浙东中新世Cinnamomum cf.bejolghota微细构造及其古环境意义

化石采自浙江天台嵊县组.该地层含丰富被子植物压型化石及少量裸子植物化石.利用叶结构分析法,依据叶脉、叶尖、叶基等稳定性特征,对一种樟科植物化石的宏观形态进行了详细的研究;结合表皮特征分析,发现该化石与现生Cinnamomum bejolghota(Buch-Ham.)Sweet在叶形、叶脉特征以及表皮脉络细胞特征、气孔器类型和毛基特征等方面均很相似,因而将当前化石定名为Cinnamomum cf.bejolghota(钝叶桂相似种).通过分析化石钝叶桂的角质层微细构造特征,结合现生钝叶桂的分布范围和生存环境恢复了浙东地区中新世的古气候和古环境特征,推测当时的气候比现在更为炎热多雨,同时认为钝叶桂从中新世至今有南迁的趋势.     

浙江天台中新统润楠属化石及其古环境指示

对浙江天台中新统下南山组6块叶片压型化石——天台润楠Machilus tiantaiensis Ding S. T. et Sun B. N. sp. nov.,并对其叶结构和表皮构造特征进行了详细的研究.其主要特征为:叶椭圆形,全缘,顶端渐尖,基部楔形,侧脉羽状;表皮细胞垂周壁浅波状,单细胞毛基多集中于脉络区;气孔下生式,气孔器短平列型,副卫细胞1~2个.通过与现生植物进行对比分析,发现当前标本的叶结构和表皮构造特征与樟科润楠属(Machilus)十分一致.利用PAST(palaeontological statistics)软件对我国润楠属68种植物叶片的形态特征进行聚类分析,结果表明当前化石与M. pingii Cheng ex Yang最为接近;角质层分析也表明二者的表皮构造最为相似.利用化石与其现存最近亲缘种M. pingii的气孔比率,恢复了天台晚中新世的古大气CO₂体积分数为407.9×10-6.此外,依据现生润楠属主要分布于亚洲东南部和南部的热带、亚热带地区,推断天台地区中新世的古气候较为温暖湿润.      

云南腾冲上新统垂丝紫荆(比较种)Cerciscf.racemosa Oliv.表皮微细构造研究

本文研究了产于滇西腾冲上新世芒棒组一种被子植物叶型化石垂丝紫荆(比较种)Cerciscf.racemosaOliv.,归于豆科紫荆属。揭示了该种叶表皮构造特征:上表皮无气孔器,表皮细胞为多边形,垂周壁平直;下表皮具环列型气孔器,表皮细胞为多边形,垂周壁平直;上下表皮有相同类型的毛基,毛基的根部由多细胞组成,毛基部的细胞特化成辐射状。该化石表皮微细构造的分析为恢复腾冲上新世古环境提供了古植物证据。     

云南腾冲上新统垂丝紫荆（比较种）Cercis cf．racemosa Oliv．表皮微细构造研究

本文研究了产于滇西腾冲上新世芒棒组一种被子植物叶型化石垂丝紫荆（比较种）Cercis cf．racemosa Oliv．，归于豆科紫荆属。揭示了该种叶表皮构造特征：上表皮无气孔器，表皮细胞为多边形，垂周壁平直；下表皮具环列型气孔器，表皮细胞为多边形，垂周壁平直；上下表皮有相同类型的毛基，毛基的根部由多细胞组成，毛基部的细胞特化成辐射状。该化石表皮微细构造的分析为恢复腾冲上新世古环境提供了古植物证据。     

