The distribution of calcareous nannofossils in the late Santonian–Late Maastrichtian deposits in the southwest of Iran (Khuzestan Province)

The first data on the distribution of calcareous nannofossils in the Behbehan section, the Kuh-e-Rish, are considered. According to the distribution of nannofossils, the Upper Cretaceous deposits of the section are subdivided into nine biostratigraphic zones. CC17 (Calculites obscurus zone) indicate the Late Santonian. Biozones CC18 (Aspidolithus parcus zone), CC19 (Calculites ovalis zone), CC20 (Ceratolithoides aculeus zone), CC21 (Quadrum sissinghii zone), and CC22 (Quadrum trifidum zone) represent the Campanian. Biozone CC23 (Tranolithus phacelosus zone) indicate the Late Campanian–Early Maastrichtian. Biozones CC24 (Reinhardtites levis zone) and CC25 (Arkhangelskiella cymbiformis zone) suggest the Middle and Late Maastrichtian, respectively. In the late Late Maastrichtian, due to decreasing in water depth at the study area, Nephrolithus frequens zone (CC26) defined in Tethysian domain was not recognized. The boundary between Gurpi–Pabdeh Formations represented a non-depositional period from the late Late Maastrichtian to the end of Early Paleocene. Also, it seems that predominant conditions of the sedimentary environment of Neotethys basin with the presence of index species calcareous nannofossils specified, which itself indicates that the warm climate and high depth of the basin in Late Santonian to Late Maastrichtian, in low latitudes has been prevalent.     

The fossil record (calcareous nannofossils and foraminifers) in single intervals of the Upper Cretaceous flysch sequence of the West Carpathians, Czechoslovakia

Studies of calcareous nannofossils and microfossils and their distribution in different intervals of the flysch rhythms in the Kloko ník brook in the Bílé Karpaty Unit of the Magura Group of nappes in the West Carpathians gave the following results. The highest species diversity of calcareous nannofossils was found in the lower parts of the Bouma Te hemipelagic intervals. The Campanian marker species Ceratolithoides aculeus and Aspidolithus parcus were found in practically all layers studied. A stratigraphically important foraminiferal fauna was obtained from the upper parts of the Bouma Te intervals, including the Campanian-Maastrichtian planktic foraminifers Globotruncana arca and Globotruncanita stuartiformis. In the non-calcareous pelagites, agglutinated species with a range from the Late Cretaceous to the Paleogene dominate.     

A review of the Upper Campanian and Maastrichtian belemnite biostratigraphy of Europe

The conventional belemnite zonation of northwest Europe includes three Upper CampanianBelemnitellazones, two Lower MaastrichtianBelemnellazones, in addition to the lower Upper MaastrichtianBelemnitella juniorZone and upper Upper MaastrichtianBelemnella casimirovensisZone. These zones are critically assessed. Recent belemnite subdivisions of the Upper Campanian and Lower Maastrichtian are reviewed. The Upper Campanian of Norfolk has been subdivided into seven informalBelemnitellazones and subzones, and the Lower Maastrichtian of northwest Germany into six formalBelemnellazones. The timespan of the Early Maastrichtian zones is estimated and the duration varies from 0.12–0.60 Ma. It is shown that the base of the basal MaastrichtianBelemnella lanceolataZone is slightly diachronous, and the base of theB. casimirovensisZone is highly diachronous across Europe.     

Late Maastrichtian litho- and ecocycles from hemipelagic deposits of eastern Sinai, Egypt

