymus sipyleus ssp rosulans have become the dominant species on degraded pastures. We hypothesized that decreases in species richness and abundance are correlated with increasing prevalence of these two species. Therefore, our objectives were to determine whether there are patterns in examined vegetation; how dominant species contribute to these patterns; and how patterns differ between grazed and ungrazed vegetation. We determined that protection from grazing increased species richness. Grazing significantly changed composition through decreasing total plant, forb, grass and F. valesiaca covers, while substantially increasing T. sipyleus cover. Topography, soil and grazing appear to impact the dominance of plant communities where F. valesiaca and T. sipyleus prevail. These two dominant species had a significant effect in shaping vegetation patterns. Based on regression analysis, alterations in species richness with changes in cover of forbs and shrubs were evident, and spatial heterogeneity of F. valesiaca and T. sipyleus indicated unstable vegetative patterns in heavily grazed pastures and successional changes in protected pastures. Our study results identify F. valesiaca and T. sipyleus as indicator species of vegetation suppression in condition assessments of degraded steppe rangelands.     

Soil water condition and small mammal spatial distribution in Inner Mongolian steppes, China

We studied the roles of soil moisture contents and vegetation structure in the spatial distribution of small mammals in the typical steppes of Inner Mongolia, China, using logistic and linear regressions of a data set collected in a 6-year study. Our results indicated that soil moisture contents remained in the most parsimonious models for Spermophilus dauricus, Cricetulus barabensis, Microtus maximowiczii, M. gregalis, and Ochotona daurica. The relative abundance of C. barabensis, M. maximowiczii, and O. daurica was inversely related to soil moisture contents, while that of M. gregalis and S. dauricus was positively related to soil moisture contents in logistic regressions. Linear regression analyses showed that soil moisture contents and the number of small mammal species were inversely related. The negative effects of wet soil were consistent at both small mammal population and community levels in the semi-arid steppes. Above-ground plant biomass and plant coverage also affected the spatial distribution of small mammals in the typical steppe of Inner Mongolia.     

科尔沁沙地植被恢复过程中群落组成及多样性演变特征

流动沙丘、固定沙丘和沙质草地均是科尔沁沙地土地沙化的产物,植被恢复重建则是该区域流沙治理的主要措施,理解植被恢复过程中群落组成及植物多样性演变特征有利于沙地植被恢复和生态重建.本研究基于连续15年对科尔沁沙地流动沙丘、固定沙丘和沙质草地长期植物群落结构调查数据,分析了 3种沙地类型植物群落年际变化特征及不同生活型植物年...     

Plant communities of central Tibetan pastures in the Alpine Steppe/Kobresia pygmaea ecotone

Eleven plant communities of the central Tibetan ecotone (31°20′-33°00′N/89°00′-92°10′E) between the Kobresia pygmaea grazing pastures of the eastern highlands and the open short grass steppe with cushion plants (“Alpine Steppe”) of the north-western highlands have been described on the basis of 189 plot-based floristically complete vegetation records. In parallel, remote sensing techniques distinguish four main land-use cover types. Our hypotheses are: (1) The plant communities of the central Tibetan highlands are grazing-adapted and resilient to degradation. (2) In contrast to grazing resilient plant functional types, the turf cover of the K. pygmaea pastures and the Kobresia schoenoides wetlands is degradable through desiccation, periglacial processes, soil-dwelling small mammals and livestock. Five grazing-related plant functional traits are introduced. Grazing tolerance is the prevalent functional type. Species with no specific protection against grazing make up not more than 6% of the total cover. Unpalatable plants cover up to 8%. Only the azonal K. schoenoides swamps - the indispensable winter grazing reserve - have been widely degraded and depleted by 75%, being replaced by Carex sagaensis grazing pastures. It can be foreseen that governmental policy of sedentarisation of nomads will lead to reduced grazing mobility and degradation of winter grazing reserves.     

Grazing response in the vegetation communities of the Kamiesberg, South Africa: Adopting a plant functional type approach

Potential grazing impacts associated with the sustained heavy grazing of a communal rangeland are explored through a plant functional type approach. An informative typology is derived that shows grazing responses. An examination of plant functional types within growth forms does not provide any additional insights, whereas looking at traits alone does demonstrate a grazing response with increases in the traits of large basal and erect leafy growth forms and losses in the traits of medium height, simple leaves and few flowers. There is also evidence of a biogeographic shift, with an increase in a succulent-dominated plant functional type at higher altitudes on the heavily grazed communal rangeland. This is attributed to the fact that in adopting the proposed universal language advocated by plant functional type practitioners some of the subtleties of this particular vegetation type are lost. For example growth form height classes are too coarse for the shorter vegetation of the Succulent Karoo. In the resultant ‘fuzzy’ groups, responses are lost. While the method adopted is informative a stronger case is made for a more simple growth form analysis traditionally adopted in rangeland analysis.     

Changes in soil nutrients, vegetation structure and herbaceous biomass in response to grazing in a semi-arid savanna of Ethiopia

The effect of grazing was studied on vegetation structure, herbaceous biomass, basal and bare ground covers, together with soil nutrient concentrations in two locations in an Ethiopian semi-arid savanna. The lightly grazed sites had significantly higher herbaceous diversity, total abundance, basal cover and aboveground biomass, and a lower percentage of bare ground compared with the heavy grazed sites. Grazing pressure had no effect on the density and number of woody species as well as on the proportion of encroaching woody species. The light grazing sites had higher organic carbon, phosphorus and exchangeable bases, and therefore a higher pH and higher electrical conductance, indicating an improved soil nutrient status compared with heavy grazing sites, mainly attributed to the higher basal cover and standing biomass at light grazed sites, and the export of nutrients through grazing and dung collection from the heavily grazed sites. There were significantly higher soil nutrients, species diversity, aboveground biomass and basal cover in the light grazing sites compared with heavy grazing sites. We concluded that changes in herbaceous vegetation, standing biomass and soil compositions are caused by interactions between grazing, soil and vegetation, and these interactions determine the transitions of semi-arid savannas.