浙东中新世Betula mioluminifera Hu et Chaney的发现及古气候重建

随着古气候和古环境研究的日渐深入,“特有种气候分析法”成为新生代定量重建古气候研究的重要方法之一。使用这一方法分析了浙江东部天台中新统下南山组的古亮叶桦（Betula mioluminifera Hu et Chaney）压型化石,研究了其叶结构特征及表皮微细构造,定量推论浙江天台中新世时年平均温度为12.5～22.1℃,春季平均温度为13.0～23.1℃,较现在温暖。我国内蒙、吉林、山东、云南等地的新近系发现过古亮叶桦,分布的北界为42°N。其现存最近亲缘种亮叶桦（Betula luminifera）生活在101～119°E,23～34°N之间,海拔500～1800 m的山区,是我国特有树种。亮叶桦分布范围的缩小显示中新世一段时期我国气候逐渐变冷的趋势。     

辽宁阜新早白垩世几种松柏类化石微细构造及地质意义

采集了辽宁阜新盆地下白垩统的3种松柏类植物化石,分别为Cephalotaxopsis haizhouensis Shang,Elatocladus cf. dunnii Miller et Lapasha和Elatides harrisii Chow。对三者的表皮微细构造进行了详细研究,其中Cephalotaxopsis haizhouensis和 Elatocladus cf. dunnii的微细构造特征总体反映了一种温暖湿润的环境条件。根据它们的表皮构造和气孔带特征,与现生种 表皮微细构造做了对比分析,结果显示现生植物Cephalotaxus sinensis和Cephalotaxus fortunei可以作为化石植物Cephalotaxopsis haizhouensis和Elatocladus cf. dunnii的最近现存对应种,通过这两种化石植物与其最近现存对应种的气孔比率定量重建了阜 新盆地早白垩世的古大气CO2浓度。     

云南腾冲上新统Platycladus tengchongensis sp.nov.表皮构造与古CO_2恢复

在云南腾冲上新统芒棒组发现了角质层保存完好的侧柏化石。通过与已知化石种及最近亲缘种的宏观形态和微细构造特征对比,建立一新种Platycladus tengchongensis P.Deng et B.N.Sun sp.nov.。基于当前新种与现生种Platycladus orientalis和化石种Platycladus yunnanensis在宏观形态及角质层特征的相似性,统计了3个种的气孔密度和气孔指数,并用气孔比率法重建了云南腾冲上新世晚期和云南梁河晚中新世的古大气CO_2浓度分别为417.9×10~(-6)(386.5×10~(-6)~471.3×10~(-6))和409.9×10~(-6)(392.2×10~(-6)~433.3×10~(-6)),结果与Berner全球碳平衡模型中CO_2浓度变化曲线上的变化范围基本相符,故表明Platycladus tengchongensis和Platycladus yunnanensis叶片可以作为测量古大气CO_2浓度变化的生物指标。根据文中恢复的云南西部上新世晚期和晚中新世古大气CO_2浓度平均值均高于现在,推测新近纪时期云南西部的气候比现在温暖。综合云南腾冲地区已有的古气候研究与化石和现生叶片角质层特征的对比分析,认为云南腾冲上新世晚期气候较为温暖湿润。此外,文中化石的发现为侧柏属自然分布的研究提供了宝贵的材料。     

陇原化石珍品

<正>临夏鸵鸟学名:Struthio linxiaensis侯连海,周忠和,张福成,王赵2005词源:以化石产地甘肃临夏为种名。产地与层位:和政地区晚中新世柳树组分类位置:鸟纲鸵鸟目鸵鸟科鸵鸟属特征:鸵鸟是现生鸟类中体形最大的鸟,仅存非洲鸵鸟(Struthio     

云南临沧晚中新世梣属翅果化石及其古植物地理学意义

梣属植物是北半球古近纪和新近纪化石记录中最具代表性的种属之一。对采自云南临沧晚中新世的一种梣属翅果化石Fraxinus cf.honshuensis进行了鉴定和描述,并通过梣属12个翅果化石种和9个现生种的形态特征的研究,分析了该属翅果形态从始新世直至现代的演化趋势及风力传播作用对翅果形态的影响。结果表明:该属翅果在晚渐新世至中中新世时翅果的风力传播能力最强,传播距离最远,并且与化石记录具有较好的对应性。地史分布表明,该属植物最初可能通过白令陆桥和北大西洋陆桥由高纬度地区向低纬度地区迁徙,最终传播到亚热带和热带区域。     