A biostratigraphical and palaeoecological survey employing calcareous nannofossils and planktonic and benthonic foraminifera has been carried out in four sections of hemipelagic marls and chalks of the Late Maastrichtian Abathomphalus mayaroensis Zone of eastern Sinai, in order to evaluate the mechanisms controlling the composition of the well preserved microfauna and nannoflora.The Abathomphalus mayaroensis Zone in eastern Sinai can be easily identified by the wide occurrence of the index fossil A. mayaroensis and can be further subdivided by the first occurrences of Plummerita reicheli (ex. P. hantkeninoides) and Micula prinsii. Microfossil abundances and lithologies are characterised by pronounced repetitive distribution patterns. These include low and high frequency fluctuations of the planktonic/benthonic (P/B) foraminiferal ratio, repetitive changes in the abundance of calcareous nannofossils and benthonic foraminifera, as well as the development of chalk-marl couplets and thinning upward chalk packets. both microfossil distribution patterns and the occurrence of rhythmites are attributed to changes in primary palaeoproductivity. Semiquantitative investigations of calcareous nannofossils and a few selected benthonic foraminifera yield evidence of the presence of high (HP) and low (LP) productivity assemblages.The interpreted HP assemblage is dominated by Glaukolithus diplogrammus, Manvitella pemmatoidea, Microrhabdulus decoratus and Micula murus and the benthonic foraminifera Neoflabellina jarvisi; the LP assemblage is characterised by Lithraphidites quadratus and Bolivinoides draco. However, further quantitative studies are necessary to reconstruct the exact composition of these assemblages and to explain deviatory developments. The chalk-marl couplets, thinning-upward chalk packets and the high frequency P/B patterns are interpreted to reflect productivity changes related to orbital forcing. These hemipelagites were deposited during the latest phase of the southern Tethyan upwelling system, which was active from the Santonian to the Late Maastrichtian with a peak in the Campanian. Termination of upwelling just before the K/T boundary also provides a good explanation for the change towards a palaeobathymetric control on foraminiferal distribution, as observed for the Palaeocene of central east Sinai.     

An integrated study of upper Campanian-lower Maastrichtian carbon isotopes and calcareous plankton biostratigraphy of the Kurdistan Region,northeastern Iraq

New carbon (δ¹³C) isotope records calibrated by planktonic bioevents provide general support for a late Campanian age assignment of the Shiranish Formation (Fm.) and its boundaries in the Dokan section (NE Iraq). The Shiranish Fm. is characterised at the base by a mid-Campanian unconformity as can be interpreted by absences of nannofossil zones CC20-21. The Shiranish Fm. then spans nannofossil biozones CC22-CC23a (UC15d-e^TP to UC16a^TP). Results obtained on carbon isotopes suggest that diagenesis affected and compromised a few carbonate samples in the uppermost 50 m of the section. However, once these samples are discarded, pristine trends suggest that the top of the section records a negative carbon isotope excursion that is interpreted as CMBa-c events that straddle the Campanian–Maastrichtian boundary. This interpretation is supported by the lowermost occurrence of planktic foraminifers Rugoglobigerina scotti and Contusotruncana contusa some 30 m above the base of the negative excursion and 10 m below a positive excursion identified as the Maastrichtian M1+ event. Discrepancies in the stratigraphic range of several planktic foraminifer bioevents are highlighted and advocate for the need of many more integrated records of planktic foraminifer and nannofossil biostratigraphy alongside carbon isotope stratigraphy in the eastern Tethys in order to improve regional and global schemes.     

Evaluation of Cretaceous–Paleogene boundary based on calcareous nannofossils in section of Pol Dokhtar, Lorestan, southwestern Iran

The Pol Dokhtar section of southern Lorestan, faulted Zagros range of southwestern Iran, contains one of the most complete Early Campanian to Danian sequences. The lack of a good fundamental paleontological study is a strong motivation for investigating calcareous nannofossils in southwestern Iran. The majority of the section is made of shale, marl, and partly of marly limestone and clay limestone, respectively. As a result of this study, 24 genera and 45 species of nannofossils have been identified and presented for the first time. This confirms the existence of biozone CC18 of zonation scheme of Sissingh (Geologie en Minjbouw 56:37–65, 1977) to NP1 of zonation of Martini, which suggests the age of Early Campanian to Danian. All Early Campanian to Danian calcareous nannofossil biozones from CC18 (equivalent to the Aspidolithus parcus zone) to NP1 (equivalent to the Markalius inversus zone) are discussed. Also, the zonal subdivision of this section based on calcareous nannofossils has shown continuity in Cretaceous/Paleocene boundary in south part of Lorestan Province. We can also learn about the predominant conditions of the studied sedimentary basin that was in fact part of the Neotethys basin with the existence of indexed species calcareous nannofossils that indicate warm climate and high water depths of the basin in low latitudes.     

