| 进化序列系统学——以二叠纪瓦岗珊瑚类为例 |
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| 引用本文： | 徐桂荣,龚淑云.进化序列系统学——以二叠纪瓦岗珊瑚类为例[J].地球科学,2000,25(1):1-10. |
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| 作者姓名： | 徐桂荣 龚淑云 |
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| 作者单位： | 中国地质大学地球科学学院，武汉 430074 |
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| 基金项目： | 国家教委博士点基金项目!(No .970 14 0 0 1)，国家自然科学基金项目!(No .49872 0 14 ) |
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| 摘 要： | 进化序列系统学是古生物学中研究物种间和类别间的系统发育关系为目的的学科, 包括研究物种或类别间的亲缘关系, 并确定它们形成的先后序列.表型-分支系统学是该系统学的基础.分支系统学的许多重要概念在该系统学中加入了时间的含义, 例如, 认为最亲近的姐妹种之间的时间间隔不会超过它们母种的生存期.姐妹群和性状镶嵌分析是进化序列研究的主要方法, 而理解性状镶嵌分布是关键.这些分析可由计算机程序实现.以二叠纪瓦岗珊瑚类作为研究进化序列系统学的实例, 该类在特提斯地区广泛分布.通过对瓦岗珊瑚类的特征分析确定了17个性状用于研究, 并且每一性状确定了几个变化状态.根据最广布的性状状态、化石记录和个体发生等原则分辨出近祖性状状态, 并根据性状在系统发育中的重要性确定性状分级.然后, 确定了姐妹群间的关系和物种间的进化序列.在姐妹群与进化序列分析相互检验中可发现性状退化、平行演化和多向分支等古生物学重要现象, 并分辨出物种的异常顺序.在检验生物地层学首次出现事件讨论中推测了一些化石首次出现的可能层位.
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| 关 键 词： | 进化序列系统学 性状镶嵌分布 二叠纪 瓦岗珊瑚类 |
| 文章编号： | 1000-2383(2000)01-0001-10 |
| 收稿时间： | 1999-06-25 |
EVOLUTIONARY SEQUENCE SYSTEMATICS: CASE ANALYSIS OF PERMIAN WAAGENOPHYLLOID CORALS |
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| Xu Guirong,Gong Shuyun,Wang Yongbiao.EVOLUTIONARY SEQUENCE SYSTEMATICS: CASE ANALYSIS OF PERMIAN WAAGENOPHYLLOID CORALS[J].Earth Science-Journal of China University of Geosciences,2000,25(1):1-10. |
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| Authors: | Xu Guirong Gong Shuyun Wang Yongbiao |
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| Abstract: | Evolutionary sequence systematics, an academic discipline in paleontology and biology, is oriented toward the research into the systematic growth association between species or taxa, including the relative relations between species or taxa, and also toward the determination of sequence of their formation. The systematics is based on the phenetic-cladistic systematics. Many important theoretical concepts in the cladistics include the time concept in this systematics. For example, the time-span between two closest sister species can not exceed the life span of their mother species. The analysis of sister-group and character mosaic distribution may serve as a major method for the research into the evolutionary sequence. However, the understanding of the character mosaic distribution is the key to this research. These analyses can be executed with the computer programs. The Permian Waagenophylloid coral fauna is taken as an example to illustrate the analysis of evolutionary sequence. These coral fossils are widespread in the Tethyan region. Seventeen characters and several changing states for each character were determined through the analysis of character features of Waagenophylloid corals. The plesiomorph character states for each character were identified in line with the principles of the widest distribution of character states, fossil records and ontogenesis. In addition, these characters are classified as 4 ranks on the basis of their important places in phylogeny. Then the relationship between sister groups and evolutionary sequence between species were determined following the theory and methodology of the evolutionary sequence systematics. In the mutual proof-tests between the sister groups and evolutionary sequence, some important palaeontological phenomena were discovered such as character degeneration, parallel evolution, and poly-directional clades. At the same time, a number of abnormal orders were identified. From the first events that occurred in the biostratigraphical test, possible stratigraphic horizons of some fossils can be deduced. |
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| Keywords: | evolutionary sequence systematics character mosaic distribution Permian Waagenophylloid corals |
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