现生河狸的起源

本文通过对来自欧洲、亚洲、北美洲的河狸属(Ｃａｓｔｏｒ)化石和现生种的头骨形态学之研究，认为河狸属的起源可上溯到中新世的Ｓｔｅｎｅｏｆｉｂｅｒ。现生欧洲河狸Ｃ．ｆｉｂｅｒ与中国化石河狸Ｃ．ａｎｄｅｒｓｓｏｎｉ有密切关系；而北美现生河狸Ｃ．ｃａｎａｄｅｎｓｉｓ则可能源于美国的化石河狸Ｃ．ｃａｌｉｆｏｒｎｉｃｕｓ。约六百多万年前，河狸属从欧亚大陆穿越白令陆桥向北美的扩张，导致了现生河狸两个种的出现。这六百多万年的隔离是造成两种现生河狸在基因库、生态、头骨形态方面分异的根本原因。     

Biomolecular Preservation of Miocene Fossil Liquidambar from Tiantai, China and Clarkia, USA: Implications for Palaeoenvironmental Study

Biomolecular characteristics of fossil Liquidambar from the Miocene Shengxian Formation, Tiantai, Zhejiang province in eastern China and Miocene Clarkia sediment, Idaho, U.S.A., are obtained by means of Pyrolysis-Gas chromatography-Mass spectrometry analysis (Py-GC-MS). Their modern equivalent (Liquidambar formosana) collected from Nanjing, eastern China, is also analyzed for comparison. The total ion chromatograms (TIC) of modern L. formosana leaves shows abundant pyrolysates of lignin, polysaccharide, aliphatic acid, and amino acid. However, only one polysaccharide molecule, i.e., levoglucosan, is retained in fossil Liquidambar leaves from the two Miocene sites with the Tiantai specimen possessing higher intensity. As expected, some lignin pyrolysates including benzene, phenol, guaiacol, vinylphenol and their derivatives, are detected in Miocene Tiantai L. miosinica. However, the lignin components with large molecular weight other than benzene and phenol compounds, are absent in Miocene Clarkia L. pachyphyllum. Abundant pristene and homologous series of n-alk-1-ene/n-alkane pairs with a long retention time are shown in pyrograms of both Tiantai and Clarkia fossils. Based upon our Py-GC-MS data, the molecular preservation in these Liquidambar leaf samples can be ranked as the following (from high to low quality respectively): modern sample >Tiantai sample >Clarkia sample. Vinyl phenol, an important compound, is present in modern and Tiantai Liquidambar leaves, while absent in Clarkia sample, implying that vinyl phenol may be sourced from lignin for Liquidambar leaves. Our data support the hypothesis of in situ polymerization for the origin of long-chain homologous series of n-alk-1-ene/n-alkane pairs as leaf alteration products. Our data have implications of the impact of leaf degradation toward biases for palaeoenvironmental reconstruction using stable carbon isotope of fossil plants, and suggest the importance of evaluating molecular preservation of fossil leaf tissues prior to the application of carbon isotope compositions of fossil leaves.     

Bamboo Leaf and Pollen Fossils from the Late Miocene of Eastern Zhejiang,China and their Phytogeological Significance