Upper Maastrichtian ammonite biostratigraphy of the Gulf Coastal Plain (Mississippi Embayment,southern USA)

The Cretaceous outcrop belt of the Mississippi Embayment in the Gulf Coastal Plain (GCP) spans the Cretaceous/Paleogene (K/Pg) boundary. A detailed reconstruction of this time interval is critical for understanding the nature of biotic and environmental changes preceding the end-Cretaceous Mass Extinction event and for deciphering the likely extinction mechanism (i.e., bolide impact versus volcanism). Eight sections encompassing the K/Pg succession across the Mississippi Embayment were analyzed using biostratigraphic sampling of ammonites, dinoflagellates, and nannofossils. An upper Maastrichtian ammonite zonation is proposed as follows, from oldest to youngest: Discoscaphites conradi Zone, D. minardi Zone, and D. iris Zone. Our study documents that the ammonite zonation established in the Atlantic Coastal Plain (ACP) extends to the GCP. This zonation is integrated with nannofossil and dinoflagellate biostratigraphy to provide a framework to more accurately determine the age relationships in this region. We demonstrate that ammonites and dinoflagellates are more reliable stratigraphic indicators in this area than nannofossils because age-diagnostic nannofossils are not consistently present within the upper Maastrichtian in the GCP. This biostratigraphic framework has the potential to become a useful tool for correlation of strata both within the GCP and between the GCP, Western Interior, and ACP. The presence of the uppermost Maastrichtian ammonite D. iris, calcareous nannofossil Micula prinsii, and dinoflagellates Palynodinium grallator and Disphaerogena carposphaeropsis suggests that the K/Pg succession in the GCP is nearly complete. Consequently, the GCP is an excellent setting for investigating fine scale temporal changes across the K/Pg boundary and ultimately elucidating the mechanisms causing extinction.     

Geodynamic evolution of the Vulcan Sub‐basin,Timor Sea,northwest Australia: A pre‐compression New Guinea analogue?

The Vulcan Sub‐basin lies immediately inboard of the incipient arc‐continent collision in the Timor Sea and comprises part of the Bonaparte Basin system, the northernmost basin on Australia's North West Shelf. Given the high level of preservation of its extensional fabric, the region can provide important analogues for the likely pre‐orogeny architecture of New Guinea, which enables a better understanding of the onset of, and response to, orogenesis. Structural restoration of regional, depth‐converted 2–D seismic lines shows that although the Late Jurassic Swan Graben is significant and contains a thick source‐rock section, the principal phase of crustal extension took place in the Triassic to Middle Jurassic. Within the Vulcan Sub‐basin, the southern Tilted Fault Block Domain records ～10% Triassic to Middle Jurassic extension, whereas <5% upper crustal extension has been measured in the northern Hourglass Domain. Similarly, while Jurassic extension in the Tilted Fault Block Domain is both deep and focused, the Hourglass Domain is expressed as a broad sag to the northeast, indicating a strong underlying basement influence on compartmentalisation. The Vulcan Sub‐basin shows four principal stages of evolution: (i) regional, evenly spaced crustal faulting and subsidence in the Triassic ‐ Middle Jurassic; (ii) focused faulting in the Late Jurassic that created grabens with uplift of the shoulders; (iii) regional subsidence from the Middle Valanginian; and (iv) minor extensional and contractional reactivation in the Mio‐Pliocene. The measured brittle extension is much less than that suggested by modelling of lithospheric subsidence, which suggests long wavelength distribution of strain in the ductile lower crust, with upper crustal extension mainly focused along the continent‐ocean boundary. Along the North West Shelf and on a smaller scale within the Vulcan Sub‐basin per se, the obvious, basement‐involved, rectilinear compartments defined by prominent offsetting of both extensional fault systems and abyssal plains have important implications for the development of the New Guinea orogen. Similar scale compartments are recognised in New Guinea and display different structural styles and hydrocarbon prospectivity. The transfer zones separating the compartments are the sites of the major copper‐gold deposits in New Guinea. Using the Vulcan Sub‐basin ‐ Timor area as an analogue, it can be seen that an arc could originally collide with a promontory, such as what is now Timor, and reactivate the lineaments allowing local extension and mineralisation. In addition, interpretation of the structure of the New Guinea Fold Belt may be aided by considering the effects of compression on the geometry of the Vulcan Sub‐basin and of the similar Carnarvon Basin and adjacent extended and broken Exmouth Plateau.     

The Campanian-Maastrichtian stage boundary in the Aktulagai section (North Caspian depression)

Distribution of belemnites and benthic foraminifers in the Campanian-Maastrichtian boundary layers of the Aktulagai section, one of Upper Cretaceous reference sections in the east of the European paleobiogeographic region (EPR) is discussed. The base of Lanceolata Beds defined by A.D. Arkhangelsky in 1912 is well-substantiated biostratigraphic level corresponding to boundary between the Campanian and Maastrichtian stages. In spacious outcrops of Upper Cretaceous deposits in the Aktulagai Plateau (Aktyubinsk region, Kazakhstan Republic), “primitive Belemnella forms” (two rostra plates) appearing above that base distinctly replace the genus Belemnitella dominant in the Campanian. Seven successive zonal assemblages of benthic foraminifers (one plate) are established in the boundary interval. The Aktulagai reference section of Upper Cretaceous sediments can be used to trace the Campanian-Maastrichtian boundary from the eastern EPR to Boreal regions of Russia based on abundant micro-and nannofossils.     

Dentaries of a caenagnathid (Dinosauria: Theropoda) from the Nemegt Formation of the Gobi Desert in Mongolia

A symphyseal region of the fused dentaries of a caenagnathid theropod is described from the Upper Cretaceous Nemegt Formation at the Bugin Tsav locality in the Mongolian Gobi Desert. In contrast to the high diversity of Caenagnathidae in the upper Campanian to Maastrichtian in North America, only specimens of a single caenagnathid, Elmisaurus raurus, have been reported in the coeval strata in Asia. Although dentaries are commonly-found bones in the fossil record of Caenagnathidae, the present specimen is the first discovery of caenagnathid dentaries from the upper Campanian to Maastrichtian in Asia. The Nemegt Formation is unique for its diverse oviraptorosaurian fauna that includes both Caenagnathidae and Oviraptoridae as well as the non-caenagnathoid Avimimus portentosus. Hypothesized coexistence of eolian and fluvial environments in the Gobi Basin during the deposition of the Nemegt Formation might explain such co-occurrence of Caenagnathidae and Oviraptoridae.     

Calcareous nannofossils from chalky limestone of upper Abderaz Formation and lower part of Abtalkh Formation in the Kopet-Dogh range NE Iran

For the first time, the calcareous nannofossils of the chalky limestone of upper Abderaz Formation and lower part of Abtalkh Formation have been studied. In this study, 83 nannoplanktonic species of 45 genera were identified and presented. A biostratigraphic study of calcareous nannofossils from this section has allowed the recognition of five calcareous nannofossil biozones of Sissingh (Geol Mijnbouw 56:37–65, 1977) CC17–CC21. On the obtained calcareous nannofossils, the age of this section is Late Santonian/Early Campanian–Early Late Campanian.     

Oceanic events and biotic effects of the Cenomanian-Turonian anoxic event, Tarfaya Basin, Morocco

Profound biotic changes accompanied the late Cenomanian δ¹³C excursion and OAE2 in planktic foraminifera in the Tarfaya Basin of Morocco. Planktic foraminifera experienced a severe turnover, though no mass extinction, beginning with the rapid δ¹³C excursion and accelerating with the influx of oxic bottom waters during the first peak and trough of the excursion. Species extinctions equaled the number of evolving species, though only the disaster opportunists Guembelitria and Hedbergella thrived along with a low oxygen tolerant benthic assemblage. The succeeding δ¹³C plateau and organic-rich black shale deposition marks the anoxic event and maximum biotic stress accompanied by a prolonged drop in diversity to just two species, the dominant (80–90%) low oxygen tolerant Heterohelix moremani and surface dweller Hedbergella planispira. After the anoxic event other species returned, but remained rare and sporadically present well into the lower Turonian, whereas Heterohelix moremani remained the single dominant species. The OAE2 biotic turnover suggests that the stress to calcareous plankton was related to changes in the watermass stratification, intensity of upwelling, nutrient flux and oxic levels in the water column driven by changes in climate and oceanic circulation. Results presented here demonstrate a 4-stage pattern of biotic response to the onset, duration, and recovery of OAE2 that is observed widely across the Tethys and its bordering epicontinental seas.     