Fossil bamboo leaves and pollen from Upper Miocene deposits of the Shengxian Formation in Tiantai and Ninghai counties,eastern Zhejiang,China represent a rare record in Asia.The distinctive pseudopetiole and parallel venation of the leaf blades and the clearly thickened annulus of the pollen aperture place them in the subfamily Bambusoideae.Morphological analysis supports the determination of these fossil leaves as belonging to the genus Bambusium and two new species are described.Bambusium latipseudopetiolus Q.J.Wang et B.N.Sun sp.nov.has a distinctly wide pseudopetiole of 0.23 cm in width and 0.40 cm in length,several vascular bundles on parallel veins,and 5–8 lateral veins on both sides of the midrib.Bambusium longipseudopetiolus Q.J.Wang et B.N.Sun sp.nov.has a distinctly long pseudopetiole of 0.60 cm in length and 0.10 cm in width,several vascular bundles on the pseudopetiole,and 6 lateral veins on both sides of the midrib.Pollen grains from the same layer provide additional evidence of bamboos.They are characterized by 32.0–51.0 μm in diameter,a round pore 3.0–4.0 μm in diameter with a broad thickened annulus 2.5–3.5 μm around,and several conspicuous secondary folds on the exine surface.The morphological analysis leads to their assignment to Graminidites bambusoides Stuchlik.These fossils are important for the study of bamboo phytogeography in China.They demonstrate that there were bamboos growing in southeastern China during the Late Miocene and that bamboos in Zhejiang begin to diversify no later than the Late Miocene.In combination with bamboo fossils from other places,it seems that bamboos had a wide distribution across southern China during the Miocene,ranging from southwestern Yun'nan to southeastern Zhejiang.     

浙江天台盆地白垩纪含恐龙骨骼及蛋化石地层

浙东天台盆地早白垩世晚期的阿普特（Aptian）期至晚白垩世早期的赛诺曼（Cenomanian）期天台群赖家组和赤城山组已发现数以千计的恐龙骨骼及蛋化石。通过对恐龙骨骼及蛋化石地层剖面实测,首次精确厘定了天台盆地恐龙动物群的出露层位。化石研究显示,恐龙生物群包括蜥脚类（sauropods）、鸭嘴龙类（hadrosaur...     

Paliurus (Paliureae,Rhamnaceae) from the Miocene of East China and its Macrofossil-based Phylogenetic and Phytogeographical History

The extant genus Paliurus Miller is divided into P. ramosissimus and P. spina-christi groups and is diagnostically characterized by distinctive orbicular-winged fruits with a trilocular （or often bilocular） ovary and basally triveined leaves with a subentire to serrate margin. Here, structurally preserved winged fruits and associated compressed leaves of Paliurus （Paliureae, Rhamnaceae） are reported from the Middle-Upper Miocene Shengxian Formation of Tiantai, Zhejiang, eastern China. The fruits can be identified as Paliurus microcarpa Xiang-Chuan Lisp. nov., the diagnostic characters of external morphology and anatomy of which differ from those of both extant and fossil generic members; the leaves can be designated as Paliurus sp. The species based on winged fruits, P. microcarpa, might represent an ancient intermediate link exclusively known across the P. ramosissimus and P. spina-christi groups, showing a closer affinity to the P. ramosissimus group based on the resemblant smaller fruits with a narrower wing, and to the P. spina-christi group based on the similar wing structure and texture, respectively. Morphological analyses of the present and previous fossil fruit records around the world indicate that diversification of Paliurus had begun in northern transpacific regions by the Late Eocene. The macrofossil records, i.e., fruits and leaves of Paliurus globally when projected on paleoenvironmental maps in view of changing continental positions through time （from Eocene to Miocene） display that the past distribution of the genus was much wider than the present and belongs to a tropical and warm temperate realm of the Northern Hemisphere. Therefore, the modern restricted disjunction in tropical and warm temperate regions of East Asia and South Europe （even eastwards West Asia） might represent glacial refugia for some of the generic ancient lineages to survive severe cold at least since the terminal Pliocene.     