New Spanish Late Cretaceous eusuchian reveals the synchronic and sympatric presence of two allodaposuchids

The recently described clade Allodaposuchidae includes European eusuchian crocodyliforms restricted to the Late Cretaceous (Campanian and Maastrichtian). A new allodaposuchid crocodyliform is here described based on two specimens from the upper Campanian–lower Maastrichtian fossil site of Lo Hueco (Cuenca, Spain). This new taxon, Agaresuchus fontisensis gen. et sp. nov., is described by two complete skulls and a lower jaw associated with one of them. This new species can be distinguished unambiguously from Lohuecosuchus megadontos, the other allodaposuchid known from the same fossil site. The presence of two allodaposuchid crocodyliforms in Lo Hueco allows the recognition of the synchronic and sympatric existence of two representatives of this clade for the first time. The new genus Agaresuchus, comprises a previously described Iberian allodaposuchid species, “Allodaposuchus” subjuniperus, as Agaresuchus subjuniperus, new combination.     

Coniacian-Santonian Planktic Stratigraphy in central Tunisia

In this study, we describe a new stratigraphy of three exposed sections in central Tunisia, integrating Coniacian and Santonian planktic foraminifera and calcareous nannoplankton, supported by ammonite and inoceramid bioevents. In the three sections, the Coniacian/Santonian (C/S) boundary lies slightly above the lowest occurrence (LO) of the calcareous nannofossil Lucianorhabdus cayeuxii, which marks nannofossil Zone CC16 and matches well with the LO of the planktic foraminifera Dicarinella asymetrica. It also lies ∼4–7 m below the LO of the inoceramid Platyceramus cycloides and the ammonite Texanites (Texanites) sp. Comparing these marker C/S bioevents with the global stratotype section, the Olazagutia section (Spain) shows that the stratigraphic range of the bioevents are variable. This observation must be taken into consideration when making regional chronostratigraphic correlations.     

Tertiary stratigraphy of Western Australia

This paper is a summary of the present knowledge of the Tertiary stratigraphy of Western Australia. Also included is new information on the Cainozoic of the Carnarvon Basin, a result of petroleum exploration in the area.

Tertiary rocks formed during more than one cycle of deposition in three basins (Eucla, Perth, and Carnarvon), and also as thin units deposited in a single transgression along the south coast. The Tertiary stratigraphy of the Bonaparte Gulf Basin is not well known.

Drilling in the Eucla Basin has encountered up to 400 m of Tertiary in the south central part, with uniform thinning towards the margins. The section begins with a middle‐upper Eocene carbonate unit which represents the dominant event in the Tertiary sedimentation in this basin. More carbonates were deposited in the late Oligocene‐early Miocene and middle Miocene.

Along the south coast, the so‐called Bremer Basin, the Plantagenet Group (up to 100 m) of siltstone, sandstone, spongolite, and minor limestone, was deposited during the late Eocene.

The Perth Basin contains up to 700 m of Tertiary sediment, formed during at least two phases of sedimentation. The upper Paleocene‐lower Eocene Kings Park Formation consists of marine shale, sandstone, and minor limestone, with a thickness of up to 450 m. The Stark Bay Formation (200 m) includes limestone, dolomite, and chert formed during the early and middle Miocene. Events after deposition of the Stark Bay Formation are not well known.

The northern Carnarvon Basin and Northwest Shelf contain by far the most voluminous Tertiary sediment known from Western Australia: 3500 m is known from BOCAL's Scott Reef No. 1. A more usual maximum thickness is 2500 m. Most sediments were laid down in four episodes, separated by unconformities: late Paleocene‐early Eocene; middle‐late Eocene; late Oligocene‐middle Miocene; and late Miocene to Recent.

The Paleocene‐early Eocene cycle consists of about 100–200 m (up to 450 m in the north) of carbonate, shale, and marl of the Cardabia Group containing rich faunas of planktonic foraminifera.

The middle‐late Eocene sediments include diverse rock types. Marine and nonmarine sandstone formed in the Merlinleigh Trough. At the same time, the Giralia Calcarenite (fauna dominated by the large foraminifer Discocyclina) and unnamed, deeper water shale, marl, and carbonate (with rich planktonic foraminiferal faunas) formed in the ocean outside the embayment. Thickness is usually of the order of 100–200 m.

The main cycle of sedimentation is the late Oligocene‐middle Miocene, during which time the Cape Range Group of carbonates formed. This contains dominantly large foraminiferal faunas, of a wide variety of shallow‐water microfacies, but recent oil exploration farther offshore has recovered outer continental shelf facies with abundant planktonic foraminifera. A minor disconformity representing N7 and perhaps parts of N6 and N8 is now thought to be widespread within the Cape Range Group. The last part of this cycle resulted in sedimentation mainly of coarse calcareous marine sandstone (unnamed), and, in the Cape Range area, of the sandstone and calcareous conglomerate of the Pilgramunna Formation. Maximum thickness encountered in WAPET wells is 900 m.

After an unconformity representing almost all the late Miocene, sedimentation began again, forming an upper Miocene‐Recent carbonate unit which includes some excellent planktonic faunas. Thickness is up to 1100 m.

Thin marine sediments of the White Mountain Formation outcrop in the Bonaparte Gulf Basin. They contain some foraminifera and a Miocene age has been suggested.     

Stratigraphy,sedimentology and tectonic evolution of the Upper Cretaceous/Paleogene succession in north Eastern Desert,Egypt

The stratigraphy, sedimentology and syn-depositional tectonic events (SdTEs) of the Upper Cretaceous/Paleogene (K–P) succession at four localities in north Eastern Desert (NED) of Egypt have been studied. These localities are distributed from south-southwest to north-northeast at Gebel Millaha, at North Wadi Qena, at Wadi El Dakhal, and at Saint Paul Monastery. Lithostratigraphically, four rock units have been recorded: Sudr Formation (Campanian–Maastrichtian); Dakhla Formation (Danian–Selandian); Tarawan Formation (Selandian–Thanetian) and Esna Formation (Thanetian–Ypresian). These rock units are not completely represented all over the study area because some of them are absent at certain sites and others have variable thicknesses. Biostratigrapgically, 18 planktonic foraminiferal zones have been recorded. These are in stratigraphic order: Globotruncana ventricosa Zone (Campanian); Gansserina gansseri, Contusotruncana contusa, Recimguembelina fructicosa, Pseudohastigerina hariaensis, Pseudohastigerina palpebra and Plummerita hantkenenoides zones (Maastrichtian); Praemurica incostans, Praemurica uncinata, Morozovella angulata and Praemurica carinata/Igorina albeari zones (Danian); Igorina albeari, Globanomanlina pseudomenradii/Parasubbotina variospira, Acarinina subsphaerica, Acarinina soldadoensis/Globanomanlina pseudomenardii and Morozovella velascoensis zones (Selandian/Thantian); and Acarinina sibaiyaensis, Pseudohastigerina wilcoxensis/Morozovella velascoensis zones (earliest Ypresian). Sedimentologically, four sedimentary facies belts forming southwest gently-dipping slope to basin transect have been detected. They include tidal flats, outer shelf, slumped continental slope and open marine hemipelagic facies. This transect can be subdivided into a stable basin plain plus outer shelf in the extreme southwestern parts; and an unstable slope shelf platform in the northeastern parts. The unstable slope shelf platform is characterized by open marine hemipelagic, fine-grained limestones and fine siliciclastic shales (Sudr, Dakhla, Tarawan and Esna formations). The northeastern parts are marked by little contents of planktonic foraminifera and dolomitized, slumped carbonates, intercalated with basinal facies. Tectonically, four remarkable syn-depositional tectonic events (SdTEs) controlled the evolution of the studied succession. These events took place strongly within the Campanian–Ypresian time interval and were still active till Late Eocene. These events took place at: the Santonian/Campanian (S/C) boundary; the Campanian/Maastrichtian (C/M) boundary; the Cretaceous/Paleogene (K/P) boundary; and the Middle Paleocene–Early Eocene interval. These tectonic events are four pronounced phases in the tectonic history of the Syrian Arc System (SAS), the collision of the Afro-Arabian and Eurasian plates as well as the closure of the Tethys Sea.     