浙东中新世菱属果实化石的数量分类

在浙东天台晚中新世嵊县组发现大量菱属果实化石。该化石保存完好,特征丰富。由于其种类多、性状冗杂不明显,菱属植物的鉴定与分类一直存在很大争议。本研究基于菱属果实化石性状特征多、便于测量记录的特点,通过聚类分析和主成分分析等数值统计方法对菱属果实进行鉴别、归类,寻找鉴定分类中的指导性性状。首先,利用上述方法对现生菱属果实进行数量分析,取得了较好结果。然后,采用同样方法对菱属果实化石进行研究,认为聚类分析和主成分分析法能够对化石种进行有效的鉴定分类,可以成功剔除冗余性状。最后,研究得出果体形状、是否有果冠、果冠是否外翻、顶部形态、肩角生长方向、腰角生长方向、腰角长度、瘤状退化腰角数目等特征是菱属果实化石鉴定的关键性状。基于以上性状,嵊县组菱属果实化石可以分为两个类群。     

A New Species of Tsuga (Pinaceae) based on Lignified Wood from the Late Miocene of Central Yunnan, China, and Its Paleoenvironmental Implications

A new species, Tsuga nanfengensis sp. nov. (Pinaceae), is described on the basis of lignified fossil wood from the late Miocene of the Xianfeng Basin, central Yunnan, southwestern China. Detailed observation of the fossil wood specimens show the following characteristics: distinct growth rings, absence of resin canals, uniseriate bordered pits in the radial wall of tracheids, ray tracheids and piceoid and cupressoid cross-field pits. These features indicate similarities to the wood of extant Tsuga canadensis, T. chinensis, and T. dumosa. According to the fossil record, Tsuga was present in Xundian County during the Miocene. Today Tsuga is drought intolerant, preferring wet conditions with no extant species growing naturally in Xundian County. The presence of Tsuga in the Miocene of Xundian County indicates a humid climate consistent with previous palaeoclimatic reconstructions showing a wetter and probably shorter dry season in the Miocene, relative to the present day. Therefore, the change in the local climate such as increasing aridity through the Miocene might explain the local extinction of Tsuga from central Yunnan.     

从浙江天台白垩纪蛋化石复原恐龙类群

浙江天台盆地白垩纪地层中,已发现了甲龙类及慢龙类等恐龙骨骼化石及数以千计的蛋化石,其中许多为恐龙蛋化石。根据国内外相同类型的蛋化石中迄今已发现的恐龙胚胎化石研究成果,可大致确定其中一部分蛋化石分别是由鸭嘴龙类、巨龙类、慢龙类、盗蛋龙类、伤齿龙类和暴龙类等恐龙所产。并发现鸭嘴龙类、巨龙类和慢龙类的圆形蛋、副圆形蛋在数量上占绝大多数,盗蛋龙的长形蛋和伤齿龙的棱柱形蛋较少,暴龙类的巨型长形蛋最少,从而反映了他们各自在生态金字塔中的位置。     

华东白垩纪鸭嘴龙类恐龙蛋化石的发现

浙江天台白垩纪地层中最常见的蛋化石类型之一是赤城山组地层（锆石U-Pb法同位素年龄约92Ma）的树枝树枝蛋（Dendroolithus dendriticus）。对比河南西峡含恐龙胚胎的蛋化石,确定其为鸭嘴龙类（Hadrosaurs）恐龙所产。鸭嘴龙类恐龙在系统分类上归鸟脚亚目（Ornithopoda Marsh,1881）鸭嘴龙超科（Hadrosauroidea Cope,1869）鸭嘴龙科（Hadrosauridae Cope,1869）,建议树枝蛋化石也相应归入圆形蛋化石亚目（Spheroolithineae账oosubord．nov．）树枝蛋化石超科（Dendroolithoidea oosuperfam．nov．）树枝蛋化石科（Dendroolithidae Zhaoand Li 1988）,以便对应研究。从已知的鸭嘴龙类恐龙产蛋尺寸推算,浙江天台的树枝树枝蛋（Dendroolithus dendriticus）为身长不超过7m的恐龙所产,个体大小类似东北一蒙古发现的萨哈林日本龙（Nipponosaurus sachaIinensis）、李氏黑龙江龙（Amurosaurus riabinini）和姜氏巴克龙（Bactrosaurus johnsoni）。该蛋在巢穴里,无序放置,至少3层,常超过20枚,蛋壳具韧性,重力作用下呈扁球形。