Dinosaur eggshells from the Santonian Milk River Formation of Alberta,Canada

The North American fossil record of dinosaur eggshells for the Cretaceous is primarily restricted to formations of the middle (Albian–Cenomanian) and uppermost (Campanian–Maastrichtian) stages, with a large gap in the record for intermediate stages. Here we describe a dinosaur eggshell assemblage from a formation that represents an intermediate and poorly fossiliferous stage of the Upper Cretaceous, the Santonian Milk River Formation of southern Alberta, Canada. The Milk River eggshell assemblage contains five eggshell taxa: Continuoolithus, Porituberoolithus, Prismatoolithus, Spheroolithus, and Triprismatoolithus. These ootaxa are most similar to those reported from younger Campanian–Maastrichtian formations of the northern Western Interior than they are to ootaxa reported from older middle Cretaceous formations (i.e., predominantly Macroelongatoolithus). Characteristics of the Milk River ootaxa indicate that they are ascribable to at least one ornithopod and four small theropod species. The taxonomic affinity of the eggshell assemblage is consistent with the dinosaur fauna known based on isolated teeth and fragmentary skeletal remains from the formation, although most ornithischians and large theropods are not represented by eggshell. Relative to the Milk River Formation eggshell, similar oospecies occurring in younger Cretaceous deposits tend to be somewhat thicker, which may reflect an increase in body size of various dinosaur lineages during the Late Cretaceous.     

塔里木盆地西部晚白垩世钙质超微化石

本文分析研究了塔里木盆地西部晚白垩世钙质超微化石,划分出Quadrumgartneri,Eprolithus floralis两个组合,从一个新的角度为探讨库克拜组和乌依塔克组的地层时代及其沉积环境提供有关的论据.     

Uintacrinus anglicus Rasmussen from the Upper Cretaceous Flamborough Chalk Formation of Yorkshire: implications for the position of the Santonian-Campanian boundary

Calyx plates of the crinoid Uintacrinus anglicus are recorded from the top and bottom of a 5 m interval, 4 m above the disappearance of calyx plates of the crinoid Marsupites testudinarius within the Flamborough Chalk Formation at Danes Dyke in North Yorkshire, UK. The U. anglicus Zone, as used here, comprises the interval from the disappearance of M. testudinarius to the disappearance of U. anglicus calyx plates, and therefore includes, but does not comprise, the total range of the index species. Two subzones are recognized in the M. testudinarius Zone; a lower subzone characterized by smooth Marsupites calyx plates and an upper subzone characterized by variably ornamented Marsupites calyx plates. A provisional study of the belemnite indicates the Gonioteuthis granulataquadrata appears in the highest part of the M. testudinarius Zone. The 1983 Copenhagen symposium on Cretaceous stage boundaries proposed that the base of the Campanian Stage should be drawn at a level close to the appearance of G. granulataquadrata. Consequently, the extinction of M. testudinarius is used to define the base of the Campanian here and U. anglicus Zone is placed in the basal Lower Campanian.     

Infrazonal biostratigraphy of the Upper Cretaceous in the East European province based on benthic foraminifers,Part 2: Santonian-Maastrichtian

A chart of infrazonal biostratigraphic subdivisions in the Cenomanian-Coniacian succession of the East European paleobiogeographic province is substantiated based on distribution of benthic foraminifers. The suggested chart characterizes successive trend of changes in ecologic assemblages of benthic foraminifers and morphologic evolution of certain agglutinated (Gaudryina, Ataxophragmium, Ataxoorbignyina, Arenobulimina, Novatrix, Voloshinovella, Orbignyina, Bolivinopsis) and secretory (Neoflabellina, Globorotalites, Stensioeina, Osangularia, Eponides, Gavelinella, Pseudovalvulineria, Pseudogavelinella, Brotzenella, Cibicides, Cibicidoides, Angulogavelinella, Falsoplanulina, Anomalinoides, Coryphostoma, Bolivinoides, Praebulimina) foraminiferal genera. The chart includes 23 biostratigraphic units (zones and subzones), most of which are recognizable over the vast territory from the Mangyshlak to southern Baltic areas. It is correlated with the acknowledged belemnite, inoceramid, ammonoid and echinoid zonations. Eight stadia of taxonomic changes in foraminiferal assemblages, which are distinguished in this work, show that principal biotic events took place across the middle-late Santonian and Santonian-Campanian boundaries, in the earliest Campanian, at the early-late Campanian boundary time, during the late and terminal Campanian, and in the mid-early Maastrichtian.